167 lines
20 KiB
XML
167 lines
20 KiB
XML
<document id="82E0A32E37D8DF770D5960065D428EE9" ID-CLB-Dataset="102846" ID-DOI="10.11646/zootaxa.5166.1.1" ID-GBIF-Dataset="7e35a6a2-7ddc-453b-99ec-20142ba284f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6876209" ID-ZooBank="17952A48-902C-47A0-A344-8B07490F3B28" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658417435455" checkinUser="plazi" docAuthor="Cadena-Castañeda, Oscar J., Braun, Holger & García, Alexander García" docDate="2022" docId="03C0C519CF23614ED4A28C3B49D5FB5A" docLanguage="en" docName="zootaxa.5166.1.1.pdf" docOrigin="Zootaxa 5166 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5166.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lichenodraculus Braun 2011" docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="FFF9BD61CF3D6151D43589634C49FF9E" masterDocTitle="The tribe Dysoniini part VI: Phylogeny, biogeography and evolutionary trends of the lichen katydid genera (Orthoptera: Tettigoniidae: Phaneropterinae). Eleventh contribution to the suprageneric organization of Neotropical phaneropterines" masterLastPageNumber="93" masterPageNumber="1" pageNumber="31" updateTime="1699361192069" updateUser="ExternalLinkService">
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<mods:title id="3292A8FA498EBCD961E0DBCD161FD2A8">The tribe Dysoniini part VI: Phylogeny, biogeography and evolutionary trends of the lichen katydid genera (Orthoptera: Tettigoniidae: Phaneropterinae). Eleventh contribution to the suprageneric organization of Neotropical phaneropterines</mods:title>
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<mods:namePart id="301F6F881CC5A3718852BA635A6ACF40">Cadena-Castañeda, Oscar J.</mods:namePart>
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<mods:nameIdentifier id="6F85F59C1341E42BF5CB69C02E07B5D2" type="email">ojccorthoptera@gmail.com</mods:nameIdentifier>
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<mods:namePart id="97DF76BF375AA9EF1C471683811A0FF6">Braun, Holger</mods:namePart>
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<mods:nameIdentifier id="AD8FBBC64C07924FA88F2B07A036B611" type="email">braun@fcnym.unlp.edu.ar</mods:nameIdentifier>
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<mods:namePart id="8F6D57A891848B7421ECD9E0F8092D75">García, Alexander García</mods:namePart>
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<mods:title id="9B86E0A9DB1A7F939CEE1A0A4385E60C">Zootaxa</mods:title>
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<mods:date id="174D1E605DBF7D870EBC9A59296B6BD5">2022</mods:date>
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<mods:number id="419D6B85C3E654D2746D6C1D03CEC3EC">2022-07-19</mods:number>
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<treatment id="03C0C519CF23614ED4A28C3B49D5FB5A" ID-DOI="http://doi.org/10.5281/zenodo.6885780" ID-GBIF-Taxon="197841201" ID-Zenodo-Dep="6885780" LSID="urn:lsid:plazi:treatment:03C0C519CF23614ED4A28C3B49D5FB5A" httpUri="http://treatment.plazi.org/id/03C0C519CF23614ED4A28C3B49D5FB5A" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<subSubSection id="C3732784CF23614FD4A28C3B4E4AFAEC" box="[151,515,1368,1395]" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="8BD6740FCF23614FD4A28C3B4E4AFAEC" blockId="30.[151,515,1368,1395]" box="[151,515,1368,1395]" pageId="30" pageNumber="31">
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<heading id="D09EC363CF23614FD4A28C3B4E4AFAEC" bold="true" box="[151,515,1368,1395]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
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<taxonomicName id="4C690F8CCF23614FD4A28C3B4E4AFAEC" ID-CoL="8NTK9" authority="Braun, 2011" authorityName="Braun" authorityYear="2011" box="[151,515,1368,1395]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="B91DA81DCF23614FD4A28C3B4E4AFAEC" bold="true" box="[151,515,1368,1395]" pageId="30" pageNumber="31">
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<emphasis id="B91DA81DCF23614FD4A28C3B4D2DFAED" bold="true" box="[151,356,1368,1395]" italics="true" pageId="30" pageNumber="31">Lichenodraculus</emphasis>
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<bibRefCitation id="EFF809FECF23614FD5598C3A4E4AFAEC" author="Braun, H." box="[364,515,1368,1395]" pageId="30" pageNumber="31" pagination="33 - 39" refId="ref38372" refString="Braun, H. (2011) The Little Lichen Dragon - an extraordinary katydid from the Ecuadorian Andes (Orthoptera, Tettigoniidae, Phaneropterinae, Dysoniini). Zootaxa, 3032 (1), 33 - 39. https: // doi. org / 10.11646 / zootaxa. 3032.1.3" type="journal article" year="2011">Braun, 2011</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3732784CF23614FD4A28CC348B6F9D4" pageId="30" pageNumber="31" type="diagnosis">
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<paragraph id="8BD6740FCF23614FD4A28CC348B6F9D4" blockId="30.[151,1437,1440,1683]" pageId="30" pageNumber="31">
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<emphasis id="B91DA81DCF23614FD4A28CC34D5BFA24" bold="true" box="[151,274,1440,1466]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
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Vertex pointed and conical in lateral view. Pronotum with a prominent pointed process on anterior margin (
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<figureCitation id="1352688ACF23614FD4FE8CA74D29FA41" box="[203,352,1476,1503]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="30" pageNumber="31">Figs. 19A–B</figureCitation>
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), sometimes on posterior margin as well (
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<figureCitation id="1352688ACF23614FD7028CA74F82FA41" box="[823,971,1476,1503]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="30" pageNumber="31">Figs. 19D–E</figureCitation>
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). Hind femora and tibiae with cylindrical spines, elongated and curving, all genicular lobes developed as long spines. Male cerci elongate and sickle-shaped or the shape resembling tweezers, internally excavated (
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<figureCitation id="1352688ACF23614FD7268F6F4FECF9B9" box="[787,933,1548,1575]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="30" pageNumber="31">Figs. 19C, F</figureCitation>
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). Ovipositor broad, strongly curved upward, margins parallel and broadly rounded, apex delicately serrate. Female cerci are slender and elongate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3732784CF23614FD4F28F3648FFF9F0" box="[199,1206,1620,1647]" pageId="30" pageNumber="31" type="type_taxon">
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||
<paragraph id="8BD6740FCF23614FD4F28F3648FFF9F0" blockId="30.[151,1437,1440,1683]" box="[199,1206,1620,1647]" pageId="30" pageNumber="31">
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||
<emphasis id="B91DA81DCF23614FD4F28F364D2BF9F0" bold="true" box="[199,354,1620,1647]" pageId="30" pageNumber="31">
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||
<typeStatus id="54D2CAADCF23614FD4F28F364D4BF9F1" box="[199,258,1621,1647]" pageId="30" pageNumber="31">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C690F8CCF23614FD55C8F374EB0F9F1" ID-CoL="6Q7ZN" authority="Braun, 2011" authorityName="Braun" authorityYear="2011" box="[361,761,1620,1647]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="matti">
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<emphasis id="B91DA81DCF23614FD55C8F374E2FF9F0" box="[361,614,1620,1646]" italics="true" pageId="30" pageNumber="31">Lichenodraculus matti</emphasis>
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<bibRefCitation id="EFF809FECF23614FD6588F364EB0F9F1" author="Braun, H." box="[621,761,1621,1647]" pageId="30" pageNumber="31" pagination="33 - 39" refId="ref38372" refString="Braun, H. (2011) The Little Lichen Dragon - an extraordinary katydid from the Ecuadorian Andes (Orthoptera, Tettigoniidae, Phaneropterinae, Dysoniini). Zootaxa, 3032 (1), 33 - 39. https: // doi. org / 10.11646 / zootaxa. 3032.1.3" type="journal article" year="2011">Braun, 2011</bibRefCitation>
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</taxonomicName>
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, by monotypy and original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3732784CF23614FD4F28F1B4840F90D" box="[199,1033,1656,1683]" pageId="30" pageNumber="31" type="distribution">
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<paragraph id="8BD6740FCF23614FD4F28F1B4840F90D" blockId="30.[151,1437,1440,1683]" box="[199,1033,1656,1683]" pageId="30" pageNumber="31">
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<emphasis id="B91DA81DCF23614FD4F28F1B4D28F90C" bold="true" box="[199,353,1656,1682]" pageId="30" pageNumber="31">Distribution.</emphasis>
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Ecuadorian Andes and Guiana Shield in
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<collectingCountry id="F37E349FCF23614FD7188F1B4FE9F90C" box="[813,928,1656,1682]" name="Venezuela" pageId="30" pageNumber="31">Venezuela</collectingCountry>
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(
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<figureCitation id="1352688ACF23614FD79A8F1A4FB5F90D" box="[943,1020,1657,1683]" captionStart="Map 4" captionStartId="31.[151,202,848,872]" captionTargetBox="[151,1436,181,824]" captionTargetId="figure-22@31.[151,1436,181,824]" captionTargetPageId="31" captionText="Map 4. Distribution map of Apolinaria hygracantha and Lichenodraculus species." figureDoi="http://doi.org/10.5281/zenodo.6885564" httpUri="https://zenodo.org/record/6885564/files/figure.png" pageId="30" pageNumber="31">Map 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF162487CF22614ED4A28A334F97FCF6" ID-DOI="http://doi.org/10.5281/zenodo.6885564" ID-Zenodo-Dep="6885564" box="[151,990,848,872]" httpUri="https://zenodo.org/record/6885564/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,202,848,872]" targetBox="[151,1436,181,824]" targetPageId="31">
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<paragraph id="8BD6740FCF22614ED4A28A334F97FCF6" blockId="31.[151,990,848,872]" box="[151,990,848,872]" pageId="31" pageNumber="32">
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<emphasis id="B91DA81DCF22614ED4A28A334CAAFCF6" bold="true" box="[151,227,848,872]" pageId="31" pageNumber="32">Map 4.</emphasis>
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Distribution map of
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<taxonomicName id="4C690F8CCF22614ED58D8A334EE5FCF6" baseAuthorityName="Karsch" baseAuthorityYear="1896" box="[440,684,848,872]" class="Insecta" family="Tettigoniidae" genus="Apolinaria" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="hygracantha">
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<emphasis id="B91DA81DCF22614ED58D8A334EE5FCF6" box="[440,684,848,872]" italics="true" pageId="31" pageNumber="32">Apolinaria hygracantha</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C690F8CCF22614ED6E88A334F97FCF6" box="[733,990,848,872]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B91DA81DCF22614ED6E88A334FCEFCF6" box="[733,903,848,872]" italics="true" pageId="31" pageNumber="32">Lichenodraculus</emphasis>
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species.
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</taxonomicName>
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</paragraph>
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</caption>
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<subSubSection id="C3732784CF22614ED4A28AD949D5FB5A" pageId="31" pageNumber="32" type="key">
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||
<paragraph id="8BD6740FCF22614ED4A28AD94E72FC4A" blockId="31.[151,571,953,980]" box="[151,571,953,980]" pageId="31" pageNumber="32">
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<emphasis id="B91DA81DCF22614ED4A28AD94E72FC4A" bold="true" box="[151,571,953,980]" pageId="31" pageNumber="32">
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Key to species of
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<taxonomicName id="4C690F8CCF22614ED55B8ADA4E72FC4A" ID-CoL="8NTK9" authorityName="Braun" authorityYear="2011" box="[366,571,953,980]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B91DA81DCF22614ED55B8ADA4E72FC4A" bold="true" box="[366,571,953,980]" italics="true" pageId="31" pageNumber="32">Lichenodraculus</emphasis>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<keyStep id="309D6CAACF22614ED4A28A9C49D5FB5A" pageId="31" pageNumber="32">
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<key id="FFF8233ECF22614ED4A28A9C49D5FB5A" pageId="31" pageNumber="32">
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<paragraph id="8BD6740FCF22614ED4A28A9C49D4FBD1" blockId="31.[151,1437,1023,1221]" pageId="31" pageNumber="32">
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<keyLead id="3098D73ACF22614ED4A28A9C49D4FBD1" pageId="31" pageNumber="32">
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1. Only prozonal spine of the pronotum developed and as long as the vertex (
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<figureCitation id="1352688ACF22614ED79F8A9C486EFB8A" box="[938,1063,1023,1045]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="31" pageNumber="32">Figs. 19A, B</figureCitation>
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); male cerci with inner branch reduced (
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<figureCitation id="1352688ACF22614ED4DC8D7F4D08FBAF" box="[233,321,1052,1074]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="31" pageNumber="32">Fig. 19C</figureCitation>
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); ovipositor of female robust, with the dorsal and ventral margins slightly denticulate from the middle to the apex....................................................................................
|
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<taxonomicName id="4C690F8CCF22614ED0F88D5949D0FBD1" ID-CoL="6Q7ZN" authority="Braun, 2011" authorityName="Braun" authorityYear="2011" box="[1229,1433,1081,1103]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="matti">
|
||
<emphasis id="B91DA81DCF22614ED0F88D594952FBD0" bold="true" box="[1229,1307,1081,1103]" italics="true" pageId="31" pageNumber="32">L. matti</emphasis>
|
||
<bibRefCitation id="EFF809FECF22614ED1158D5949D0FBD1" author="Braun, H." box="[1312,1433,1082,1103]" pageId="31" pageNumber="32" pagination="33 - 39" refId="ref38372" refString="Braun, H. (2011) The Little Lichen Dragon - an extraordinary katydid from the Ecuadorian Andes (Orthoptera, Tettigoniidae, Phaneropterinae, Dysoniini). Zootaxa, 3032 (1), 33 - 39. https: // doi. org / 10.11646 / zootaxa. 3032.1.3" type="journal article" year="2011">Braun, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD6740FCF22614ED4A28D3449D5FB5A" blockId="31.[151,1437,1023,1221]" pageId="31" pageNumber="32">
|
||
<keyLead id="3098D73ACF22614ED4A28D3449D5FB5A" pageId="31" pageNumber="32">
|
||
- Both prozonal and metazonal spine of the pronotum developed, of similar size and not longer than the vertex in male, in female the pronotal spines are almost as long as the vertex (
|
||
<figureCitation id="1352688ACF22614ED6EC8D174F07FB17" box="[729,846,1140,1162]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="31" pageNumber="32">Fig. 19D, E</figureCitation>
|
||
); male cerci with inner branch almost as long as outer branch and with sclerotized margin (
|
||
<figureCitation id="1352688ACF22614ED5CF8DF14E06FB39" box="[506,591,1170,1192]" captionStart="FIGURE 19" captionStartId="33.[151,250,1880,1904]" captionTargetBox="[151,1421,182,1855]" captionTargetId="figure-22@33.[151,1436,181,1855]" captionTargetPageId="33" captionText="FIGURE 19. Lichenodraculus species. A–C. L. matti and D–F. L. holgeri. A, D. Male holotype in lateral view. B, E. Head and pronotum. C, F. Cerci in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6876259" httpUri="https://zenodo.org/record/6876259/files/figure.png" pageId="31" pageNumber="32">Fig. 19F</figureCitation>
|
||
); ovipositor slender, dorsal and ventral margins crenulated from middle to apex.............................................................................
|
||
<taxonomicName id="4C690F8CCF22614ED0738DCC49D5FB5A" ID-CoL="6PVZN" authority="Cadena-Castaneda, 2013" authorityName="Cadena-Castaneda" authorityYear="2013" box="[1094,1436,1198,1221]" class="Insecta" family="Tettigoniidae" genus="Lichenodraculus" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="holgeri">
|
||
<emphasis id="B91DA81DCF22614ED0738DCC48E1FB5A" bold="true" box="[1094,1192,1198,1220]" italics="true" pageId="31" pageNumber="32">L. holgeri</emphasis>
|
||
Cadena-Castañeda, 2013
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |