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<document id="1B3FFD80EAA078FF0CD89709F979288C" ID-DOI="10.5281/zenodo.180873" ID-GBIF-Dataset="45a1d960-62bf-4451-8fef-7a43b9269f0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="180873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459818278956" checkinUser="plazi" docAuthor="González, Guillermo, Corrêa, Geovan Henrique &amp; Almeida, Lúcia Massutti De" docDate="2008" docId="03C08152FFE3FFA92F9D5FFCFB3EF481" docLanguage="en" docName="zt01704p046.pdf" docOrigin="Zootaxa 1704" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpasus quadrifolium Gonzalez, Correa &amp;Almeida, sp. nov." docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="FFF9F92AFFE1FFAD2F0A5F21FF93F753" masterDocTitle="A new species of Harpasus Mulsant (Coleoptera, Coccinellidae, Chilocorinae) from Peru" masterLastPageNumber="46" masterPageNumber="42" pageNumber="44" updateTime="1698221629041" updateUser="plazi">
<mods:mods id="A24999CEA714198DE025ACD62DC8273F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5D63C238C08E1040B49EAC4C3613D25E">
<mods:title id="7F16D7D809C98F0ABA0DB053CE02030A">A new species of Harpasus Mulsant (Coleoptera, Coccinellidae, Chilocorinae) from Peru</mods:title>
</mods:titleInfo>
<mods:name id="19461CD536E8AB0599E7698807447490" type="personal">
<mods:role id="3031C1B3280662538654CEF352014099">
<mods:roleTerm id="E3B4DA47E1B37F79C047E85596107811">Author</mods:roleTerm>
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<mods:namePart id="97A63B3438B8E441F109ADF50EE5A302">González, Guillermo</mods:namePart>
</mods:name>
<mods:name id="856EC8F6B9E62896090AC61F139037F0" type="personal">
<mods:role id="8A2711D601C3E3EA12DDFCCF1349E96D">
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<mods:namePart id="5A1373D0F8443EDF777C00DC82AECC12">Corrêa, Geovan Henrique</mods:namePart>
</mods:name>
<mods:name id="F5CB45AAF878054D094DB937F967C019" type="personal">
<mods:role id="68CA90EA41533ACFBB88A30BE49C74DB">
<mods:roleTerm id="06637342797D9FBFA97C0B6A306E9BC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB585E18982FC989CF1B888FBC74A4AB">Almeida, Lúcia Massutti De</mods:namePart>
</mods:name>
<mods:typeOfResource id="BACAE6B1FAD3FCD715BB5A0AC872815A">text</mods:typeOfResource>
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<mods:titleInfo id="5FF5638101060B4E785D418807C27F95">
<mods:title id="2DD547545E4B0C117DE0C9243A99EFDA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3890F713766D8792CA5A951A638293AE">
<mods:date id="76D4224BFACB89C9B58908898741D445">2008</mods:date>
<mods:detail id="49AFBAC987C93407C83D10E53F4E08C8" type="volume">
<mods:number id="666CB5B9E223AEE14AC3E272931D64D1">1704</mods:number>
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<mods:start id="8C859D4677F357C5433E938860E204C1">42</mods:start>
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<mods:classification id="B699A17A85A697545F7608458CDEEF64">journal article</mods:classification>
<mods:identifier id="DCACC95637FAB257E206DBED3B046946" type="DOI">10.5281/zenodo.180873</mods:identifier>
<mods:identifier id="47AAAAB3F2C6FFA349483EC32DB971DE" type="GBIF-Dataset">45a1d960-62bf-4451-8fef-7a43b9269f0d</mods:identifier>
<mods:identifier id="614679DA3D66FDC3A94FDFCC70748E80" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BB8E9EDF8ACFDD1ECF1BFB8EA5E8598B" type="Zenodo-Dep">180873</mods:identifier>
</mods:mods>
<treatment id="03C08152FFE3FFA92F9D5FFCFB3EF481" ID-DOI="http://doi.org/10.5281/zenodo.6229949" ID-GBIF-Taxon="119355180" ID-Zenodo-Dep="6229949" LSID="urn:lsid:plazi:treatment:03C08152FFE3FFA92F9D5FFCFB3EF481" httpUri="http://treatment.plazi.org/id/03C08152FFE3FFA92F9D5FFCFB3EF481" lastPageId="4" lastPageNumber="46" pageId="2" pageNumber="44">
<subSubSection id="C37363CFFFE3FFAF2F9D5FFCFE81F649" pageId="2" pageNumber="44" type="nomenclature">
<paragraph id="8BD63044FFE3FFAF2F9D5FFCFC17F7A4" blockId="2.[151,900,221,282]" box="[151,900,221,247]" pageId="2" pageNumber="44">
<heading id="D09E8728FFE3FFAF2F9D5FFCFC17F7A4" bold="true" box="[151,900,221,247]" fontSize="11" level="1" pageId="2" pageNumber="44" reason="1">
<emphasis id="B91DEC56FFE3FFAF2F9D5FFCFC17F7A4" bold="true" box="[151,900,221,247]" pageId="2" pageNumber="44">
<taxonomicName id="4C694BC7FFE3FFAF2F9D5FFCFC8CF7A4" authority="Gonzalez, Correa &amp;Almeida" authorityName="Gonzalez, Correa &amp;Almeida" box="[151,799,221,247]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="quadrifolium" status="sp. nov.">
<emphasis id="B91DEC56FFE3FFAF2F9D5FFCFE20F7A4" bold="true" box="[151,435,221,247]" italics="true" pageId="2" pageNumber="44">Harpasus quadrifolium</emphasis>
González, Corrêa &amp;Almeida
</taxonomicName>
<taxonomicNameLabel id="A22E512DFFE3FFAF2C2D5FFCFC17F7A4" box="[807,900,221,247]" pageId="2" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD63044FFE3FFAF2F9D5E21FE81F649" blockId="2.[151,900,221,282]" box="[151,274,256,282]" pageId="2" pageNumber="44">
<figureCitation id="13522CC1FFE3FFAF2F9D5E21FE81F649" box="[151,274,256,282]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 11" captionStart-2="FIGURES 12 – 15" captionStartId-0="1.[151,269,1859,1883]" captionStartId-1="2.[151,269,1693,1717]" captionStartId-2="3.[151,269,1744,1768]" captionTargetBox-0="[287,1291,645,1827]" captionTargetBox-1="[273,1313,453,1680]" captionTargetBox-2="[425,1162,425,1733]" captionTargetId-0="figure@1.[264,1323,637,1833]" captionTargetId-1="figure@2.[264,1323,446,1685]" captionTargetId-2="figure@3.[415,1171,424,1736]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." captionText-1="FIGURES 7 – 11. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho." captionText-2="FIGURES 12 – 15. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli." httpUri-0="https://zenodo.org/record/180874/files/figure.png" httpUri-1="https://zenodo.org/record/180875/files/figure.png" httpUri-2="https://zenodo.org/record/180876/files/figure.png" pageId="2" pageNumber="44">Figs. 1–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37363CFFFE3FFAF2F9D5E6EFB2FF6C1" pageId="2" pageNumber="44" type="diagnosis">
<paragraph id="8BD63044FFE3FFAF2F9D5E6EFB2FF6C1" blockId="2.[151,1436,335,402]" pageId="2" pageNumber="44">
<emphasis id="B91DEC56FFE3FFAF2F9D5E6EFE84F63A" bold="true" box="[151,279,335,361]" pageId="2" pageNumber="44">Diagnosis.</emphasis>
Oblong semicircular, black with the lateral and frontal border of pronotum orange brown. Elytra with lateral border and four orange brown maculae pattern, appearing as a four leaf clover.
</paragraph>
</subSubSection>
<caption id="DF1660CCFFE3FFAF2F9D59BCFCDEF184" httpUri="https://zenodo.org/record/180875/files/figure.png" pageId="2" pageNumber="44" targetBox="[273,1313,453,1680]" targetPageId="2">
<paragraph id="8BD63044FFE3FFAF2F9D59BCFCDEF184" blockId="2.[151,1436,1693,1751]" pageId="2" pageNumber="44">
<emphasis id="B91DEC56FFE3FFAF2F9D59BCFEDBF1E6" bold="true" box="[151,328,1693,1717]" pageId="2" pageNumber="44">FIGURES 7–11</emphasis>
.
<taxonomicName id="4C694BC7FFE3FFAF2E5F59BFFC1DF1E6" authority="Gonzalez, Correa &amp; Almeida" authorityName="Gonzalez, Correa &amp; Almeida" box="[341,910,1694,1717]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="quadrifolium" status="sp. nov.">
<emphasis id="B91DEC56FFE3FFAF2E5F59BFFDDDF1E6" box="[341,590,1694,1717]" italics="true" pageId="2" pageNumber="44">Harpasus quadrifolium</emphasis>
González, Corrêa &amp; Almeida
</taxonomicName>
<emphasis id="B91DEC56FFE3FFAF2C9E59BCFC70F1E6" bold="true" box="[916,995,1693,1717]" pageId="2" pageNumber="44">
<taxonomicNameLabel id="A22E512DFFE3FFAF2C9E59BCFC70F1E6" box="[916,995,1693,1717]" pageId="2" pageNumber="44" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho.
</paragraph>
</caption>
<subSubSection id="C37363CFFFE3FFA92FCC5824FC33F549" lastPageId="4" lastPageNumber="46" pageId="2" pageNumber="44" type="materials_examined">
<paragraph id="8BD63044FFE3FFAE2FCC5824FF56F629" blockId="2.[151,1437,1797,2024]" lastBlockId="3.[151,1437,152,378]" lastPageId="3" lastPageNumber="45" pageId="2" pageNumber="44">
<emphasis id="B91DEC56FFE3FFAF2FCC5824FED3F04C" bold="true" box="[198,320,1797,1823]" pageId="2" pageNumber="44">
<typeStatus id="54D28EE6FFE3FFAF2FCC5824FEA8F04C" box="[198,315,1797,1823]" pageId="2" pageNumber="44" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Length: 2.80–3.00 mm. Width: 2.50–2.70 mm. Body rounded, convex, punctuation uniformly distributed on pronotum and elytra (
<figureCitation id="13522CC1FFE3FFAF2D9E580FFD4AF01B" box="[660,729,1838,1864]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 1</figureCitation>
). Head dark brown (
<figureCitation id="13522CC1FFE3FFAF2CC7580FFB87F01B" box="[973,1044,1838,1864]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 5</figureCitation>
), antenna (fig. 8) and mouthparts light or dark brown (
<figureCitation id="13522CC1FFE3FFAF2E805877FE41F023" box="[394,466,1878,1904]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 2</figureCitation>
). Pronotum black, with antero-lateral margins light brown (
<figureCitation id="13522CC1FFE3FFAF2B865877FA9CF023" box="[1164,1295,1878,1904]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 1, 3, 5</figureCitation>
). Scutellum black (
<figureCitation id="13522CC1FFE3FFAF2FEC585FFEDFF0CB" box="[230,332,1918,1944]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 1, 5</figureCitation>
). Elytra black with raised humeral callosity, lateral margin slightly reflexed, light brown (
<figureCitation id="13522CC1FFE3FFAF2A66585FFF71F093" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="2" pageNumber="44">Fig. 1, 3, 5</figureCitation>
). Each elytron with two light brown maculae: the first, subtriangular close to suture and base of elytron with an emargination above the humeral callus; the second, slightly smaller than first and subtriangular, near the suture and external margin of elytron (
<figureCitation id="13522CC1FFE2FFAE2D985FB9FCAAF7E1" box="[658,825,152,178]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="3" pageNumber="45">Fig. 1, 3, 4, 5</figureCitation>
). Epipleura and abdominal sterna light brown, epipleura glabrous and abdominal sternites with small setae and small punctures. The center of pro, meso and metathorax dark brown (
<figureCitation id="13522CC1FFE2FFAE2EBD5FC9FE6CF651" box="[439,511,232,258]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="3" pageNumber="45">Fig. 2</figureCitation>
). Legs dark brown, robust. Abdomen with six visible sternites in male, the fifth a little larger than the second, third or fourth with posterior margin emarginate, the sixth sternite small (
<figureCitation id="13522CC1FFE2FFAE2A665E31FF3BF601" captionStart="FIGURES 7 – 11" captionStartId="2.[151,269,1693,1717]" captionTargetBox="[273,1313,453,1680]" captionTargetId="figure@2.[264,1323,446,1685]" captionTargetPageId="2" captionText="FIGURES 7 – 11. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho." httpUri="https://zenodo.org/record/180875/files/figure.png" pageId="3" pageNumber="45">Fig. 7</figureCitation>
); five sternites in female, fifth is larger than the second, third or fourth, with posterior margin rounded (
<figureCitation id="13522CC1FFE2FFAE2A675E19FF2BF629" captionStart="FIGURES 12 – 15" captionStartId="3.[151,269,1744,1768]" captionTargetBox="[425,1162,425,1733]" captionTargetId="figure@3.[415,1171,424,1736]" captionTargetPageId="3" captionText="FIGURES 12 – 15. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli." httpUri="https://zenodo.org/record/180876/files/figure.png" pageId="3" pageNumber="45">Fig. 12</figureCitation>
).
</paragraph>
<caption id="DF1660CCFFE2FFAE2F9D59F1FDCEF059" httpUri="https://zenodo.org/record/180876/files/figure.png" pageId="3" pageNumber="45" targetBox="[425,1162,425,1733]" targetPageId="3">
<paragraph id="8BD63044FFE2FFAE2F9D59F1FDCEF059" blockId="3.[151,1436,1744,1802]" pageId="3" pageNumber="45">
<emphasis id="B91DEC56FFE2FFAE2F9D59F1FECDF1BB" bold="true" box="[151,350,1744,1768]" pageId="3" pageNumber="45">FIGURES 12–15.</emphasis>
<taxonomicName id="4C694BC7FFE2FFAE2E6E59F1FC0DF1B4" authority="Gonzalez, Correa &amp; Almeida" authorityName="Gonzalez, Correa &amp; Almeida" box="[356,926,1744,1767]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="45" phylum="Arthropoda" rank="species" species="quadrifolium" status="sp. nov.">
<emphasis id="B91DEC56FFE2FFAE2E6E59F1FDCDF1B4" box="[356,606,1744,1767]" italics="true" pageId="3" pageNumber="45">Harpasus quadrifolium</emphasis>
González, Corrêa &amp; Almeida
</taxonomicName>
<emphasis id="B91DEC56FFE2FFAE2CAF59F1FC60F1BB" bold="true" box="[933,1011,1744,1768]" pageId="3" pageNumber="45">
<taxonomicNameLabel id="A22E512DFFE2FFAE2CAF59F1FC60F1BB" box="[933,1011,1744,1768]" pageId="3" pageNumber="45" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli.
</paragraph>
</caption>
<paragraph id="8BD63044FFE2FFA92FCC581CFBFEF7E1" blockId="3.[151,1436,1853,1999]" lastBlockId="4.[151,1436,152,978]" lastPageId="4" lastPageNumber="46" pageId="3" pageNumber="45">
Male genitalia. Median lobe symmetrical, lanceolate, broad at basal third abruptly narrowing at apex, 1/3 shorter than paramere; trabes rather stout, a little longer than median lobe and a little shorter than paramere; paramere constricted at base and wider in apical 1/3, with dense and long pubescence at lateral sides and apex (
<figureCitation id="13522CC1FFE2FFAE2F955894FF7BF09C" box="[159,232,1973,1999]" captionStart="FIGURES 7 – 11" captionStartId="2.[151,269,1693,1717]" captionTargetBox="[273,1313,453,1680]" captionTargetId="figure@2.[264,1323,446,1685]" captionTargetPageId="2" captionText="FIGURES 7 – 11. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho." httpUri="https://zenodo.org/record/180875/files/figure.png" pageId="3" pageNumber="45">Fig. 9</figureCitation>
). In lateral view paramere with apex strongly bent (
<figureCitation id="13522CC1FFE2FFAE2C405894FC31F09C" box="[842,930,1973,1999]" captionStart="FIGURES 7 – 11" captionStartId="2.[151,269,1693,1717]" captionTargetBox="[273,1313,453,1680]" captionTargetId="figure@2.[264,1323,446,1685]" captionTargetPageId="2" captionText="FIGURES 7 – 11. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho." httpUri="https://zenodo.org/record/180875/files/figure.png" pageId="3" pageNumber="45">Fig. 10</figureCitation>
). Sipho long, uniform diameter throughout most of length, strongly sclerotized, bent, with apex membranous, twisted (
<figureCitation id="13522CC1FFE5FFA92B035FB9FBCCF7E1" box="[1033,1119,152,178]" captionStart="FIGURES 7 – 11" captionStartId="2.[151,269,1693,1717]" captionTargetBox="[273,1313,453,1680]" captionTargetId="figure@2.[264,1323,446,1685]" captionTargetPageId="2" captionText="FIGURES 7 – 11. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 7, male abdomen; 8, antenna; male genitalia: 9, tegmen ventral view; 10, tegmen lateral view; 11, sipho." httpUri="https://zenodo.org/record/180875/files/figure.png" pageId="4" pageNumber="46">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BD63044FFE5FFA92FCC5FE1FD9DF679" blockId="4.[151,1436,152,978]" pageId="4" pageNumber="46">
Female genitalia. Coxites long, about 3.8 times longer than wide, subtriangular, slender, apex rounded; bursa copulatrix large (
<figureCitation id="13522CC1FFE5FFA92EA45FC9FD94F651" box="[430,519,232,258]" captionStart="FIGURES 12 – 15" captionStartId="3.[151,269,1744,1768]" captionTargetBox="[425,1162,425,1733]" captionTargetId="figure@3.[415,1171,424,1736]" captionTargetPageId="3" captionText="FIGURES 12 – 15. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli." httpUri="https://zenodo.org/record/180876/files/figure.png" pageId="4" pageNumber="46">Fig. 13</figureCitation>
); infundibulum long, cup-shaped (
<figureCitation id="13522CC1FFE5FFA92CA25FC9FB92F651" box="[936,1025,232,258]" captionStart="FIGURES 12 – 15" captionStartId="3.[151,269,1744,1768]" captionTargetBox="[425,1162,425,1733]" captionTargetId="figure@3.[415,1171,424,1736]" captionTargetPageId="3" captionText="FIGURES 12 – 15. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli." httpUri="https://zenodo.org/record/180876/files/figure.png" pageId="4" pageNumber="46">Fig. 14</figureCitation>
); styli small with three long setae (
<figureCitation id="13522CC1FFE5FFA92F955E31FF65F679" box="[159,246,272,298]" captionStart="FIGURES 12 – 15" captionStartId="3.[151,269,1744,1768]" captionTargetBox="[425,1162,425,1733]" captionTargetId="figure@3.[415,1171,424,1736]" captionTargetPageId="3" captionText="FIGURES 12 – 15. Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 12, female abdomen; female genitalia: 13, ovipositor; 14, infundibulum; 15, styli." httpUri="https://zenodo.org/record/180876/files/figure.png" pageId="4" pageNumber="46">Fig. 15</figureCitation>
). Spermatheca missing.
</paragraph>
<paragraph id="8BD63044FFE5FFA92FCF5E16FC33F549" blockId="4.[151,1436,152,978]" pageId="4" pageNumber="46">
<emphasis id="B91DEC56FFE5FFA92FCF5E16FEE9F602" bold="true" box="[197,378,311,337]" pageId="4" pageNumber="46">
<typeStatus id="54D28EE6FFE5FFA92FCF5E16FE93F602" box="[197,256,311,337]" pageId="4" pageNumber="46">Type</typeStatus>
Material
</emphasis>
.
<typeStatus id="54D28EE6FFE5FFA92E9B5E19FD90F601" box="[401,515,312,338]" pageId="4" pageNumber="46" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F37E70D4FFE5FFA92D195E19FDF1F601" box="[531,610,312,338]" name="Peru" pageId="4" pageNumber="46">PERU</collectingCountry>
. [Chanchamayo,
<date id="FFD71684FFE5FFA92C325E19FC3EF601" box="[824,941,312,338]" pageId="4" pageNumber="46" value="1986-01-07">07/I/1986</date>
, Arellano Coll./ UA
<date id="FFD71684FFE5FFA92BBD5E19FA8CF601" box="[1207,1311,312,338]" pageId="4" pageNumber="46" value="2005" valueMax="2086">2005-86</date>
], (
<figureCitation id="13522CC1FFE5FFA92A4A5E19FA1EF601" box="[1344,1421,312,338]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1859,1883]" captionTargetBox="[287,1291,645,1827]" captionTargetId="figure@1.[264,1323,637,1833]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Habitus of Harpasus quadrifolium González, Corrêa &amp; Almeida sp. nov.; 1, dorsal view; 2, ventral view; 3, lateral view; 4, posterior view; 5, frontal view; 6, specimen labels." httpUri="https://zenodo.org/record/180874/files/figure.png" pageId="4" pageNumber="46">Fig. 6</figureCitation>
), male, deposited at the Museo de Entomología Klaus Raven Büller, Universidad Nacional Agraria La Molina, Lima,
<collectingCountry id="F37E70D4FFE5FFA92FEE5EA9FE89F6F1" box="[228,282,392,418]" name="Peru" pageId="4" pageNumber="46">Perú</collectingCountry>
(
<collectionCode id="ED78A881FFE5FFA92E235EA9FE0FF6F1" box="[297,412,392,418]" pageId="4" pageNumber="46">UNALM</collectionCode>
).
<typeStatus id="54D28EE6FFE5FFA92EA45EA9FDB6F6F1" box="[430,549,392,418]" pageId="4" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F37E70D4FFE5FFA92D3A5EA9FDECF6F1" box="[560,639,392,418]" name="Peru" pageId="4" pageNumber="46">PERU</collectingCountry>
. [Chanchamayo,
<date id="FFD71684FFE5FFA92C405EA9FC2CF6F1" box="[842,959,392,418]" pageId="4" pageNumber="46" value="1986-01-07">07/I/1986</date>
, Arellano Coll./ UA
<date id="FFD71684FFE5FFA92BBF5EA9FA8FF6F1" box="[1205,1308,392,418]" pageId="4" pageNumber="46" value="2005" valueMax="2086">2005-86</date>
],
<specimenCount id="9D6FFBCDFFE5FFA92A255EA9FA0BF6F1" box="[1327,1432,392,418]" pageId="4" pageNumber="46" type="female">2 female</specimenCount>
, deposited at the Museo de Entomología Klaus Raven Büller, Universidad Nacional Agraria La Molina, Lima,
<collectingCountry id="F37E70D4FFE5FFA92F9D5EF9FF5EF6A1" box="[151,205,472,498]" name="Peru" pageId="4" pageNumber="46">Perú</collectingCountry>
(
<collectionCode id="ED78A881FFE5FFA92FD65EF9FEDCF6A1" box="[220,335,472,498]" pageId="4" pageNumber="46">UNALM</collectionCode>
); same data and locality,
<specimenCount id="9D6FFBCDFFE5FFA92D7F5EF9FD4DF6A1" box="[629,734,472,498]" pageId="4" pageNumber="46" type="female">1 female</specimenCount>
, deposited at the Coleção de Entomológica Pe. J. S. Moure, Universidade Federal do Paraná, Curitiba, Paraná,
<collectingCountry id="F37E70D4FFE5FFA92DE05D21FCA1F549" box="[746,818,512,538]" name="Brazil" pageId="4" pageNumber="46">Brazil</collectingCountry>
(
<collectionCode id="ED78A881FFE5FFA92C4B5D21FC07F549" box="[833,916,512,538]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="4" pageNumber="46">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37363CFFFE5FFA92FCF5D06FC8AF4F9" pageId="4" pageNumber="46" type="discussion">
<paragraph id="8BD63044FFE5FFA92FCF5D06FC8AF4F9" blockId="4.[151,1436,152,978]" pageId="4" pageNumber="46">
<emphasis id="B91DEC56FFE5FFA92FCF5D06FEADF512" bold="true" box="[197,318,551,577]" pageId="4" pageNumber="46">Remarks.</emphasis>
<taxonomicName id="4C694BC7FFE5FFA92E4D5D09FC2EF511" authority="Gonzalez, Correa &amp; Almeida" authorityName="Gonzalez, Correa &amp; Almeida" box="[327,957,552,578]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="quadrifolium" status="sp. nov.">
<emphasis id="B91DEC56FFE5FFA92E4D5D09FDC9F512" box="[327,602,552,577]" italics="true" pageId="4" pageNumber="46">Harpasus quadrifolium</emphasis>
González, Corrêa &amp; Almeida
</taxonomicName>
<emphasis id="B91DEC56FFE5FFA92CCF5D06FBB0F512" bold="true" box="[965,1059,551,577]" pageId="4" pageNumber="46">
<taxonomicNameLabel id="A22E512DFFE5FFA92CCF5D06FBB0F512" box="[965,1059,551,577]" pageId="4" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a pattern of maculae which is distinct from the other species.
<taxonomicName id="4C694BC7FFE5FFA92D2D5D71FC75F539" authority="Almeida &amp; Carvalho, 2006" authorityName="Almeida &amp; Carvalho" authorityYear="2006" box="[551,998,592,618]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="aureus">
<emphasis id="B91DEC56FFE5FFA92D2D5D71FD08F53A" box="[551,667,592,617]" italics="true" pageId="4" pageNumber="46">H. aureus</emphasis>
<bibRefCitation id="EFF84DB5FFE5FFA92DA95D71FC75F539" author="Almeida" box="[675,998,592,618]" pageId="4" pageNumber="46" refString="Almeida, L. M. &amp; R. C. Z. de Carvalho. (2006) A new Brazilian species of Harpasus Mulsant (Coleoptera, Coccinellidae, Chilocorinae), a predator of Tinocallis kahawaluokalani (Kirkaldy) (Hemiptera, Aphididae). Zootaxa, 1195, 31 - 37." type="journal article" year="2006">Almeida &amp; Carvalho, 2006</bibRefCitation>
</taxonomicName>
is unicolorous, bright golden brown, and yellowish at the lateral margin of the elytra. In
<taxonomicName id="4C694BC7FFE5FFA92DE05D59FB13F5C1" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[746,1152,632,658]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="zonatus">
<emphasis id="B91DEC56FFE5FFA92DE05D59FC2EF5C2" box="[746,957,632,657]" italics="true" pageId="4" pageNumber="46">Harpasus zonatus</emphasis>
(
<bibRefCitation id="EFF84DB5FFE5FFA92CC15D59FBE4F5C1" author="Mulsant" box="[971,1143,632,658]" pageId="4" pageNumber="46" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physique et Naturelles d'Agriculture et d'Industrie, 2, 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
)
</taxonomicName>
the integument is bright golden brown, with a dark brown stripe on sutural and lateral margin of each elytron; in
<taxonomicName id="4C694BC7FFE5FFA92BAF5D81FE8AF5B1" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" class="Insecta" family="Coccinellidae" genus="Orcus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="eversmanni">
<emphasis id="B91DEC56FFE5FFA92BAF5D81FAC3F5EA" box="[1189,1360,672,697]" italics="true" pageId="4" pageNumber="46">H. eversmanni</emphasis>
(
<bibRefCitation id="EFF84DB5FFE5FFA92A555D81FE82F5B1" author="Mulsant" pageId="4" pageNumber="46" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physique et Naturelles d'Agriculture et d'Industrie, 2, 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
)
</taxonomicName>
the integument is reddish at lateral margin with a blue-metallic area at the disc of each elytron; in
<emphasis id="B91DEC56FFE5FFA92F9D5DD1FEC0F45A" box="[151,339,752,777]" italics="true" pageId="4" pageNumber="46">
<taxonomicName id="4C694BC7FFE5FFA92F9D5DD1FEDCF45A" box="[151,335,752,777]" class="Insecta" family="Coccinellidae" genus="Orcus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="pallidilabris">H. pallidilabris</taxonomicName>
,
</emphasis>
the
<typeStatus id="54D28EE6FFE5FFA92E8C5DD1FE2BF459" box="[390,440,752,778]" pageId="4" pageNumber="46">type</typeStatus>
species of the genus, the integument is bluish-green, without any maculae or stripes. The male genitalia of the new species are similar to
<taxonomicName id="4C694BC7FFE5FFA92DFF5C39FCF4F462" box="[757,871,792,817]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="aureus">
<emphasis id="B91DEC56FFE5FFA92DFF5C39FCF4F462" box="[757,871,792,817]" italics="true" pageId="4" pageNumber="46">H. aureus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694BC7FFE5FFA92C955C39FB88F462" box="[927,1051,792,817]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="zonatus">
<emphasis id="B91DEC56FFE5FFA92C955C39FB88F462" box="[927,1051,792,817]" italics="true" pageId="4" pageNumber="46">H. zonatus</emphasis>
</taxonomicName>
except that the apex of the sipho is twisted in
<taxonomicName id="4C694BC7FFE5FFA92E245C61FE78F40A" box="[302,491,832,857]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="quadrifolium">
<emphasis id="B91DEC56FFE5FFA92E245C61FE78F40A" box="[302,491,832,857]" italics="true" pageId="4" pageNumber="46">H. quadrifolium</emphasis>
</taxonomicName>
and truncate in
<taxonomicName id="4C694BC7FFE5FFA92DA15C61FC8DF40A" box="[683,798,832,857]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="aureus">
<emphasis id="B91DEC56FFE5FFA92DA15C61FC8DF40A" box="[683,798,832,857]" italics="true" pageId="4" pageNumber="46">H. aureus</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C694BC7FFE5FFA92C475C61FC5FF40A" box="[845,972,832,857]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="zonatus">
<emphasis id="B91DEC56FFE5FFA92C475C61FC5FF40A" box="[845,972,832,857]" italics="true" pageId="4" pageNumber="46">H. zonatus</emphasis>
</taxonomicName>
the paramere is wider than in the other species. The female genitalia are similar to
<taxonomicName id="4C694BC7FFE5FFA92D905C49FC9DF4D2" box="[666,782,872,897]" class="Insecta" family="Coccinellidae" genus="Harpasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="aureus">
<emphasis id="B91DEC56FFE5FFA92D905C49FC9DF4D2" box="[666,782,872,897]" italics="true" pageId="4" pageNumber="46">H. aureus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694BC7FFE5FFA92C425C49FC67F4D2" box="[840,1012,872,897]" class="Insecta" family="Coccinellidae" genus="Orcus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="46" phylum="Arthropoda" rank="species" species="eversmanni">
<emphasis id="B91DEC56FFE5FFA92C425C49FC67F4D2" box="[840,1012,872,897]" italics="true" pageId="4" pageNumber="46">H. eversmanni</emphasis>
</taxonomicName>
, but differ in the shape of the bursa copulatrix, and in possessing longer and wider coxites.
</paragraph>
</subSubSection>
<subSubSection id="C37363CFFFE5FFA92FCF5C96FB3EF481" box="[197,1197,951,978]" pageId="4" pageNumber="46" type="etymology">
<paragraph id="8BD63044FFE5FFA92FCF5C96FB3EF481" blockId="4.[151,1436,152,978]" box="[197,1197,951,978]" pageId="4" pageNumber="46">
<emphasis id="B91DEC56FFE5FFA92FCF5C96FED9F482" bold="true" box="[197,330,951,977]" pageId="4" pageNumber="46">Etymology</emphasis>
. This species is named after the pattern of maculae like a four leaf clover.
</paragraph>
</subSubSection>
</treatment>
</document>
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