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<document id="8305AD302081945B923627D5EDC69A52" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE41B62DDD9E4620FA8302A4" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius longipenis S. Tarasov 2017, new species" docType="treatment" docUuid="9BE1948F-A942-492A-A07A-278B88EE090D" docUuidSource="ZooBank" docVersion="12" lastPageNumber="129" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="129" updateTime="1698469310636" updateUser="ExternalLinkService">
<mods:mods id="25E5B9943DB708022AC6AA219E122144" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="78775083F1591BC7BEB0751B198F1CA4">
<mods:title id="F3711A32D77744F734F23998B8C412A3">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:name id="153B13DD47433F020FF2889FD29A4892" type="personal">
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<mods:roleTerm id="2E8ED3B8CB6EF490F2451D7FB0633E49">Author</mods:roleTerm>
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<mods:namePart id="04EF132E59EAD783D4F19EC69E3091DA">Tarasov, Sergei</mods:namePart>
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<mods:title id="1A9191D19BF156BC72B82456A709FB6D">Zootaxa</mods:title>
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<mods:part id="F7ADE9D1D21A408016EC376DDB6B8121">
<mods:date id="0E3D696C3A5D6D383DAFD226FAD2C322">2017</mods:date>
<mods:detail id="296D38AB27BD2DA7BE3A49B315218128" type="pubDate">
<mods:number id="A6091A47A4A56FC04B3746AF4F712426">2017-10-04</mods:number>
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<mods:number id="2C613238FE298C5DEE4FCD831B59D162">4329</mods:number>
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<mods:classification id="A20EF2B894A61FE7F817403A7155F150">journal article</mods:classification>
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<mods:identifier id="A6056B059FEA035B944CB8CBC99123A2" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<mods:identifier id="599A3184C217DADE1C843E10C51ABEA1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C187E1AE41B62DDD9E4620FA8302A4" ID-DOI="http://doi.org/10.5281/zenodo.6010830" ID-GBIF-Taxon="135106139" ID-Zenodo-Dep="6010830" LSID="urn:lsid:zoobank.org:act:9BE1948F-A942-492A-A07A-278B88EE090D" httpUri="http://treatment.plazi.org/id/03C187E1AE41B62DDD9E4620FA8302A4" lastPageId="29" lastPageNumber="129" pageId="28" pageNumber="129">
<subSubSection id="C372657CAE41B62CDD9E4620FE9903DB" pageId="28" pageNumber="129" type="nomenclature">
<paragraph id="8BD736F7AE41B62CDD9E4620FD6803FA" blockId="28.[151,967,367,463]" box="[151,691,367,394]" pageId="28" pageNumber="129">
<heading id="D09F819BAE41B62CDD9E4620FD6803FA" bold="true" box="[151,691,367,394]" fontSize="11" level="1" pageId="28" pageNumber="129" reason="1">
<emphasis id="B91CEAE5AE41B62CDD9E4620FD6803FA" bold="true" box="[151,691,367,394]" pageId="28" pageNumber="129">
<taxonomicName id="4C684D74AE41B62CDD9E4620FDC603FA" ID-CoL="6TNM7" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[151,541,367,394]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="longipenis" status="sp. nov.">
<emphasis id="B91CEAE5AE41B62CDD9E4620FE6A03F9" bold="true" box="[151,433,367,393]" italics="true" pageId="28" pageNumber="129">Parachorius longipenis</emphasis>
Tarasov
</taxonomicName>
<taxonomicNameLabel id="A22F579EAE41B62CDF2D463FFD6803FA" box="[548,691,368,394]" pageId="28" pageNumber="129" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736F7AE41B62CDD9E46DDFE9903DB" blockId="28.[151,967,367,463]" box="[151,322,402,427]" pageId="28" pageNumber="129">
(
<figureCitation id="13532A72AE41B62CDD9646DDFF2703DB" box="[159,252,402,427]" captionStart="FIGURE 15" captionStartId="29.[151,250,1247,1269]" captionTargetBox="[151,1436,267,1223]" captionTargetId="figure@29.[151,1436,267,1225]" captionTargetPageId="29" captionText="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri="https://zenodo.org/record/1001947/files/figure.png" pageId="28" pageNumber="129">Figs. 15</figureCitation>
,
<figureCitation id="13532A72AE41B62CDC0146DDFEE103DB" box="[264,314,402,427]" captionStart="FIGURE 26" captionStartId="43.[151,250,1905,1927]" captionTargetBox="[151,1436,265,1825]" captionTargetId="figure@43.[151,1436,265,1825]" captionTargetPageId="43" captionText="FIGURE 26. Distribution maps of Parachorius species. (A) Parachorius hookeri. (B) Parachorius humeralis. (C) Parachorius javanus. (D) Parachorius longipenis. (E) Parachorius maruyamai. (F) Parachorius newthayerae. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study." httpUri="https://zenodo.org/record/1001969/files/figure.png" pageId="28" pageNumber="129">26D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDD9E46F9FC1C03BF" box="[151,967,438,463]" pageId="28" pageNumber="129" type="description">
<paragraph id="8BD736F7AE41B62CDD9E46F9FC1C03BF" blockId="28.[151,967,367,463]" box="[151,967,438,463]" pageId="28" pageNumber="129">urn:lsid:zoobank.org:act:9BE1948F-A942-492A-A07A-278B88EE090D</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDD9E46B1FDC70067" box="[151,540,510,535]" pageId="28" pageNumber="129" type="materials_examined">
<paragraph id="8BD736F7AE41B62CDD9E46B1FDC70067" blockId="28.[151,1437,510,2011]" box="[151,540,510,535]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDD9E46B1FEF60067" bold="true" box="[151,301,510,535]" pageId="28" pageNumber="129">Type locality</emphasis>
. Hoa Binh, Vietnam.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDDCE456DFB00002E" pageId="28" pageNumber="129" type="etymology">
<paragraph id="8BD736F7AE41B62CDDCE456DFB00002E" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE456DFE95004B" bold="true" box="[199,334,546,571]" pageId="28" pageNumber="129">Etymology.</emphasis>
The name of this species derives from the Latin words &quot;longus&quot; and &quot;penis&quot; reflecting the fact that this species has a relatively elongated penis in comparison to the similar species
<taxonomicName id="4C684D74AE41B62CD9144508FB0E002F" authorityName="Tarasov" authorityYear="2017" box="[1053,1237,582,607]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="solodovnikovi">
<emphasis id="B91CEAE5AE41B62CD9144508FB0E002F" box="[1053,1237,582,607]" italics="true" pageId="28" pageNumber="129">P. solodovnikovi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDDCE4525FBDA00D7" pageId="28" pageNumber="129" type="distribution">
<paragraph id="8BD736F7AE41B62CDDCE4525FBDA00D7" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE4525FEBB00F3" bold="true" box="[199,352,618,643]" pageId="28" pageNumber="129">Distribution.</emphasis>
The species is known from
<collectingCountry id="F37F7667AE41B62CDF944525FD2500F3" box="[669,766,618,643]" name="Vietnam" pageId="28" pageNumber="129">Vietnam</collectingCountry>
,
<collectingCountry id="F37F7667AE41B62CDE034524FC9800F2" box="[778,835,619,642]" name="Laos" pageId="28" pageNumber="129">Laos</collectingCountry>
, and
<collectingCountry id="F37F7667AE41B62CDE774525FC3E00F3" box="[894,997,618,643]" name="Thailand" pageId="28" pageNumber="129">Thailand</collectingCountry>
. It is abundant in collections compared to other species of
<taxonomicName id="4C684D74AE41B62CDC6345C1FE2D00D7" box="[362,502,654,679]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE41B62CDC6345C1FE2D00D7" box="[362,502,654,679]" italics="true" pageId="28" pageNumber="129">Parachorius</emphasis>
</taxonomicName>
and therefore, likely widespread in Indochina.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDDCE45FDFC000147" pageId="28" pageNumber="129" type="diagnosis">
<paragraph id="8BD736F7AE41B62CDDCE45FDFC000147" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE45FDFE9A00BB" bold="true" box="[199,321,690,715]" pageId="28" pageNumber="129">Diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="4C684D74AE41B62CDFA245FBFCD000BB" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[683,779,690,715]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="nudus">
<emphasis id="B91CEAE5AE41B62CDFA245FBFCD000BB" box="[683,779,690,715]" italics="true" pageId="28" pageNumber="129">P. nudus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C684D74AE41B62CDE1345FBFC0800BB" authorityName="Tarasov" authorityYear="2017" box="[794,979,690,715]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="solodovnikovi">
<emphasis id="B91CEAE5AE41B62CDE1345FBFC0800BB" box="[794,979,690,715]" italics="true" pageId="28" pageNumber="129">P. solodovnikovi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C684D74AE41B62CD91A45FBFB6000BB" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[1043,1211,690,715]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91CEAE5AE41B62CD91A45FBFB6000BB" box="[1043,1211,690,715]" italics="true" pageId="28" pageNumber="129">P. peninsularis</emphasis>
</taxonomicName>
. From them and all other
<taxonomicName id="4C684D74AE41B62CDDD14599FEBF009F" box="[216,356,726,751]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE41B62CDDD14599FEBF009F" box="[216,356,726,751]" italics="true" pageId="28" pageNumber="129">Parachorius</emphasis>
</taxonomicName>
, it can be separated by (1) protibia with first tooth spatulate, bent downward and notably larger than second tooth, (2) first tooth also looks bifurcated as it merged with denticles located between first and second teeth, (3) aedeagal parameres elongate with horn-like upper lobes dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62CDDCE440DFC3E075F" pageId="28" pageNumber="129" type="description">
<paragraph id="8BD736F7AE41B62CDDCE440DFBEF012B" blockId="28.[151,1437,510,2011]" box="[199,1076,834,859]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE440DFE82012B" bold="true" box="[199,345,834,859]" pageId="28" pageNumber="129">Description.</emphasis>
<emphasis id="B91CEAE5AE41B62CDC56440DFE44012B" box="[351,415,834,859]" italics="true" pageId="28" pageNumber="129">Body.</emphasis>
Body black, length
<quantity id="4C909B12AE41B62CDF88440DFCD0012A" box="[641,779,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.6" metricValueMin="3.8" pageId="28" pageNumber="129" unit="mm" value="4.699999999999999" valueMax="5.6" valueMin="3.8">3.8–5.6 mm</quantity>
. Elytra uniformly colored.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE4429FBAD010F" blockId="28.[151,1437,510,2011]" box="[199,1142,870,895]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE4429FED0010F" box="[199,267,870,895]" italics="true" pageId="28" pageNumber="129">Head.</emphasis>
Clypeus with two teeth, outer margin of each tooth distinctly notched basally.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE44C3FB7601B7" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE44C3FEE601D3" box="[199,317,908,931]" italics="true" pageId="28" pageNumber="129">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 0.3– 1.0 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE449DFD1B019A" blockId="28.[151,1437,510,2011]" box="[199,704,978,1003]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE449DFECE019B" box="[199,277,978,1003]" italics="true" pageId="28" pageNumber="129">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE44B8FD94067F" blockId="28.[151,1437,510,2011]" box="[199,591,1014,1039]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE44B8FEC9067E" box="[199,274,1015,1038]" italics="true" pageId="28" pageNumber="129">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE4353FE520697" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE4353FED80643" box="[199,259,1052,1075]" italics="true" pageId="28" pageNumber="129">Legs.</emphasis>
Profemora with almost straight or sinuate anterior margin. Protibia apical spur reaching protarsomere 2. First tooth notably modified in males, spatulate, bent downward and notably larger than second tooth, with which it is almost merged; first tooth may look bifurcate because it is merged with denticles located between first and second teeth. Metafemoral posterior margin not serrate, it varies from simple to dilate with produced angle in basal half of metafemur. Metatibia produced in small angle apically, slightly sinuate, inner margin denticulate with relatively small teeth.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE43BBFBBD077B" blockId="28.[151,1437,510,2011]" box="[199,1126,1266,1291]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE43BBFEB9077B" box="[199,354,1268,1291]" italics="true" pageId="28" pageNumber="129">Metasternum.</emphasis>
Transverse basal ridge of metasternum reduced and extremely short.
</paragraph>
<paragraph id="8BD736F7AE41B62CDDCE4259FC3E075F" blockId="28.[151,1437,510,2011]" box="[199,997,1302,1327]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE4259FEE0075F" box="[199,315,1302,1327]" italics="true" pageId="28" pageNumber="129">Aedeagus.</emphasis>
Parameres symmetrical, with horn-like upper lobes dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE41B62DDDCE4275FA8302A4" lastPageId="29" lastPageNumber="130" pageId="28" pageNumber="129" type="materials_examined">
<paragraph id="8BD736F7AE41B62CDDCE4275FE9A0707" blockId="28.[151,1437,510,2011]" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDDCE4275FEE00723" bold="true" box="[199,315,1338,1363]" pageId="28" pageNumber="129">
<typeStatus id="54D38855AE41B62CDDCE4275FEED0723" box="[199,310,1338,1363]" pageId="28" pageNumber="129" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="13532A72AE41B62CDC804275FE370723" box="[393,492,1338,1363]" captionStart="FIGURE 15" captionStartId="29.[151,250,1247,1269]" captionTargetBox="[151,1436,267,1223]" captionTargetId="figure@29.[151,1436,267,1225]" captionTargetPageId="29" captionText="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri="https://zenodo.org/record/1001947/files/figure.png" pageId="28" pageNumber="129">Fig. 15F</figureCitation>
). Profemur with sinuate anterior margin. First tooth bifurcate. Metafemoral posterior margin dilated.
</paragraph>
<paragraph id="8BD736F7AE41B62DDDCE42CDFA8302A4" blockId="28.[151,1437,510,2011]" lastBlockId="29.[151,1436,151,213]" lastPageId="29" lastPageNumber="130" pageId="28" pageNumber="129">
<materialsCitation id="3B003CAAAE41B62CDDCE42CDFD4C07EB" ID-GBIF-Occurrence="1658411473" box="[199,663,1410,1435]" country="Vietnam" location="Vietnam" pageId="28" pageNumber="129" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEAE5AE41B62CDDCE42CDFDFF07EB" bold="true" box="[199,548,1410,1435]" pageId="28" pageNumber="129">
Material examined.
<typeStatus id="54D38855AE41B62CDCBE42CDFDFF07EB" box="[439,548,1410,1435]" pageId="28" pageNumber="129" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F37F7667AE41B62CDF3A42CDFD4C07EB" box="[563,663,1410,1435]" name="Vietnam" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDF3A42CDFD4C07EB" bold="true" box="[563,663,1410,1435]" pageId="28" pageNumber="129">Vietnam</emphasis>
</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B003CAAAE41B62CDFA142CDFB3107CF" ID-GBIF-Occurrence="1658411489" collectionCode="MNHN" country="Vietnam" latitude="20.828333" location="Cooman" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Hoa Binh" typeStatus="holotype">
<emphasis id="B91CEAE5AE41B62CDFA142CDFD2C07EB" bold="true" box="[680,759,1410,1435]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDFA142CDFD2C07EB" box="[680,759,1410,1435]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
</emphasis>
, &quot;
<collectingRegion id="49ACF815AE41B62CDE0442CCFC5F07EB" box="[781,900,1410,1435]" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="129">Hoa Binh</collectingRegion>
[,]
<collectingMunicipality id="6BB3AC8DAE41B62CDE9442CDFC3707EB" box="[925,1004,1410,1435]" pageId="28" pageNumber="129">Tonkin</collectingMunicipality>
[northern
<collectingCountry id="F37F7667AE41B62CD96B42CDFB1C07EB" box="[1122,1223,1410,1435]" name="Vietnam" pageId="28" pageNumber="129">Vietnam</collectingCountry>
,] [leg.] de
<location id="8EB7602CAE41B62CD83442CCFA4707EB" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD83442CCFA4707EB" box="[1341,1436,1411,1435]" country="Vietnam" latitude="20.828333" longLatPrecision="870" longitude="105.333336" municipality="Tonkin" name="Cooman" pageId="28" pageNumber="129" stateProvince="Hoa Binh">Cooman</location>
/ 57 /
<typeStatus id="54D38855AE41B62CDDD442E8FEAA07CF" box="[221,369,1447,1471]" pageId="28" pageNumber="129" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C684D74AE41B62CDC7342E9FD2607CE" authority="S. Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[378,765,1446,1471]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="129" phylum="Arthropoda" rank="species" species="longipenis">Parachorius longipenis S. Tarasov</taxonomicName>
det. 2010&quot;,
<geoCoordinate id="EE5C5030AE41B62CDE8542E8FBDB07CF" box="[908,1024,1447,1471]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="92" value="20.828333">20°49.7'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CD90442E9FBAE07CF" box="[1037,1141,1446,1471]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="925" value="105.333336">105°20'E</geoCoordinate>
(
<collectionCode id="ED79AE32AE41B62CD98C42E8FB3A07CF" box="[1157,1249,1447,1471]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="129">MNHN</collectionCode>
)
</materialsCitation>
.
<emphasis id="B91CEAE5AE41B62CD9FF42E9FF3E0793" bold="true" pageId="28" pageNumber="129">Paratypes (45 males)</emphasis>
:
<materialsCitation id="3B003CAAAE41B62CDDFC4285FC240793" ID-GBIF-Occurrence="1658411477" box="[245,1023,1482,1507]" collectionCode="MNHN" country="Laos" location="Bonnote" municipality="Cambodge" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CDDFC4285FE630793" bold="true" box="[245,440,1482,1507]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDDFC4285FE9E0793" box="[245,325,1482,1507]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDC444285FE630793" box="[333,440,1482,1507]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingCountry id="F37F7667AE41B62CDCC14284FE250792" box="[456,510,1483,1506]" name="Laos" pageId="28" pageNumber="129">Laos</collectingCountry>
ou
<collectingMunicipality id="6BB3AC8DAE41B62CDF254285FD730793" box="[556,680,1482,1507]" pageId="28" pageNumber="129">Cambodge</collectingMunicipality>
, on 1911 (
<location id="8EB7602CAE41B62CDE294284FC590793" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CDE294284FC590793" box="[800,898,1483,1507]" country="Laos" municipality="Cambodge" name="Bonnote" pageId="28" pageNumber="129">Bonnote</location>
) (
<collectionCode id="ED79AE32AE41B62CDE934284FC2D0793" box="[922,1014,1483,1507]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="129">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CD9054285FA58045B" ID-GBIF-Occurrence="1658411468" collectingDate="2008-06-02" collectingDateMax="2008-06-19" collectingDateMin="2008-06-02" collectionCode="ANIMT, FMNH" collectorName="Pedersen &amp; M. Thayer &amp; A. Newton" country="Laos" elevation="190" latitude="18.098717" location="Houayang National Park" longLatPrecision="1" longitude="102.6688" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="paratype">
<collectingCountry id="F37F7667AE41B62CD9054285FB9E0793" box="[1036,1093,1482,1507]" name="Laos" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CD9054285FB9E0793" bold="true" box="[1036,1093,1482,1507]" pageId="28" pageNumber="129">Laos</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE41B62CD95C4285FAC20793" bold="true" box="[1109,1305,1482,1507]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CD95C4285FB7D0793" box="[1109,1190,1482,1507]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CD9A74285FAC20793" box="[1198,1305,1482,1507]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<location id="8EB7602CAE41B62CD8204284FEE70477" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD8204284FEE70477" country="Laos" latitude="18.098717" longLatPrecision="1" longitude="102.6688" name="Houayang National Park" pageId="28" pageNumber="129" stateProvince="Vientiane">Houayang National Park</location>
, approximately
<locationDeviation id="9A652FD5AE41B62CDF0942A1FCCD0477" box="[512,790,1518,1543]" pageId="28" pageNumber="129">
<quantity id="4C909B12AE41B62CDF0942A1FD940477" box="[512,591,1518,1543]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="129" unit="km" value="15.0">15 km</quantity>
N of
<collectingRegion id="49ACF815AE41B62CDFA142A1FCCD0477" box="[680,790,1518,1543]" country="Laos" name="Viangchan" pageId="28" pageNumber="129">Vientiane</collectingRegion>
</locationDeviation>
,
<collectionCode id="ED79AE32AE41B62CDE2042A0FC5E0477" box="[809,901,1519,1543]" pageId="28" pageNumber="129">ANIMT</collectionCode>
site 1225,
<collectingMethod id="52294EE0AE41B62CD91142A1FB790476" box="[1048,1186,1518,1543]" pageId="28" pageNumber="129">carrion trap</collectingMethod>
(squid),
<geoCoordinate id="EE5C5030AE41B62CD81F42A1FA4C0477" box="[1302,1431,1518,1543]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="1" value="18.098717">18°5.923'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDD9E415CFEE2045A" box="[151,313,1555,1579]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="1" value="102.6688">102°40.128' E</geoCoordinate>
,
<elevation id="0045D1C4AE41B62CDC4D415CFE56045A" box="[324,397,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="129" unit="m" value="190.0">
<quantity id="4C909B12AE41B62CDC4D415CFE56045A" box="[324,397,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="129" unit="m" value="190.0">190 m</quantity>
</elevation>
, tropical handwood forest, on
<collectingDate id="EF92E9DFAE41B62CDFE8415DFCA9045B" box="[737,882,1554,1579]" pageId="28" pageNumber="129" value="2008-06-02" valueMax="2008-06-19" valueMin="2008-06-02">
<date id="FFD61037AE41B62CDFE8415DFCA9045B" box="[737,882,1554,1579]" pageId="28" pageNumber="129" value="2008-06-02" valueMax="2008-06-19" valueMin="2008-06-02">2-19/vi/2008</date>
</collectingDate>
(J.
<collectorName id="269D5321AE41B62CDE92415DFBD8045B" box="[923,1027,1554,1579]" pageId="28" pageNumber="129">Pedersen</collectorName>
,
<collectorName id="269D5321AE41B62CD907415CFB5D045B" box="[1038,1158,1554,1579]" pageId="28" pageNumber="129">M. Thayer</collectorName>
,
<collectorName id="269D5321AE41B62CD998415CFAD6045B" box="[1169,1293,1555,1579]" pageId="28" pageNumber="129">A. Newton</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CD82D415CFAA7045B" box="[1316,1404,1555,1579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="28" pageNumber="129">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CD887415DFDC40403" ID-GBIF-Occurrence="1658411476" collectingDate="2008-06-12" collectingDateMax="2008-06-19" collectingDateMin="2008-06-12" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="Laos" elevation="200" latitude="18.097666" location="Houayang forest" longLatPrecision="9" longitude="102.67578" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Vientiane" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CD887415DFE98043F" bold="true" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CD887415DFF0A043F" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDDD14179FE98043F" box="[216,323,1590,1615]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE41B62CDC5A4179FE1A043F" box="[339,449,1590,1615]" country="Laos" name="Viangchan" pageId="28" pageNumber="129">Vientiane</collectingRegion>
,
<location id="8EB7602CAE41B62CDCC44178FD50043F" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CDCC44178FD50043F" box="[461,651,1590,1615]" country="Laos" latitude="18.097666" longLatPrecision="9" longitude="102.67578" name="Houayang forest" pageId="28" pageNumber="129" stateProvince="Vientiane">Houayang forest</location>
,
<geoCoordinate id="EE5C5030AE41B62CDF9E4179FCD0043F" box="[663,779,1590,1615]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="9" value="18.097666">18°5.86'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDE114179FC69043F" box="[792,946,1590,1615]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="1" value="102.67578">102°40.547'E</geoCoordinate>
,
<elevation id="0045D1C4AE41B62CDEB64178FBD2043E" box="[959,1033,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="129" unit="m" value="200.0">
<quantity id="4C909B12AE41B62CDEB64178FBD2043E" box="[959,1033,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="129" unit="m" value="200.0">200 m</quantity>
</elevation>
, disturbed forest, on
<collectingDate id="EF92E9DFAE41B62CD9F04179FA46043F" box="[1273,1437,1590,1615]" pageId="28" pageNumber="129" value="2008-06-12" valueMax="2008-06-19" valueMin="2008-06-12">
<date id="FFD61037AE41B62CD9F04179FA46043F" box="[1273,1437,1590,1615]" pageId="28" pageNumber="129" value="2008-06-12" valueMax="2008-06-19" valueMin="2008-06-12">12–19/vi/2008</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDDA94114FEFB0402" box="[160,288,1627,1651]" pageId="28" pageNumber="129">A. Newton</collectorName>
,
<collectorName id="269D5321AE41B62CDC244114FE7C0403" box="[301,423,1626,1651]" pageId="28" pageNumber="129">M. Thayer</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDCC94114FDCC0403" box="[448,535,1627,1651]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="28" pageNumber="129">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CDF254115FC7604E7" ID-GBIF-Occurrence="1658411474" collectingDate="2007-09-01" collectionCode="ZMUC" collectorName="H. Enghoff" country="Thailand" latitude="15.7" location="Pa Him Ngam" longLatPrecision="1284" longitude="101.38333" municipality="Thep Sathit district" pageId="28" pageNumber="129" specimenCount="2" specimenCount-male="2" stateProvince="Chaiyaphum province" typeStatus="paratype">
<collectingCountry id="F37F7667AE41B62CDF254115FD420403" box="[556,665,1626,1651]" name="Thailand" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDF254115FD420403" bold="true" box="[556,665,1626,1651]" pageId="28" pageNumber="129">Thailand</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE41B62CDFA34115FCA10403" bold="true" box="[682,890,1626,1651]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDFA34115FD200403" box="[682,763,1626,1651]" pageId="28" pageNumber="129" type="male">2 male</specimenCount>
<typeStatus id="54D38855AE41B62CDE0D4115FCA10403" box="[772,890,1626,1651]" pageId="28" pageNumber="129" type="paratype">paratypes</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE41B62CDE824115FB530403" box="[907,1160,1626,1651]" country="Thailand" name="Chaiyaphum" pageId="28" pageNumber="129">Chaiyaphum province</collectingRegion>
,
<collectingMunicipality id="6BB3AC8DAE41B62CD99C4115FAA80403" box="[1173,1395,1626,1651]" pageId="28" pageNumber="129">Thep Sathit district</collectingMunicipality>
,
<location id="8EB7602CAE41B62CD8894114FEC104E7" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD8894114FEC104E7" country="Thailand" latitude="15.7" longLatPrecision="1284" longitude="101.38333" municipality="Thep Sathit district" name="Pa Him Ngam" pageId="28" pageNumber="129" stateProvince="Chaiyaphum province">Pa Him Ngam</location>
,
<geoCoordinate id="EE5C5030AE41B62CDC214131FE5C04E7" box="[296,391,1662,1687]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="925" value="15.7">15°42'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDC9C4130FE2404E7" box="[405,511,1663,1687]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="925" value="101.38333">101°23'E</geoCoordinate>
, on
<collectingDate id="EF92E9DFAE41B62CDF3D4131FD4404E7" box="[564,671,1662,1687]" pageId="28" pageNumber="129" value="2007-09-01">
<date id="FFD61037AE41B62CDF3D4131FD4404E7" box="[564,671,1662,1687]" pageId="28" pageNumber="129" value="2007-09-01">1/ix/2007</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDFBB4130FCEF04E7" box="[690,820,1662,1687]" pageId="28" pageNumber="129">H. Enghoff</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDE474130FC7E04E7" box="[846,933,1663,1687]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="28" pageNumber="129">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CDEB24131FE4704CB" ID-GBIF-Occurrence="1658411471" collectingDate="1929" collectingDateMax="1933" collectingDateMin="1929" collectionCode="ZMUC" collectorName="Poul Fogh" country="Thailand" location="Siam" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CDEB24131FB5A04E7" bold="true" box="[955,1153,1662,1687]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDEB24131FBD704E7" box="[955,1036,1662,1687]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CD91F4131FB5A04E7" box="[1046,1153,1662,1687]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
, Prae
<collectingCountry id="F37F7667AE41B62CD9C74131FAD104E7" box="[1230,1290,1662,1687]" name="Thailand" pageId="28" pageNumber="129">Siam</collectingCountry>
, on
<collectingDate id="EF92E9DFAE41B62CD8364130FA4704E7" box="[1343,1436,1663,1687]" pageId="28" pageNumber="129" value="1929" valueMax="1933">
<date id="FFD61037AE41B62CD8364130FA4704E7" box="[1343,1436,1663,1687]" pageId="28" pageNumber="129" value="1929" valueMax="1933">1929-33</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDD9741EDFEC504CB" box="[158,286,1698,1723]" pageId="28" pageNumber="129">Poul Fogh</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDC3441ECFE4F04CB" box="[317,404,1699,1723]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="28" pageNumber="129">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CDCB941EDFAEF04AF" ID-GBIF-Occurrence="1658411479" collectingDate="2005-07-04" collectingDateMax="2005-09-06" collectingDateMin="2005-07-04" collectionCode="OUMNH" collectorName="Limestone Karst &amp; L. Hayes" country="Vietnam" elevation="290" latitude="22.4175" location="Ba Be National Park" longLatPrecision="21" longitude="105.63306" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan Province" typeStatus="paratype">
<collectingCountry id="F37F7667AE41B62CDCB941EDFDCF04CB" box="[432,532,1698,1723]" name="Vietnam" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDCB941EDFDCF04CB" bold="true" box="[432,532,1698,1723]" pageId="28" pageNumber="129">Vietnam</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE41B62CDF2441EDFD2404CB" bold="true" box="[557,767,1698,1723]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDF2441EDFD5F04CB" box="[557,644,1698,1723]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDF9A41EDFD2404CB" box="[659,767,1698,1723]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE41B62CDE1C41ECFC2E04CB" box="[789,1013,1698,1723]" country="Vietnam" name="Bac Kan" pageId="28" pageNumber="129">
Bac
<collectingRegion id="49ACF815AE41B62CDE5941ECFC2E04CB" box="[848,1013,1698,1723]" inferredBySuffix="true" pageId="28" pageNumber="129">Kan Province</collectingRegion>
</collectingRegion>
,
<location id="8EB7602CAE41B62CD90141ECFAD004CB" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD90141ECFAD004CB" box="[1032,1291,1698,1723]" country="Vietnam" latitude="22.4175" longLatPrecision="21" longitude="105.63306" name="Ba Be National Park" pageId="28" pageNumber="129" stateProvince="Bac Kan Province">Ba Be National Park</location>
,
<geoCoordinate id="EE5C5030AE41B62CD81641EDFA4C04CB" box="[1311,1431,1698,1723]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="15" value="22.4175">22°25'3&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDD9E4189FEF304AF" box="[151,296,1734,1759]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="15" value="105.63306">105°37'59&quot;E</geoCoordinate>
,
<elevation id="0045D1C4AE41B62CDC324188FE5004AE" box="[315,395,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="28" pageNumber="129" unit="m" value="290.0">
<quantity id="4C909B12AE41B62CDC324188FE5004AE" box="[315,395,1735,1759]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="28" pageNumber="129" unit="m" value="290.0">290 m</quantity>
</elevation>
,
<collectorName id="269D5321AE41B62CDC974189FDB804AE" box="[414,611,1734,1759]" pageId="28" pageNumber="129">Limestone Karst</collectorName>
, disturbed forest, on
<collectingDate id="EF92E9DFAE41B62CDE624189FBC304AF" box="[875,1048,1734,1759]" pageId="28" pageNumber="129" value="2005-07-04" valueMax="2005-09-06" valueMin="2005-07-04">
<date id="FFD61037AE41B62CDE624189FBC304AF" box="[875,1048,1734,1759]" pageId="28" pageNumber="129" value="2005-07-04" valueMax="2005-09-06" valueMin="2005-07-04">4/vii–6/ix/2005</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CD9264188FB4504AF" box="[1071,1182,1735,1759]" pageId="28" pageNumber="129">L. Hayes</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CD9B24188FAF004AF" box="[1211,1323,1735,1759]" pageId="28" pageNumber="129">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CD84F4189FD1F0557" ID-GBIF-Occurrence="1658411486" collectingDate="2005-07-04" collectingDateMax="2005-09-06" collectingDateMin="2005-07-04" collectionCode="OUMNH" collectorName="Limestone Karst &amp; L. Hayes" country="Vietnam" elevation="315" latitude="22.417221" location="Ba Be National Park" longLatPrecision="21" longitude="105.633606" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan Province" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CD84F4189FED90573" bold="true" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CD84F4189FA4704AF" box="[1350,1436,1734,1759]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDD9E41A5FED90573" box="[151,258,1770,1795]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE41B62CDC1A41A4FE3C0573" box="[275,487,1770,1795]" country="Vietnam" name="Bac Kan" pageId="28" pageNumber="129">
Bac
<collectingRegion id="49ACF815AE41B62CDC4141A4FE3C0573" box="[328,487,1770,1795]" inferredBySuffix="true" pageId="28" pageNumber="129">Kan Province</collectingRegion>
</collectingRegion>
,
<location id="8EB7602CAE41B62CDCFD41A4FD3D0573" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CDCFD41A4FD3D0573" box="[500,742,1770,1795]" country="Vietnam" latitude="22.417221" longLatPrecision="21" longitude="105.633606" name="Ba Be National Park" pageId="28" pageNumber="129" stateProvince="Bac Kan Province">Ba Be National Park</location>
,
<geoCoordinate id="EE5C5030AE41B62CDFFD41A5FCB60573" box="[756,877,1770,1795]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="15" value="22.417221">22°25'2&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDE7241A5FC260573" box="[891,1021,1770,1795]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="15" value="105.633606">105°38'1&quot;E</geoCoordinate>
,
<elevation id="0045D1C4AE41B62CD90541A5FB830572" box="[1036,1112,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="28" pageNumber="129" unit="m" value="315.0">
<quantity id="4C909B12AE41B62CD90541A5FB830572" box="[1036,1112,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="3.15" pageId="28" pageNumber="129" unit="m" value="315.0">315 m</quantity>
</elevation>
,
<collectorName id="269D5321AE41B62CD96F41A5FAFE0572" box="[1126,1317,1770,1795]" pageId="28" pageNumber="129">Limestone Karst</collectorName>
, disturbed forest, on
<collectingDate id="EF92E9DFAE41B62CDC024041FE630557" box="[267,440,1806,1831]" pageId="28" pageNumber="129" value="2005-07-04" valueMax="2005-09-06" valueMin="2005-07-04">
<date id="FFD61037AE41B62CDC024041FE630557" box="[267,440,1806,1831]" pageId="28" pageNumber="129" value="2005-07-04" valueMax="2005-09-06" valueMin="2005-07-04">4/vii–6/ix/2005</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDCC34040FDE80557" box="[458,563,1807,1831]" pageId="28" pageNumber="129">L. Hayes</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDF424040FD600557" box="[587,699,1807,1831]" pageId="28" pageNumber="129">OUMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CDFD84041FB5F0557" ID-GBIF-Occurrence="1658411472" box="[721,1156,1806,1831]" country="Vietnam" location="Dong Nai Province" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Dong Nai Province" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CDFD84041FC4E0557" bold="true" box="[721,917,1806,1831]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDFD84041FCFA0557" box="[721,801,1806,1831]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDE234041FC4E0557" box="[810,917,1806,1831]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE41B62CDEAC4040FB5F0557" box="[933,1156,1806,1831]" country="Vietnam" name="Dong Nai" pageId="28" pageNumber="129">
Dong
<collectingRegion id="49ACF815AE41B62CDEE54041FB5F0557" box="[1004,1156,1806,1831]" inferredBySuffix="true" pageId="28" pageNumber="129">Nai Province</collectingRegion>
</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B003CAAAE41B62CD9984040FA4C053B" ID-GBIF-Occurrence="1658411490" collectingDate="2005-05-21" collectingDateMax="2005-06-16" collectingDateMin="2005-05-21" collectionCode="ZMUM" collectorName="D. Fedorenko &amp; Hoa Binh" country="Vietnam" latitude="11.418333" location="Nam Cat Tien National Park" longLatPrecision="129" longitude="107.42667" pageId="28" pageNumber="129" specimenCount="13" specimenCount-male="13" stateProvince="Hoa Binh" typeStatus="paratype">
<location id="8EB7602CAE41B62CD9984040FF16053B" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD9984040FF16053B" country="Vietnam" latitude="11.418333" longLatPrecision="129" longitude="107.42667" name="Nam Cat Tien National Park" pageId="28" pageNumber="129" stateProvince="Hoa Binh">Nam Cat Tien National Park</location>
,
<geoCoordinate id="EE5C5030AE41B62CDDD0407DFE97053B" box="[217,332,1842,1867]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="92" value="11.418333">11°25.1'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDC51407DFE03053B" box="[344,472,1842,1867]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="92" value="107.42667">107°25.6'E</geoCoordinate>
, forest, on
<collectingDate id="EF92E9DFAE41B62CDF5F407DFCD4053B" box="[598,783,1842,1867]" pageId="28" pageNumber="129" value="2005-05-21" valueMax="2005-06-16" valueMin="2005-05-21">
<date id="FFD61037AE41B62CDF5F407DFCD4053B" box="[598,783,1842,1867]" pageId="28" pageNumber="129" value="2005-05-21" valueMax="2005-06-16" valueMin="2005-05-21">21/v–16/vi/2005</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDE29407CFC66053B" box="[800,957,1842,1867]" pageId="28" pageNumber="129">D. Fedorenko</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDEDD407CFBF5053B" box="[980,1070,1843,1867]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="28" pageNumber="129">ZMUM</collectionCode>
)
<emphasis id="B91CEAE5AE41B62CD937407DFAC2053B" bold="true" box="[1086,1305,1842,1867]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CD937407DFB40053B" box="[1086,1179,1842,1867]" pageId="28" pageNumber="129" type="male">13 male</specimenCount>
<typeStatus id="54D38855AE41B62CD9AA407DFAC2053B" box="[1187,1305,1842,1867]" pageId="28" pageNumber="129" type="paratype">paratypes</typeStatus>
</emphasis>
,
<collectorName id="269D5321AE41B62CD821407CFA4C053B" box="[1320,1431,1842,1867]" pageId="28" pageNumber="129">
<collectingRegion id="49ACF815AE41B62CD821407CFA4C053B" box="[1320,1431,1842,1867]" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="129">Hoa Binh</collectingRegion>
</collectorName>
</materialsCitation>
,
<materialsCitation id="3B003CAAAE41B62CDD9E4019FE7205E3" ID-GBIF-Occurrence="1658411482" collectingDate="1923" collectingDateMax="1928" collectingDateMin="1923" collectionCode="MNHN" collectorName="A. De Cooman" country="Vietnam" latitude="20.828333" location="Tonkin" longLatPrecision="870" longitude="105.333336" pageId="28" pageNumber="129" specimenCount="1" typeStatus="paratype">
<location id="8EB7602CAE41B62CDD9E4019FF3D051F" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CDD9E4019FF3D051F" box="[151,230,1878,1903]" country="Vietnam" latitude="20.828333" longLatPrecision="870" longitude="105.333336" name="Tonkin" pageId="28" pageNumber="129">Tonkin</location>
[northern
<collectingCountry id="F37F7667AE41B62CDC564019FE1F051F" box="[351,452,1878,1903]" name="Vietnam" pageId="28" pageNumber="129">Vietnam</collectingCountry>
],
<geoCoordinate id="EE5C5030AE41B62CDCDF4018FD90051F" box="[470,587,1879,1903]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="92" value="20.828333">20°49.7'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDF514019FD1A051F" box="[600,705,1878,1903]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="925" value="105.333336">105°20'E</geoCoordinate>
, on [date specified only for two specimens:
<collectingDate id="EF92E9DFAE41B62CD9B54018FAB2051F" box="[1212,1385,1878,1903]" pageId="28" pageNumber="129" value="1923" valueMax="1928">
<date id="FFD61037AE41B62CD9B54018FAB2051F" box="[1212,1385,1878,1903]" pageId="28" pageNumber="129" value="1923" valueMax="1928">1923 and 1928</date>
</collectingDate>
] (
<collectorName id="269D5321AE41B62CD8894018FEFC05E3" pageId="28" pageNumber="129">A. De Cooman</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDC4C4034FE7A05E3" box="[325,417,1915,1939]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="129">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62CDCB54035FCFD05C7" ID-GBIF-Occurrence="1658411467" collectingDate="1901-09" collectionCode="MNHN" collectorName="A. Weiss" country="Vietnam" latitude="21.816668" location="Central Environs de Tuyen-quan" longLatPrecision="1262" longitude="105.21667" municipality="Tonkin" pageId="28" pageNumber="129" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B91CEAE5AE41B62CDCB54035FD5605E3" bold="true" box="[444,653,1914,1939]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDCB54035FDC805E3" box="[444,531,1914,1939]" pageId="28" pageNumber="129" type="male">1 male</specimenCount>
<typeStatus id="54D38855AE41B62CDF2B4035FD5605E3" box="[546,653,1914,1939]" pageId="28" pageNumber="129" type="paratype">paratype</typeStatus>
</emphasis>
,
<collectingMunicipality id="6BB3AC8DAE41B62CDFAA4035FD2905E3" box="[675,754,1914,1939]" pageId="28" pageNumber="129">Tonkin</collectingMunicipality>
[northern
<collectingCountry id="F37F7667AE41B62CDE714035FC0105E3" box="[888,986,1914,1939]" name="Vietnam" pageId="28" pageNumber="129">Vietnam</collectingCountry>
]
<location id="8EB7602CAE41B62CDEE64035FAA605E3" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CDEE64035FAA605E3" box="[1007,1405,1914,1939]" country="Vietnam" latitude="21.816668" longLatPrecision="1262" longitude="105.21667" municipality="Tonkin" name="Central Environs de Tuyen-quan" pageId="28" pageNumber="129">Central Environs de Tuyen-quan</location>
[g],
<geoCoordinate id="EE5C5030AE41B62CDD9E40D0FF2D05C7" box="[151,246,1951,1975]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="925" value="21.816668">21°49'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDC0F40D1FEB405C7" box="[262,367,1950,1975]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="925" value="105.21667">105°13'E</geoCoordinate>
, on vii–
<collectingDate id="EF92E9DFAE41B62CDCDD40D1FDF205C7" box="[468,553,1950,1975]" bridgedPair="/" pageId="28" pageNumber="129" value="1901-09">
<date id="FFD61037AE41B62CDCDD40D1FDF205C7" box="[468,553,1950,1975]" bridgedPair="/" pageId="28" pageNumber="129" value="1901-09">ix/1901</date>
</collectingDate>
(
<collectorName id="269D5321AE41B62CDF3540D0FD7C05C7" box="[572,679,1950,1975]" pageId="28" pageNumber="129">A. Weiss</collectorName>
) (
<collectionCode id="ED79AE32AE41B62CDFCB40D0FCC505C7" box="[706,798,1951,1975]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="129">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE41B62DDE3C40D1FDAA02C0" ID-GBIF-Occurrence="1658411484" collectingDate="2008-06-19" collectingDateMax="2008-06-20" collectingDateMin="2008-06-19" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; J. Pedersen" country="Laos" elevation="180" lastPageId="29" lastPageNumber="130" latitude="18.09815" location="Houayang National Park" longLatPrecision="1" longitude="102.67535" pageId="28" pageNumber="129" specimenCount="16" specimenCount-male="16" stateProvince="Vientiane" typeStatus="paratype">
<collectingCountry id="F37F7667AE41B62CDE3C40D1FCB405C7" box="[821,879,1950,1975]" name="Laos" pageId="28" pageNumber="129">
<emphasis id="B91CEAE5AE41B62CDE3C40D1FCB405C7" bold="true" box="[821,879,1950,1975]" pageId="28" pageNumber="129">Laos</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE41B62CDE8B40D1FBBE05C7" bold="true" box="[898,1125,1950,1975]" pageId="28" pageNumber="129">
<specimenCount id="9D6EFD7EAE41B62CDE8B40D1FC3F05C7" box="[898,996,1950,1975]" pageId="28" pageNumber="129" type="male">16 male</specimenCount>
<typeStatus id="54D38855AE41B62CDEE640D1FBBE05C7" box="[1007,1125,1950,1975]" pageId="28" pageNumber="129" type="paratype">paratypes</typeStatus>
</emphasis>
,
<location id="8EB7602CAE41B62CD97E40D0FA4305C7" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE41B62CD97E40D0FA4305C7" box="[1143,1432,1950,1975]" country="Laos" latitude="18.09815" longLatPrecision="1" longitude="102.67535" name="Houayang National Park" pageId="28" pageNumber="129" stateProvince="Vientiane">Houayang National Park</location>
, approximately
<locationDeviation id="9A652FD5AE41B62CDC48408DFDE705AB" box="[321,572,1986,2011]" pageId="28" pageNumber="129">
<quantity id="4C909B12AE41B62CDC48408DFE5105AB" box="[321,394,1986,2011]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="129" unit="km" value="15.0">15 km</quantity>
N of
<collectingRegion id="49ACF815AE41B62CDCC7408DFDE705AB" box="[462,572,1986,2011]" country="Laos" name="Viangchan" pageId="28" pageNumber="129">Vientiane</collectingRegion>
</locationDeviation>
,
<geoCoordinate id="EE5C5030AE41B62CDF40408DFD1005AB" box="[585,715,1986,2011]" direction="north" orientation="latitude" pageId="28" pageNumber="129" precision="1" value="18.09815">18°5.889'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE41B62CDFD1408DFCA805AB" box="[728,883,1986,2011]" direction="east" orientation="longitude" pageId="28" pageNumber="129" precision="1" value="102.67535">102°40.521'E</geoCoordinate>
,
<elevation id="0045D1C4AE41B62CDE89408CFC1105AA" box="[896,970,1987,2011]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="129" unit="m" value="180.0">
<quantity id="4C909B12AE41B62CDE89408CFC1105AA" box="[896,970,1987,2011]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="129" unit="m" value="180.0">180 m</quantity>
</elevation>
, secondary rain forest, on
<collectingDate id="EF92E9DFAE41B62CD9F0408DFA4605AB" box="[1273,1437,1986,2011]" pageId="28" pageNumber="129" value="2008-06-19" valueMax="2008-06-20" valueMin="2008-06-19">
<date id="FFD61037AE41B62CD9F0408DFA4605AB" box="[1273,1437,1986,2011]" pageId="28" pageNumber="129" value="2008-06-19" valueMax="2008-06-20" valueMin="2008-06-19">19–20/vi/2008</date>
</collectingDate>
(
<collectorName id="269D5321AE40B62DDDA947D7FEBF02C0" box="[160,356,151,176]" pageId="29" pageNumber="130">A. Solodovnikov</collectorName>
,
<collectorName id="269D5321AE40B62DDC7D47D7FE2C02C0" box="[372,503,151,176]" pageId="29" pageNumber="130">J. Pedersen</collectorName>
) (
<collectionCode id="ED79AE32AE40B62DDF1B47D7FDB202C0" box="[530,617,152,176]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="29" pageNumber="130">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B003CAAAE40B62DDF8947D8FB8B02C0" ID-GBIF-Occurrence="1658411488" box="[640,1104,151,176]" country="Vietnam" location="Hoa Binh" pageId="29" pageNumber="130" specimenCount="6" specimenCount-male="6" stateProvince="Hoa Binh" typeStatus="paratype">
<collectingCountry id="F37F7667AE40B62DDF8947D8FD3F02C0" box="[640,740,151,176]" name="Vietnam" pageId="29" pageNumber="130">
<emphasis id="B91CEAE5AE40B62DDF8947D8FD3F02C0" bold="true" box="[640,740,151,176]" pageId="29" pageNumber="130">Vietnam</emphasis>
</collectingCountry>
:
<emphasis id="B91CEAE5AE40B62DDFF147D8FC1702C0" bold="true" box="[760,972,151,176]" pageId="29" pageNumber="130">
<specimenCount id="9D6EFD7EAE40B62DDFF147D8FC9002C0" box="[760,843,151,176]" pageId="29" pageNumber="130" type="male">6 male</specimenCount>
<typeStatus id="54D38855AE40B62DDE5F47D8FC1702C0" box="[854,972,151,176]" pageId="29" pageNumber="130" type="paratype">paratypes</typeStatus>
</emphasis>
,
<collectingRegion id="49ACF815AE40B62DDED747D7FB8B02C0" box="[990,1104,151,176]" country="Vietnam" name="Hoa Binh" pageId="29" pageNumber="130">Hoa Binh</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B003CAAAE40B62DD96947D8FA8F02A4" ID-GBIF-Occurrence="1658411483" collectingDate="1923" collectingDateMax="1928" collectingDateMin="1923" collectionCode="MNHN" collectorName="A. De Cooman" country="Vietnam" latitude="20.828333" location="Tonkin" longLatPrecision="870" longitude="105.333336" pageId="29" pageNumber="130" specimenCount="1" typeStatus="paratype">
<location id="8EB7602CAE40B62DD96947D8FB7402C0" LSID="urn:lsid:plazi:treatment:03C187E1AE41B62DDD9E4620FA8302A4:8EB7602CAE40B62DD96947D8FB7402C0" box="[1120,1199,151,176]" country="Vietnam" latitude="20.828333" longLatPrecision="870" longitude="105.333336" name="Tonkin" pageId="29" pageNumber="130">Tonkin</location>
[northern
<collectingCountry id="F37F7667AE40B62DD82447D8FA4902C0" box="[1325,1426,151,176]" name="Vietnam" pageId="29" pageNumber="130">Vietnam</collectingCountry>
],
<geoCoordinate id="EE5C5030AE40B62DDD9E47F3FED002A4" box="[151,267,188,212]" direction="north" orientation="latitude" pageId="29" pageNumber="130" precision="92" value="20.828333">20°49.7'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE40B62DDC1F47F3FE5B02A5" box="[278,384,188,213]" direction="east" orientation="longitude" pageId="29" pageNumber="130" precision="925" value="105.333336">105°20'E</geoCoordinate>
, on [date specified only for two specimens:
<collectingDate id="EF92E9DFAE40B62DDE6647F3FBC302A4" box="[879,1048,188,213]" pageId="29" pageNumber="130" value="1923" valueMax="1928">
<date id="FFD61037AE40B62DDE6647F3FBC302A4" box="[879,1048,188,213]" pageId="29" pageNumber="130" value="1923" valueMax="1928">1923 and 1928</date>
</collectingDate>
] (
<collectorName id="269D5321AE40B62DD92747F3FB0102A4" box="[1070,1242,188,212]" pageId="29" pageNumber="130">A. De Cooman</collectorName>
) (
<collectionCode id="ED79AE32AE40B62DD9E647F3FA9002A4" box="[1263,1355,188,212]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="130">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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