651 lines
79 KiB
XML
651 lines
79 KiB
XML
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<mods:title id="9D919B5A4ED4337F50520CFA0E614C81">A new Liopeltis Fitzinger, 1843 (Squamata: Colubridae) from Pulau Tioman, Peninsular Malaysia</mods:title>
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<taxonomicName id="4C1C97C4FFFDBB1E60F6F924819BF2EE" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[151,384,1711,1738]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="475" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
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<emphasis id="B9683055FFFDBB1E60F6F924819BF2EE" bold="true" box="[151,384,1711,1738]" italics="true" pageId="3" pageNumber="475">Liopeltis tiomanica</emphasis>
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<taxonomicName id="4C1C97C4FFFDBB1E60F6F9658158F322" box="[151,323,1774,1798]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="475" phylum="Chordata" rank="species" species="tricolor">
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<emphasis id="B9683055FFFDBB1E60F6F9658158F322" box="[151,323,1774,1798]" italics="true" pageId="3" pageNumber="475">Liopeltis tricolor</emphasis>
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(part): J.L.
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<bibRefCitation id="EF8D91B6FFFDBB1E61DDF96582ACF323" author="Grismer" box="[444,695,1774,1799]" etAl="et al." firstAuthor="Grismer" pageId="3" pageNumber="475" pagination="247 - 279" refId="ref5729" refString="Grismer, J. L., Grismer, L. L., Das, I., Yaakob, N. S., Liat, L. B., Leong, T. M., Youmans, T. M. & Kaiser, H. (2004) Species diversity and checklist of the herpetofauna of Pulau Tioman, Peninsular Malaysia, with a preliminary overview of habitat utilization. Asiatic Herpetological Research, 10, 247 - 279." type="journal article" year="2004">
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Grismer
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<emphasis id="B9683055FFFDBB1E6274F9648252F322" box="[533,585,1774,1798]" italics="true" pageId="3" pageNumber="475">et al.</emphasis>
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2004: 275
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;
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<treatmentCitation id="0ABDCA56FFFDBB1E62A3F9658398F323" author="Grismer, L. L." box="[706,899,1774,1799]" page="203" pageId="3" pageNumber="475" year="2011">
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<bibRefCitation id="EF8D91B6FFFDBB1E62A3F9658398F323" author="Grismer" box="[706,899,1774,1799]" firstAuthor="Grismer" pageId="3" pageNumber="475" refId="ref5915" refString="Grismer, L. L. (2011) Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago, Peninsular Malaysia. Edition Chimaira, Frankfurt, 258 pp." type="book" year="2011">Grismer 2011: 203</bibRefCitation>
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.
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<taxonomicName id="4C1C97C4FFFDBB1E60F6F886814BF301" box="[151,336,1805,1829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="475" phylum="Chordata" rank="species" species="tricolour">
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<emphasis id="B9683055FFFDBB1E60F6F886814BF301" box="[151,336,1805,1829]" italics="true" pageId="3" pageNumber="475">Liopeltis tricolour</emphasis>
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[
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<emphasis id="B9683055FFFDBB1E613EF8858160F301" box="[351,379,1806,1829]" italics="true" pageId="3" pageNumber="475">sic</emphasis>
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] (part):
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<treatmentCitation id="0ABDCA56FFFDBB1E61B1F88782D0F301" author="Grismer, L. L. & Youmans, T. M. & Wood, Jr. P. L. & Grismer, J. L." box="[464,715,1804,1829]" page="178" pageId="3" pageNumber="475" year="2006">
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<bibRefCitation id="EF8D91B6FFFDBB1E61B1F88782D0F301" author="Grismer" box="[464,715,1804,1829]" etAl="et al." firstAuthor="Grismer" pageId="3" pageNumber="475" pagination="157 - 180" refId="ref6168" refString="Grismer, L. L., Youmans, T. M., Wood, Jr. P. L. & Grismer, J. L. (2006) Checklist of the herpetofauna of the Seribuat archipelago, West Malaysia with comments on biogeography, natural history, and adaptive types. Raffles Bulletin of Zoology, 54 (1), 157 - 180." type="journal article" year="2006">
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Grismer
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<emphasis id="B9683055FFFDBB1E6248F8858247F301" box="[553,604,1805,1829]" italics="true" pageId="3" pageNumber="475">et al.</emphasis>
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2006: 178
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.
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<materialsCitation id="3B74E61AFFFDBB1E60F6F8C582C0F394" ID-GBIF-Occurrence="2608670273" collectingDate="2002-08-11" collectionCode="LSUHC" collectorName="Jesse L. Grismer & Perry L. Wood, Jr." country="Malaysia" county="Peninsular" elevation="618" latitude="2.779" location="Tioman Island" longLatPrecision="1" longitude="104.15703" municipality="Pulau Tioman" pageId="3" pageNumber="475" specimenCode="LSUHC 5037" specimenCount="1" stateProvince="Pahang State" typeStatus="holotype">
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<typeStatus id="54A752E5FFFDBB1E60F6F8C5811DF34C" box="[151,262,1870,1896]" pageId="3" pageNumber="475" type="holotype">Holotype</typeStatus>
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.
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<specimenCode id="DBBA443CFFFDBB1E6175F8C581AFF34C" box="[276,436,1869,1896]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="3" pageNumber="475">LSUHC 5037</specimenCode>
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,
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<specimenCount id="9D1A27CEFFFDBB1E61A0F8C58255F34C" box="[449,590,1870,1896]" pageId="3" pageNumber="475" type="adult">adult female</specimenCount>
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(
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<figureCitation id="1327F0C2FFFDBB1E623EF8C582D5F34C" box="[607,718,1870,1896]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1124,1149]" captionStartId-1="6.[151,250,1265,1290]" captionStartId-2="7.[151,250,1860,1885]" captionTargetBox-0="[151,1436,181,1099]" captionTargetBox-1="[158,1429,181,1240]" captionTargetBox-2="[192,1382,184,1824]" captionTargetId-0="figure@5.[151,1436,181,1099]" captionTargetId-1="figure@6.[158,1429,181,1240]" captionTargetId-2="figure@7.[192,1395,181,1835]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Holotype (LSUHC 5037) of Liopeltis tiomanica sp. nov. in life. Photograph by LLG." captionText-1="FIGURE 3. Holotype (LSUHC 5037) Liopeltis tiomanica sp. nov. in preservative. A. dorsal view of body. B. Ventral view of body. C. Lateral view of head. D. Dorsal view of head. E. Ventral view of head." captionText-2="FIGURE 4. Fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 1,152 aligned characters of the complete mitochondrial cytochrome b gene and parts of its flanking tRNA genes from colubrid snakes of the genera Liopeltis, Gongylosoma and Sibynophis. Numbers above nodes are Bayesian posterior probabilities. Numbers at terminal tips are GenBank accession numbers. Voucher and locality data for samples are provided in Table 2. The holotype of Liopeltis tiomanica sp. nov. in life is shown in the image." figureDoi-0="http://doi.org/10.5281/zenodo.3765617" figureDoi-1="http://doi.org/10.5281/zenodo.3765619" figureDoi-2="http://doi.org/10.5281/zenodo.3765621" httpUri-0="https://zenodo.org/record/3765617/files/figure.png" httpUri-1="https://zenodo.org/record/3765619/files/figure.png" httpUri-2="https://zenodo.org/record/3765621/files/figure.png" pageId="3" pageNumber="475">Figs. 2–4</figureCitation>
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),
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<collectingCounty id="62C294CBFFFDBB1E6285F8C58341F34C" box="[740,858,1870,1896]" pageId="3" pageNumber="475">Peninsular</collectingCounty>
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<collectingCountry id="F30BACD7FFFDBB1E6302F8C583D7F34C" box="[867,972,1870,1896]" name="Malaysia" pageId="3" pageNumber="475">Malaysia</collectingCountry>
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,
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,
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<collectingMunicipality id="6BC7763DFFFDBB1E641CF8C58500F34C" box="[1149,1307,1870,1896]" pageId="3" pageNumber="475">Pulau Tioman</collectingMunicipality>
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(=
|
||
<location id="8EC3BA9CFFFDBB1E6527F8C580FBF3A9" LSID="urn:lsid:plazi:treatment:03B55D51FFFDBB1860F6F924835BF355:8EC3BA9CFFFDBB1E6527F8C580FBF3A9" country="Malaysia" county="Peninsular" latitude="2.779" longLatPrecision="1" longitude="104.15703" municipality="Pulau Tioman" name="Tioman Island" pageId="3" pageNumber="475" stateProvince="Pahang State">Tioman Island</location>
|
||
),
|
||
<location id="8EC3BA9CFFFDBB1E6093F8F981B8F3A8" LSID="urn:lsid:plazi:treatment:03B55D51FFFDBB1860F6F924835BF355:8EC3BA9CFFFDBB1E6093F8F981B8F3A8" box="[242,419,1906,1932]" country="Malaysia" county="Peninsular" latitude="2.779" longLatPrecision="1" longitude="104.15703" municipality="Pulau Tioman" name="Gunung Kajang" pageId="3" pageNumber="475" stateProvince="Pahang State">Gunung Kajang</location>
|
||
trail,
|
||
<geoCoordinate id="EE288A80FFFDBB1E6182F8F98246F3A8" box="[483,605,1906,1932]" degrees="2.77900" direction="north" orientation="latitude" pageId="3" pageNumber="475" precision="1" value="2.779">2.77900°N</geoCoordinate>
|
||
<geoCoordinate id="EE288A80FFFDBB1E6205F8FA82EDF3A8" box="[612,758,1905,1932]" degrees="104.15703" direction="east" orientation="longitude" pageId="3" pageNumber="475" precision="1" value="104.15703">104.15703°E</geoCoordinate>
|
||
(
|
||
<location id="8EC3BA9CFFFDBB1E6365F8F98349F3A8" LSID="urn:lsid:plazi:treatment:03B55D51FFFDBB1860F6F924835BF355:8EC3BA9CFFFDBB1E6365F8F98349F3A8" box="[772,850,1906,1932]" country="Malaysia" county="Peninsular" latitude="2.779" longLatPrecision="1" longitude="104.15703" municipality="Pulau Tioman" name="Datum" pageId="3" pageNumber="475" stateProvince="Pahang State">Datum</location>
|
||
WGS84),
|
||
<quantity id="4CE441A2FFFDBB1E63A8F8F98414F3A9" box="[969,1039,1906,1933]" metricMagnitude="2" metricUnit="m" metricValue="6.18" pageId="3" pageNumber="475" unit="m" value="618.0">
|
||
<elevation id="00310B74FFFDBB1E63A8F8F98414F3A9" box="[969,1039,1906,1933]" metricMagnitude="2" metricUnit="m" metricValue="6.18" pageId="3" pageNumber="475" unit="m" value="618.0">618 m</elevation>
|
||
</quantity>
|
||
elev., coll.
|
||
<date id="FFA2CA87FFFDBB1E64EDF8F98520F3A8" box="[1164,1339,1906,1932]" pageId="3" pageNumber="475" value="2002-08-11">
|
||
<collectingDate id="EFE6336FFFFDBB1E64EDF8F98520F3A8" box="[1164,1339,1906,1932]" pageId="3" pageNumber="475" value="2002-08-11">11 August 2002</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26E98991FFFDBB1E6502F8F8810FF394" pageId="3" pageNumber="475">Jesse L. Grismer</collectorName>
|
||
,
|
||
<collectorName id="26E98991FFFDBB1E617EF81C81F5F395" box="[287,494,1942,1969]" pageId="3" pageNumber="475">Perry L. Wood, Jr.</collectorName>
|
||
, and L. Lee Grismer.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306BFCCFFFDBB1E60A6F8318550F3DC" pageId="3" pageNumber="475" type="etymology">
|
||
<paragraph id="8BA3EC47FFFDBB1E60A6F8318550F3DC" blockId="3.[151,1437,1869,2041]" pageId="3" pageNumber="475">
|
||
<emphasis id="B9683055FFFDBB1E60A6F8318155F3F0" bold="true" box="[199,334,1978,2004]" pageId="3" pageNumber="475">Etymology.</emphasis>
|
||
The specific epithet refers to the new species’
|
||
<typeStatus id="54A752E5FFFDBB1E633BF8308391F3F1" box="[858,906,1979,2005]" pageId="3" pageNumber="475">type</typeStatus>
|
||
and only known locality on the island of Pulau Tioman. The specific epithet is feminine, in agreement with the gender of the genus (
|
||
<bibRefCitation id="EF8D91B6FFFDBB1E642EF8558524F3DC" author="Poyarkov" box="[1103,1343,2014,2040]" etAl="et al." firstAuthor="Poyarkov" pageId="3" pageNumber="475" pagination="1647 - 1672" refId="ref7003" refString="Poyarkov, N. A., Nguyen, T. V. & Vogel, G. (2019) A new species of the genus Liopeltis Fitzinger 1843 from Vietnam (Squamata: Colubridae). Journal of Natural History, 53, 1647 - 1672. https: // doi. org / 10.1080 / 00222933.2019.1656784" type="journal article" year="2019">
|
||
Poyarkov
|
||
<emphasis id="B9683055FFFDBB1E64A2F85484E0F3DC" box="[1219,1275,2014,2040]" italics="true" pageId="3" pageNumber="475">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306BFCCFFFABB1960A6FF13856EF4DF" pageId="4" pageNumber="476" type="diagnosis">
|
||
<paragraph id="8BA3EC47FFFABB1960A6FF13856EF4DF" blockId="4.[151,1437,152,683]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9683055FFFABB1960A6FF138159F497" bold="true" box="[199,322,152,179]" pageId="4" pageNumber="476">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="4C1C97C4FFFABB1961B4FF12822CF497" authorityName="Fitzinger" authorityYear="1843" box="[469,567,153,179]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9683055FFFABB1961B4FF12822CF497" box="[469,567,153,179]" italics="true" pageId="4" pageNumber="476">Liopeltis</emphasis>
|
||
</taxonomicName>
|
||
having the combination of the nasal scale fused with the internasal scale; preoculars two; ventrals 161; distinct black lateral cephalic stripe extending from rostral through eye to approximately
|
||
<quantity id="4CE441A2FFFABB1960F6FF6A80FDF4DF" box="[151,230,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="476" unit="mm" value="30.0">30 mm</quantity>
|
||
behind neck before fading; four longitudinal stripes on dorsum; and venter immaculate, without stripes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306BFCCFFFABB1860A6FE8F850FF1C2" lastPageId="5" lastPageNumber="477" pageId="4" pageNumber="476" type="description">
|
||
<paragraph id="8BA3EC47FFFABB1960A6FE8F8175F68F" blockId="4.[151,1437,152,683]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9683055FFFABB1960A6FE8F8273F53B" bold="true" box="[199,616,260,287]" pageId="4" pageNumber="476">
|
||
Description of
|
||
<typeStatus id="54A752E5FFFABB196116FE8E81C5F53B" box="[375,478,261,287]" pageId="4" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1327F0C2FFFABB19618CFE8F8241F53B" box="[493,602,260,287]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,250,1124,1149]" captionStartId-1="6.[151,250,1265,1290]" captionTargetBox-0="[151,1436,181,1099]" captionTargetBox-1="[158,1429,181,1240]" captionTargetId-0="figure@5.[151,1436,181,1099]" captionTargetId-1="figure@6.[158,1429,181,1240]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Holotype (LSUHC 5037) of Liopeltis tiomanica sp. nov. in life. Photograph by LLG." captionText-1="FIGURE 3. Holotype (LSUHC 5037) Liopeltis tiomanica sp. nov. in preservative. A. dorsal view of body. B. Ventral view of body. C. Lateral view of head. D. Dorsal view of head. E. Ventral view of head." figureDoi-0="http://doi.org/10.5281/zenodo.3765617" figureDoi-1="http://doi.org/10.5281/zenodo.3765619" httpUri-0="https://zenodo.org/record/3765617/files/figure.png" httpUri-1="https://zenodo.org/record/3765619/files/figure.png" pageId="4" pageNumber="476">Figs. 2–3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Adult female; head slightly distinct from neck, triangular in dorsal view; snout weakly pointed in lateral view, projecting beyond lower jaw; eye large, pupil round; body slender; tail thin. Rostral wide, clearly visible from above; nasal partially divided, portion anterior to nostril continuous with internasal, portion posterior to nostril with horizontal suture; loreal absent; two preoculars, upper rectangular, approximately 25% size of lower, trapezoidal preocular; two postoculars, squarish or rounded, upper scale slightly larger than lower scale; eight supralabials, 4
|
||
<superScript id="7C69410FFFFABB196265FE33820AF5E2" attach="left" box="[516,529,440,454]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
and 5
|
||
<superScript id="7C69410FFFFABB196234FE338279F5E2" attach="left" box="[597,610,440,454]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
in contact with eye, 2
|
||
<superScript id="7C69410FFFFABB196336FE33836DF5F7" attach="none" box="[855,886,440,467]" fontSize="6" pageId="4" pageNumber="476">nd–</superScript>
|
||
4
|
||
<superScript id="7C69410FFFFABB1963E2FE33838BF5E2" attach="left" box="[899,912,440,454]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
in contact with lower preocular, 5
|
||
<superScript id="7C69410FFFFABB19656FFE338500F5E2" attach="left" box="[1294,1307,440,454]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
(right side) and 5
|
||
<superScript id="7C69410FFFFABB1960B5FE5780EBF5D3" attach="none" box="[212,240,476,503]" fontSize="6" pageId="4" pageNumber="476">th–</superScript>
|
||
6
|
||
<superScript id="7C69410FFFFABB19609CFE578111F5CE" attach="left" box="[253,266,476,490]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
(left side) in contact with lower postocular, 7
|
||
<superScript id="7C69410FFFFABB196363FE578314F5CE" attach="left" box="[770,783,476,490]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
largest; eight infralabials, 5
|
||
<superScript id="7C69410FFFFABB196427FE578448F5CE" attach="left" box="[1094,1107,476,490]" fontSize="6" pageId="4" pageNumber="476">th</superScript>
|
||
largest; one elongate anterior temporal; two subequal posterior temporals, shorter than anterior temporal; two internasals; two prefrontals; single frontal, approximately 63% length of parietals, with one supraocular on each side; two large, elongate parietals; anterior pair and posterior pairs of chin shields subequal in length, anterior pair broader; dorsal scales smooth, in rows 15:15:13, apical pits absent; 161 ventrals; cloacal plate divided; 119 paired subcaudals. Scale counts are summarized in
|
||
<tableCitation id="C69ED9FCFFFABB196174FD1A8171F68F" box="[277,362,657,683]" captionStart="TABLE 2" captionStartId="4.[151,239,730,754]" captionTargetPageId="4" captionText="TABLE 2. Morphological characters of Liopeltis tiomanica sp. nov. from Pulau Tioman, Peninsular Malaysia, and its sister taxon L. philippina from the Palawan Pleistocene Aggregate Island Complex (PAIC) of the western Philippines." httpUri="http://table.plazi.org/id/DF63BCCFFFFABB1960F6FD51859BF733" pageId="4" pageNumber="476" tableUuid="DF63BCCFFFFABB1960F6FD51859BF733">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF63BCCFFFFABB1960F6FD51859BF733" ID-Table-UUID="DF63BCCFFFFABB1960F6FD51859BF733" httpUri="http://table.plazi.org/id/DF63BCCFFFFABB1960F6FD51859BF733" pageId="4" pageNumber="476" targetBox="[166,1418,808,1639]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BA3EC47FFFABB1960F6FD51859BF733" blockId="4.[151,1436,730,791]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9683055FFFABB1960F6FD518117F6D7" bold="true" box="[151,268,730,755]" pageId="4" pageNumber="476">TABLE 2.</emphasis>
|
||
Morphological characters of
|
||
<taxonomicName id="4C1C97C4FFFABB19622EFD50833BF6D6" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[591,800,731,755]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFABB19622EFD50833BF6D6" box="[591,800,731,755]" italics="true" pageId="4" pageNumber="476">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFABB196349FD508364F6D7" bold="true" box="[808,895,731,755]" pageId="4" pageNumber="476">
|
||
<taxonomicNameLabel id="A25B8D2EFFFABB196349FD508364F6D7" box="[808,895,731,755]" pageId="4" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Pulau Tioman, Peninsular Malaysia, and its sister taxon
|
||
<taxonomicName id="4C1C97C4FFFABB196179FD7481B9F733" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[280,418,767,791]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFABB196179FD7481B9F733" box="[280,418,767,791]" italics="true" pageId="4" pageNumber="476">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
from the Palawan Pleistocene Aggregate Island Complex (PAIC) of the western Philippines.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA3EC47FFFABB196126FCA38554F243" pageId="4" pageNumber="476">
|
||
<table id="F91C1EE7FFFA44E260C7FCA38591F243" box="[166,1418,808,1639]" gridcols="6" gridrows="17" pageId="4" pageNumber="476">
|
||
<tr id="352CEE05FFFA44E260C7FCA38591F7A1" box="[166,1418,808,901]" gridrow="0" pageId="4" pageNumber="476" rowspan-0="1">
|
||
<th id="76FD8779FFFA44E2615CFCA381E4F7A1" box="[317,511,808,901]" gridcol="1" gridrow="0" pageId="4" pageNumber="476">
|
||
<taxonomicName id="4C1C97C4FFFABB196126FCA381BCF719" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[327,423,808,829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFABB196126FCA381BCF719" box="[327,423,808,829]" italics="true" pageId="4" pageNumber="476">tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFABB1961CCFCA381EEF719" bold="true" box="[429,501,808,829]" pageId="4" pageNumber="476">
|
||
<taxonomicNameLabel id="A25B8D2EFFFABB1961CCFCA381EEF719" box="[429,501,808,829]" pageId="4" pageNumber="476" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype female LSUHC 5037
|
||
</th>
|
||
<th id="76FD8779FFFA44E26244FCA382EAF7A1" box="[549,753,808,901]" gridcol="2" gridrow="0" pageId="4" pageNumber="476">
|
||
<taxonomicName id="4C1C97C4FFFABB19623BFCA382A6F719" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[602,701,808,829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFABB19623BFCA382A6F719" box="[602,701,808,829]" italics="true" pageId="4" pageNumber="476">philippina</emphasis>
|
||
</taxonomicName>
|
||
Male FMNH 53372
|
||
</th>
|
||
<th id="76FD8779FFFA44E26379FCA383C1F7A1" box="[792,986,808,901]" gridcol="3" gridrow="0" pageId="4" pageNumber="476">
|
||
<taxonomicName id="4C1C97C4FFFABB196329FCA383B0F719" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[840,939,808,829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFABB196329FCA383B0F719" box="[840,939,808,829]" italics="true" pageId="4" pageNumber="476">philippina</emphasis>
|
||
</taxonomicName>
|
||
Female CAS 15284
|
||
</th>
|
||
<th id="76FD8779FFFA44E2639FFCA384D1F7A1" box="[1022,1226,808,901]" gridcol="4" gridrow="0" pageId="4" pageNumber="476">
|
||
<taxonomicName id="4C1C97C4FFFABB196452FCA3848DF719" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[1075,1174,808,829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFABB196452FCA3848DF719" box="[1075,1174,808,829]" italics="true" pageId="4" pageNumber="476">philippina</emphasis>
|
||
</taxonomicName>
|
||
Male KU 327731
|
||
</th>
|
||
<th id="76FD8779FFFA44E2648EFCA38591F7A1" box="[1263,1418,808,901]" gridcol="5" gridrow="0" pageId="4" pageNumber="476">
|
||
<taxonomicName id="4C1C97C4FFFABB19656AFCA38575F719" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[1291,1390,808,829]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="476" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFABB19656AFCA38575F719" box="[1291,1390,808,829]" italics="true" pageId="4" pageNumber="476">philippina</emphasis>
|
||
</taxonomicName>
|
||
Male BoS 940*
|
||
</th>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FC1C8591F789" box="[166,1418,919,941]" gridrow="1" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FC1C8106F789" box="[166,285,919,941]" gridcol="0" gridrow="1" pageId="4" pageNumber="476">Internasals</th>
|
||
<td id="76FD8779FFFA44E2615CFC1C81E4F789" box="[317,511,919,941]" gridcol="1" gridrow="1" pageId="4" pageNumber="476">2, fused to nasal</td>
|
||
<td id="76FD8779FFFA44E26244FC1C82EAF789" box="[549,753,919,941]" gridcol="2" gridrow="1" pageId="4" pageNumber="476">2, fused to nasal</td>
|
||
<td id="76FD8779FFFA44E26379FC1C83C1F789" box="[792,986,919,941]" gridcol="3" gridrow="1" pageId="4" pageNumber="476">2, fused to nasal</td>
|
||
<td id="76FD8779FFFA44E2639FFC1C84D1F789" box="[1022,1226,919,941]" gridcol="4" gridrow="1" pageId="4" pageNumber="476">2, fused to nasal</td>
|
||
<td id="76FD8779FFFA44E2648EFC1C8591F789" box="[1263,1418,919,941]" gridcol="5" gridrow="1" pageId="4" pageNumber="476">2, fused to nasal</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FC358591F7F0" box="[166,1418,958,980]" gridrow="2" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FC358106F7F0" box="[166,285,958,980]" gridcol="0" gridrow="2" pageId="4" pageNumber="476">Prefrontals</th>
|
||
<td id="76FD8779FFFA44E2615CFC3581E4F7F0" box="[317,511,958,980]" gridcol="1" gridrow="2" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26244FC3582EAF7F0" box="[549,753,958,980]" gridcol="2" gridrow="2" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26379FC3583C1F7F0" box="[792,986,958,980]" gridcol="3" gridrow="2" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2639FFC3584D1F7F0" box="[1022,1226,958,980]" gridcol="4" gridrow="2" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2648EFC358591F7F0" box="[1263,1418,958,980]" gridcol="5" gridrow="2" pageId="4" pageNumber="476">2</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FC6E8591F7DF" box="[166,1418,997,1019]" gridrow="3" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FC6E8106F7DF" box="[166,285,997,1019]" gridcol="0" gridrow="3" pageId="4" pageNumber="476">Frontal</th>
|
||
<td id="76FD8779FFFA44E2615CFC6E81E4F7DF" box="[317,511,997,1019]" gridcol="1" gridrow="3" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E26244FC6E82EAF7DF" box="[549,753,997,1019]" gridcol="2" gridrow="3" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E26379FC6E83C1F7DF" box="[792,986,997,1019]" gridcol="3" gridrow="3" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E2639FFC6E84D1F7DF" box="[1022,1226,997,1019]" gridcol="4" gridrow="3" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E2648EFC6E8591F7DF" box="[1263,1418,997,1019]" gridcol="5" gridrow="3" pageId="4" pageNumber="476">1</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FB868591F007" box="[166,1418,1037,1059]" gridrow="4" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FB868106F007" box="[166,285,1037,1059]" gridcol="0" gridrow="4" pageId="4" pageNumber="476">Parietals</th>
|
||
<td id="76FD8779FFFA44E2615CFB8681E4F007" box="[317,511,1037,1059]" gridcol="1" gridrow="4" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26244FB8682EAF007" box="[549,753,1037,1059]" gridcol="2" gridrow="4" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26379FB8683C1F007" box="[792,986,1037,1059]" gridcol="3" gridrow="4" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2639FFB8684D1F007" box="[1022,1226,1037,1059]" gridcol="4" gridrow="4" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2648EFB868591F007" box="[1263,1418,1037,1059]" gridcol="5" gridrow="4" pageId="4" pageNumber="476">2</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FBBF8591F06E" box="[166,1418,1076,1098]" gridrow="5" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FBBF8106F06E" box="[166,285,1076,1098]" gridcol="0" gridrow="5" pageId="4" pageNumber="476">Temporals</th>
|
||
<td id="76FD8779FFFA44E2615CFBBF81E4F06E" box="[317,511,1076,1098]" gridcol="1" gridrow="5" pageId="4" pageNumber="476">1+2</td>
|
||
<td id="76FD8779FFFA44E26244FBBF82EAF06E" box="[549,753,1076,1098]" gridcol="2" gridrow="5" pageId="4" pageNumber="476">1+2</td>
|
||
<td id="76FD8779FFFA44E26379FBBF83C1F06E" box="[792,986,1076,1098]" gridcol="3" gridrow="5" pageId="4" pageNumber="476">1+2</td>
|
||
<td id="76FD8779FFFA44E2639FFBBF84D1F06E" box="[1022,1226,1076,1098]" gridcol="4" gridrow="5" pageId="4" pageNumber="476">1+2</td>
|
||
<td id="76FD8779FFFA44E2648EFBBF8591F06E" box="[1263,1418,1076,1098]" gridcol="5" gridrow="5" pageId="4" pageNumber="476">1+2</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FBD08591F09D" box="[166,1418,1115,1209]" gridrow="6" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FBD08106F09D" box="[166,285,1115,1209]" gridcol="0" gridrow="6" pageId="4" pageNumber="476">Loreal</th>
|
||
<td id="76FD8779FFFA44E2615CFBD081E4F09D" box="[317,511,1115,1209]" gridcol="1" gridrow="6" pageId="4" pageNumber="476">Absent, prefrontal in contact with supralabial 2</td>
|
||
<td id="76FD8779FFFA44E26244FBD082EAF09D" box="[549,753,1115,1209]" gridcol="2" gridrow="6" pageId="4" pageNumber="476">Absent, prefrontal in contact with supralabials 2–3</td>
|
||
<td id="76FD8779FFFA44E26379FBD083C1F09D" box="[792,986,1115,1209]" gridcol="3" gridrow="6" pageId="4" pageNumber="476">Absent, prefrontal in contact with supralabial 2</td>
|
||
<td id="76FD8779FFFA44E2639FFBD084D1F09D" box="[1022,1226,1115,1209]" gridcol="4" gridrow="6" pageId="4" pageNumber="476">Absent, prefrontal in contact with supralabials 2–3</td>
|
||
<td id="76FD8779FFFA44E2648EFBD08591F09D" box="[1263,1418,1115,1209]" gridcol="5" gridrow="6" pageId="4" pageNumber="476">Absent</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FB408591F0C5" box="[166,1418,1227,1249]" gridrow="7" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FB408106F0C5" box="[166,285,1227,1249]" gridcol="0" gridrow="7" pageId="4" pageNumber="476">Preoculars</th>
|
||
<td id="76FD8779FFFA44E2615CFB4081E4F0C5" box="[317,511,1227,1249]" gridcol="1" gridrow="7" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26244FB4082EAF0C5" box="[549,753,1227,1249]" gridcol="2" gridrow="7" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E26379FB4083C1F0C5" box="[792,986,1227,1249]" gridcol="3" gridrow="7" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E2639FFB4084D1F0C5" box="[1022,1226,1227,1249]" gridcol="4" gridrow="7" pageId="4" pageNumber="476">1</td>
|
||
<td id="76FD8779FFFA44E2648EFB408591F0C5" box="[1263,1418,1227,1249]" gridcol="5" gridrow="7" pageId="4" pageNumber="476">1</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FB798591F12C" box="[166,1418,1266,1288]" gridrow="8" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FB798106F12C" box="[166,285,1266,1288]" gridcol="0" gridrow="8" pageId="4" pageNumber="476">Postoculars</th>
|
||
<td id="76FD8779FFFA44E2615CFB7981E4F12C" box="[317,511,1266,1288]" gridcol="1" gridrow="8" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26244FB7982EAF12C" box="[549,753,1266,1288]" gridcol="2" gridrow="8" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E26379FB7983C1F12C" box="[792,986,1266,1288]" gridcol="3" gridrow="8" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2639FFB7984D1F12C" box="[1022,1226,1266,1288]" gridcol="4" gridrow="8" pageId="4" pageNumber="476">2</td>
|
||
<td id="76FD8779FFFA44E2648EFB798591F12C" box="[1263,1418,1266,1288]" gridcol="5" gridrow="8" pageId="4" pageNumber="476">2</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FA928591F10B" box="[166,1418,1305,1327]" gridrow="9" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FA928106F10B" box="[166,285,1305,1327]" gridcol="0" gridrow="9" pageId="4" pageNumber="476">Supralabials</th>
|
||
<td id="76FD8779FFFA44E2615CFA9281E4F10B" box="[317,511,1305,1327]" gridcol="1" gridrow="9" pageId="4" pageNumber="476">8 (4,5)</td>
|
||
<td id="76FD8779FFFA44E26244FA9282EAF10B" box="[549,753,1305,1327]" gridcol="2" gridrow="9" pageId="4" pageNumber="476">8 (4,5)</td>
|
||
<td id="76FD8779FFFA44E26379FA9283C1F10B" box="[792,986,1305,1327]" gridcol="3" gridrow="9" pageId="4" pageNumber="476">8 (4,5)</td>
|
||
<td id="76FD8779FFFA44E2639FFA9284D1F10B" box="[1022,1226,1305,1327]" gridcol="4" gridrow="9" pageId="4" pageNumber="476">8 (4,5)</td>
|
||
<td id="76FD8779FFFA44E2648EFA928591F10B" box="[1263,1418,1305,1327]" gridcol="5" gridrow="9" pageId="4" pageNumber="476">8 (4,5)</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FACA8591F173" box="[166,1418,1345,1367]" gridrow="10" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FACA8106F173" box="[166,285,1345,1367]" gridcol="0" gridrow="10" pageId="4" pageNumber="476">Infralabials</th>
|
||
<td id="76FD8779FFFA44E2615CFACA81E4F173" box="[317,511,1345,1367]" gridcol="1" gridrow="10" pageId="4" pageNumber="476">8</td>
|
||
<td id="76FD8779FFFA44E26244FACA82EAF173" box="[549,753,1345,1367]" gridcol="2" gridrow="10" pageId="4" pageNumber="476">8</td>
|
||
<td id="76FD8779FFFA44E26379FACA83C1F173" box="[792,986,1345,1367]" gridcol="3" gridrow="10" pageId="4" pageNumber="476">8</td>
|
||
<td id="76FD8779FFFA44E2639FFACA84D1F173" box="[1022,1226,1345,1367]" gridcol="4" gridrow="10" pageId="4" pageNumber="476">8</td>
|
||
<td id="76FD8779FFFA44E2648EFACA8591F173" box="[1263,1418,1345,1367]" gridcol="5" gridrow="10" pageId="4" pageNumber="476">8</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FAE38591F15A" box="[166,1418,1384,1406]" gridrow="11" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FAE38106F15A" box="[166,285,1384,1406]" gridcol="0" gridrow="11" pageId="4" pageNumber="476">Dorsals</th>
|
||
<td id="76FD8779FFFA44E2615CFAE381E4F15A" box="[317,511,1384,1406]" gridcol="1" gridrow="11" pageId="4" pageNumber="476">15:15:13</td>
|
||
<td id="76FD8779FFFA44E26244FAE382EAF15A" box="[549,753,1384,1406]" gridcol="2" gridrow="11" pageId="4" pageNumber="476">15:15:13</td>
|
||
<td id="76FD8779FFFA44E26379FAE383C1F15A" box="[792,986,1384,1406]" gridcol="3" gridrow="11" pageId="4" pageNumber="476">15:15:13</td>
|
||
<td id="76FD8779FFFA44E2639FFAE384D1F15A" box="[1022,1226,1384,1406]" gridcol="4" gridrow="11" pageId="4" pageNumber="476">15:15:13</td>
|
||
<td id="76FD8779FFFA44E2648EFAE38591F15A" box="[1263,1418,1384,1406]" gridcol="5" gridrow="11" pageId="4" pageNumber="476">15</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FA048591F181" box="[166,1418,1423,1445]" gridrow="12" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FA048106F181" box="[166,285,1423,1445]" gridcol="0" gridrow="12" pageId="4" pageNumber="476">Ventrals</th>
|
||
<td id="76FD8779FFFA44E2615CFA0481E4F181" box="[317,511,1423,1445]" gridcol="1" gridrow="12" pageId="4" pageNumber="476">161</td>
|
||
<td id="76FD8779FFFA44E26244FA0482EAF181" box="[549,753,1423,1445]" gridcol="2" gridrow="12" pageId="4" pageNumber="476">139</td>
|
||
<td id="76FD8779FFFA44E26379FA0483C1F181" box="[792,986,1423,1445]" gridcol="3" gridrow="12" pageId="4" pageNumber="476">150</td>
|
||
<td id="76FD8779FFFA44E2639FFA0484D1F181" box="[1022,1226,1423,1445]" gridcol="4" gridrow="12" pageId="4" pageNumber="476">142</td>
|
||
<td id="76FD8779FFFA44E2648EFA048591F181" box="[1263,1418,1423,1445]" gridcol="5" gridrow="12" pageId="4" pageNumber="476">140</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FA3C8591F1E9" box="[166,1418,1463,1485]" gridrow="13" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FA3C8106F1E9" box="[166,285,1463,1485]" gridcol="0" gridrow="13" pageId="4" pageNumber="476">Subcaudals</th>
|
||
<td id="76FD8779FFFA44E2615CFA3C81E4F1E9" box="[317,511,1463,1485]" gridcol="1" gridrow="13" pageId="4" pageNumber="476">119</td>
|
||
<td id="76FD8779FFFA44E26244FA3C82EAF1E9" box="[549,753,1463,1485]" gridcol="2" gridrow="13" pageId="4" pageNumber="476">115</td>
|
||
<td id="76FD8779FFFA44E26379FA3C83C1F1E9" box="[792,986,1463,1485]" gridcol="3" gridrow="13" pageId="4" pageNumber="476">110</td>
|
||
<td id="76FD8779FFFA44E2639FFA3C84D1F1E9" box="[1022,1226,1463,1485]" gridcol="4" gridrow="13" pageId="4" pageNumber="476">116</td>
|
||
<td id="76FD8779FFFA44E2648EFA3C8591F1E9" box="[1263,1418,1463,1485]" gridcol="5" gridrow="13" pageId="4" pageNumber="476">119</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7FA558591F23C" box="[166,1418,1502,1560]" gridrow="14" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7FA558106F23C" box="[166,285,1502,1560]" gridcol="0" gridrow="14" pageId="4" pageNumber="476">Snout-vent (mm)</th>
|
||
<td id="76FD8779FFFA44E2615CFA5581E4F23C" box="[317,511,1502,1560]" gridcol="1" gridrow="14" pageId="4" pageNumber="476">304</td>
|
||
<td id="76FD8779FFFA44E26244FA5582EAF23C" box="[549,753,1502,1560]" gridcol="2" gridrow="14" pageId="4" pageNumber="476">308</td>
|
||
<td id="76FD8779FFFA44E26379FA5583C1F23C" box="[792,986,1502,1560]" gridcol="3" gridrow="14" pageId="4" pageNumber="476">341</td>
|
||
<td id="76FD8779FFFA44E2639FFA5584D1F23C" box="[1022,1226,1502,1560]" gridcol="4" gridrow="14" pageId="4" pageNumber="476">258</td>
|
||
<td id="76FD8779FFFA44E2648EFA558591F23C" box="[1263,1418,1502,1560]" gridcol="5" gridrow="14" pageId="4" pageNumber="476">390</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7F9A28591F21B" box="[166,1418,1577,1599]" gridrow="15" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7F9A28106F21B" box="[166,285,1577,1599]" gridcol="0" gridrow="15" pageId="4" pageNumber="476">Tail (mm)</th>
|
||
<td id="76FD8779FFFA44E2615CF9A281E4F21B" box="[317,511,1577,1599]" gridcol="1" gridrow="15" pageId="4" pageNumber="476">187</td>
|
||
<td id="76FD8779FFFA44E26244F9A282EAF21B" box="[549,753,1577,1599]" gridcol="2" gridrow="15" pageId="4" pageNumber="476">220</td>
|
||
<td id="76FD8779FFFA44E26379F9A283C1F21B" box="[792,986,1577,1599]" gridcol="3" gridrow="15" pageId="4" pageNumber="476">186</td>
|
||
<td id="76FD8779FFFA44E2639FF9A284D1F21B" box="[1022,1226,1577,1599]" gridcol="4" gridrow="15" pageId="4" pageNumber="476">175</td>
|
||
<td id="76FD8779FFFA44E2648EF9A28591F21B" box="[1263,1418,1577,1599]" gridcol="5" gridrow="15" pageId="4" pageNumber="476">250</td>
|
||
</tr>
|
||
<tr id="352CEE05FFFA44E260C7F9DB8591F243" box="[166,1418,1616,1639]" gridrow="16" pageId="4" pageNumber="476">
|
||
<th id="76FD8779FFFA44E260C7F9DB8106F243" box="[166,285,1616,1639]" gridcol="0" gridrow="16" pageId="4" pageNumber="476">Total (mm)</th>
|
||
<td id="76FD8779FFFA44E2615CF9DB81E4F243" box="[317,511,1616,1639]" gridcol="1" gridrow="16" pageId="4" pageNumber="476">491</td>
|
||
<td id="76FD8779FFFA44E26244F9DB82EAF243" box="[549,753,1616,1639]" gridcol="2" gridrow="16" pageId="4" pageNumber="476">528</td>
|
||
<td id="76FD8779FFFA44E26379F9DB83C1F243" box="[792,986,1616,1639]" gridcol="3" gridrow="16" pageId="4" pageNumber="476">527</td>
|
||
<td id="76FD8779FFFA44E2639FF9DB84D1F243" box="[1022,1226,1616,1639]" gridcol="4" gridrow="16" pageId="4" pageNumber="476">433</td>
|
||
<td id="76FD8779FFFA44E2648EF9DB8591F243" box="[1263,1418,1616,1639]" gridcol="5" gridrow="16" pageId="4" pageNumber="476">640</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BA3EC47FFFABB1960F6F9FF8572F2A8" blockId="4.[151,1385,1652,1676]" box="[151,1385,1652,1676]" pageId="4" pageNumber="476">
|
||
*Data from
|
||
<bibRefCitation id="EF8D91B6FFFABB196179F9FF81C1F2A8" author="Taylor" box="[280,474,1652,1676]" firstAuthor="Taylor" pageId="4" pageNumber="476" refId="ref7483" refString="Taylor, E. H. (1922) The Snakes of the Philippine Islands. Bureau of Printing, Manila, Philippines, 312 pp. https: // doi. org / 10.5962 / bhl. title. 55346" type="book" year="1922">Taylor (1922: 164</bibRefCitation>
|
||
, “No. 940, Bureau of Science Collection;” specimen destroyed during World War II).
|
||
</paragraph>
|
||
<paragraph id="8BA3EC47FFFABB1960A6F93181C8F389" blockId="4.[151,1437,1722,1965]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9683055FFFABB1960A6F931825BF2F1" bold="true" box="[199,576,1722,1749]" pageId="4" pageNumber="476">
|
||
Color of
|
||
<typeStatus id="54A752E5FFFABB19614EF930818DF2F1" box="[303,406,1723,1749]" pageId="4" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
in life (
|
||
<figureCitation id="1327F0C2FFFABB19618CF9318229F2F1" box="[493,562,1722,1749]" captionStart="FIGURE 2" captionStartId="5.[151,250,1124,1149]" captionTargetBox="[151,1436,181,1099]" captionTargetId="figure@5.[151,1436,181,1099]" captionTargetPageId="5" captionText="FIGURE 2. Holotype (LSUHC 5037) of Liopeltis tiomanica sp. nov. in life. Photograph by LLG." figureDoi="http://doi.org/10.5281/zenodo.3765617" httpUri="https://zenodo.org/record/3765617/files/figure.png" pageId="4" pageNumber="476">Fig. 2</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Dorsal surface of head bronze-brown (Grayish Horn Color 268); chin, supralabials, and infralabials cream (Cream White 52); black stripe on lateral side of head and neck separates dorsal and ventral coloration, stripe slightly wider than one dorsal scale, extending from rostral though eye to approximately
|
||
<quantity id="4CE441A2FFFABB1960F6F8AC80FDF365" box="[151,230,1831,1857]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="476" unit="mm" value="30.0">30 mm</quantity>
|
||
posterior to neck, gradually fading; dorsum bronze (Grayish Horn Color 268) with four longitudinal brown (Olive-Brown 278) stripes, two dorsal stripes extending from neck to tail, two lateral stripes extending from area posterior of black head stripe to level of cloacal plate; venter uniform pale cream color (Cream White 52) with small dark spots on ventral scales.
|
||
</paragraph>
|
||
<caption id="DF63BCCFFFFBBB1860F6FBEF8461F059" ID-DOI="http://doi.org/10.5281/zenodo.3765617" ID-Zenodo-Dep="3765617" box="[151,1146,1124,1149]" httpUri="https://zenodo.org/record/3765617/files/figure.png" pageId="5" pageNumber="477" startId="5.[151,250,1124,1149]" targetBox="[151,1436,181,1099]" targetPageId="5">
|
||
<paragraph id="8BA3EC47FFFBBB1860F6FBEF8461F059" blockId="5.[151,1146,1124,1149]" box="[151,1146,1124,1149]" pageId="5" pageNumber="477">
|
||
<emphasis id="B9683055FFFBBB1860F6FBEF8108F059" bold="true" box="[151,275,1124,1149]" pageId="5" pageNumber="477">FIGURE 2.</emphasis>
|
||
Holotype (LSUHC 5037) of
|
||
<taxonomicName id="4C1C97C4FFFBBB18625CFBEE831AF059" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[573,769,1125,1149]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB18625CFBEE831AF059" box="[573,769,1125,1149]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB186366FBEE8343F059" bold="true" box="[775,856,1125,1149]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB186366FBEE8343F059" box="[775,856,1125,1149]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life. Photograph by LLG.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA3EC47FFFBBB1860A6FB218291F0CD" blockId="5.[151,1437,1194,1906]" pageId="5" pageNumber="477">
|
||
<emphasis id="B9683055FFFBBB1860A6FB2182DDF0E1" bold="true" box="[199,710,1194,1221]" pageId="5" pageNumber="477">
|
||
Color of
|
||
<typeStatus id="54A752E5FFFBBB186156FB208185F0E1" box="[311,414,1195,1221]" pageId="5" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
in preservative (
|
||
<figureCitation id="1327F0C2FFFBBB18620FFB2182A3F0E1" box="[622,696,1194,1221]" captionStart="FIGURE 3" captionStartId="6.[151,250,1265,1290]" captionTargetBox="[158,1429,181,1240]" captionTargetId="figure@6.[158,1429,181,1240]" captionTargetPageId="6" captionText="FIGURE 3. Holotype (LSUHC 5037) Liopeltis tiomanica sp. nov. in preservative. A. dorsal view of body. B. Ventral view of body. C. Lateral view of head. D. Dorsal view of head. E. Ventral view of head." figureDoi="http://doi.org/10.5281/zenodo.3765619" httpUri="https://zenodo.org/record/3765619/files/figure.png" pageId="5" pageNumber="477">Fig. 3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Dorsum faded to grayish green (Olive Horn Color 16); dorsal stripes faded to light brown (Clay Color 18).
|
||
</paragraph>
|
||
<paragraph id="8BA3EC47FFFBBB1860A6FB78850FF1C2" blockId="5.[151,1437,1194,1906]" pageId="5" pageNumber="477">
|
||
<emphasis id="B9683055FFFBBB1860A6FB78819EF129" bold="true" box="[199,389,1267,1293]" pageId="5" pageNumber="477">Molecular data.</emphasis>
|
||
The standard deviation of split frequencies among the four Bayesian runs was 0.003717 and the Estimated Sample Sizes (ESS) of parameters were ≥ 3,390. The
|
||
<typeStatus id="54A752E5FFFBBB1863E9FA9C83F1F115" box="[904,1002,1303,1329]" pageId="5" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1C97C4FFFBBB18646FFA9C84FDF115" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[1038,1254,1303,1329]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB18646FFA9C84FDF115" box="[1038,1254,1303,1329]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB18648CFA9C855EF115" bold="true" box="[1261,1349,1303,1329]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB18648CFA9C855EF115" box="[1261,1349,1303,1329]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was recovered with strong support (Bayesian posterior probability 1.00) to be the sister taxon of
|
||
<taxonomicName id="4C1C97C4FFFBBB18641AFAB78510F171" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[1147,1291,1339,1365]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB18641AFAB78510F171" box="[1147,1291,1339,1365]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1327F0C2FFFBBB18657BFAB08545F171" box="[1306,1374,1339,1365]" captionStart="FIGURE 4" captionStartId="7.[151,250,1860,1885]" captionTargetBox="[192,1382,184,1824]" captionTargetId="figure@7.[192,1395,181,1835]" captionTargetPageId="7" captionText="FIGURE 4. Fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 1,152 aligned characters of the complete mitochondrial cytochrome b gene and parts of its flanking tRNA genes from colubrid snakes of the genera Liopeltis, Gongylosoma and Sibynophis. Numbers above nodes are Bayesian posterior probabilities. Numbers at terminal tips are GenBank accession numbers. Voucher and locality data for samples are provided in Table 2. The holotype of Liopeltis tiomanica sp. nov. in life is shown in the image." figureDoi="http://doi.org/10.5281/zenodo.3765621" httpUri="https://zenodo.org/record/3765621/files/figure.png" pageId="5" pageNumber="477">Fig. 4</figureCitation>
|
||
). The
|
||
<taxonomicName id="4C1C97C4FFFBBB1860F6FAEB813FF15D" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[151,292,1376,1401]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB1860F6FAEB813FF15D" box="[151,292,1376,1401]" italics="true" pageId="5" pageNumber="477">L. tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB18614AFAD4819FF15D" bold="true" box="[299,388,1375,1401]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB18614AFAD4819FF15D" box="[299,388,1375,1401]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
+
|
||
<taxonomicName id="4C1C97C4FFFBBB1861C0FAEB822AF15D" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[417,561,1375,1401]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB1861C0FAEB822AF15D" box="[417,561,1375,1401]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
clade was recovered with strong support (Bayesian posterior probability 1.00) to be nested within a clade containing
|
||
<taxonomicName id="4C1C97C4FFFBBB186224FA0F82A2F1B9" box="[581,697,1411,1437]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB186224FA0F82A2F1B9" box="[581,697,1411,1437]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C97C4FFFBBB18628EFA0F8361F1B9" baseAuthorityName="Sclater" baseAuthorityYear="1891" box="[751,890,1411,1437]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="stoliczkae">
|
||
<emphasis id="B9683055FFFBBB18628EFA0F8361F1B9" box="[751,890,1411,1437]" italics="true" pageId="5" pageNumber="477">L. stoliczkae</emphasis>
|
||
</taxonomicName>
|
||
, although the relationships among
|
||
<taxonomicName id="4C1C97C4FFFBBB186565FA0F8563F1B9" box="[1284,1400,1411,1437]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB186565FA0F8563F1B9" box="[1284,1400,1411,1437]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C97C4FFFBBB1865E7FA0F811EF1E5" baseAuthorityName="Sclater" baseAuthorityYear="1891" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="stoliczkae">
|
||
<emphasis id="B9683055FFFBBB1865E7FA0F811EF1E5" italics="true" pageId="5" pageNumber="477">L. stoliczkae</emphasis>
|
||
</taxonomicName>
|
||
, and the
|
||
<taxonomicName id="4C1C97C4FFFBBB186111FA2381E4F1E5" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[368,511,1448,1473]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB186111FA2381E4F1E5" box="[368,511,1448,1473]" italics="true" pageId="5" pageNumber="477">L. tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB186269FA2C8278F1E5" bold="true" box="[520,611,1447,1473]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB186269FA2C8278F1E5" box="[520,611,1447,1473]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
+
|
||
<taxonomicName id="4C1C97C4FFFBBB1862E2FA23830EF1E5" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[643,789,1447,1473]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB1862E2FA23830EF1E5" box="[643,789,1447,1473]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
clade were not resolved (
|
||
<figureCitation id="1327F0C2FFFBBB18645DFA2C8499F1E5" box="[1084,1154,1447,1473]" captionStart="FIGURE 4" captionStartId="7.[151,250,1860,1885]" captionTargetBox="[192,1382,184,1824]" captionTargetId="figure@7.[192,1395,181,1835]" captionTargetPageId="7" captionText="FIGURE 4. Fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 1,152 aligned characters of the complete mitochondrial cytochrome b gene and parts of its flanking tRNA genes from colubrid snakes of the genera Liopeltis, Gongylosoma and Sibynophis. Numbers above nodes are Bayesian posterior probabilities. Numbers at terminal tips are GenBank accession numbers. Voucher and locality data for samples are provided in Table 2. The holotype of Liopeltis tiomanica sp. nov. in life is shown in the image." figureDoi="http://doi.org/10.5281/zenodo.3765621" httpUri="https://zenodo.org/record/3765621/files/figure.png" pageId="5" pageNumber="477">Fig. 4</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1C97C4FFFBBB1864F9FA2C8569F1E5" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[1176,1394,1447,1473]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB1864F9FA2C8569F1E5" box="[1176,1394,1447,1473]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB18651AFA2C80D3F1C1" bold="true" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB18651AFA2C80D3F1C1" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
had uncorrected
|
||
<emphasis id="B9683055FFFBBB1861EBFA478183F1C1" box="[394,408,1484,1509]" italics="true" pageId="5" pageNumber="477">p</emphasis>
|
||
-distances in cyt
|
||
<emphasis id="B9683055FFFBBB186234FA408278F1C1" box="[597,611,1483,1509]" italics="true" pageId="5" pageNumber="477">b</emphasis>
|
||
of 9.6% from
|
||
<taxonomicName id="4C1C97C4FFFBBB186366FA47838CF1C1" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[775,919,1483,1509]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB186366FA47838CF1C1" box="[775,919,1483,1509]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
and 12.9–13.3% from
|
||
<taxonomicName id="4C1C97C4FFFBBB1864FBFA478516F1C1" box="[1178,1293,1483,1509]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB1864FBFA478516F1C1" box="[1178,1293,1483,1509]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306BFCCFFFBBB1860A6FA658530F275" pageId="5" pageNumber="477" type="distribution">
|
||
<paragraph id="8BA3EC47FFFBBB1860A6FA658530F275" blockId="5.[151,1437,1194,1906]" pageId="5" pageNumber="477">
|
||
<emphasis id="B9683055FFFBBB1860A6FA65831BF22E" bold="true" box="[199,768,1518,1546]" pageId="5" pageNumber="477">Distribution, natural history, and conservation.</emphasis>
|
||
<taxonomicName id="4C1C97C4FFFBBB186368FA6483F8F22D" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[777,995,1519,1545]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB186368FA6483F8F22D" box="[777,995,1519,1545]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB18638AFA64845DF22D" bold="true" box="[1003,1094,1519,1545]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB18638AFA64845DF22D" box="[1003,1094,1519,1545]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="54A752E5FFFBBB186500FA6480DCF20A" pageId="5" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
specimen collected from Pulau Tioman (
|
||
<figureCitation id="1327F0C2FFFBBB1862F7F99882CCF209" box="[662,727,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[151,250,1823,1848]" captionTargetBox="[180,1409,186,1791]" captionTargetId="figure@2.[171,1416,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Map showing localities of material of Liopeltis examined in this study. The base map was generated using a global void filled grid obtained from the General Bathymetric Chart of the Oceans (GEBCO; https://www.gebco.net)." figureDoi="http://doi.org/10.5281/zenodo.3765615" httpUri="https://zenodo.org/record/3765615/files/figure.png" pageId="5" pageNumber="477">Fig. 1</figureCitation>
|
||
). The specimen was found in the afternoon while perched on a small tree branch
|
||
<quantity id="4CE441A2FFFBBB18613EF9BC8191F276" box="[351,394,1591,1618]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="477" unit="m" value="2.0">2 m</quantity>
|
||
above the ground in primary hill dipterocarp forest (J.
|
||
<bibRefCitation id="EF8D91B6FFFBBB186393F9BC84D4F275" author="Grismer, J. L. & Grismer, L. L. & Das, I. & Yaakob, N. S. & Liat, L. B. & Leong, T. M. & Youmans, T. M. & Kaiser, H." box="[1010,1231,1591,1617]" pageId="5" pageNumber="477" pagination="247 - 279" refId="ref5729" refString="Grismer, J. L., Grismer, L. L., Das, I., Yaakob, N. S., Liat, L. B., Leong, T. M., Youmans, T. M. & Kaiser, H. (2004) Species diversity and checklist of the herpetofauna of Pulau Tioman, Peninsular Malaysia, with a preliminary overview of habitat utilization. Asiatic Herpetological Research, 10, 247 - 279." type="journal article" year="2004">
|
||
Grismer
|
||
<emphasis id="B9683055FFFBBB186434F9B38496F275" box="[1109,1165,1591,1617]" italics="true" pageId="5" pageNumber="477">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
,
|
||
<figureCitation id="1327F0C2FFFBBB1864BBF9BC8506F275" box="[1242,1309,1590,1617]" captionStart="FIGURE 5" captionStartId="9.[151,250,1439,1464]" captionTargetBox="[151,1436,561,1414]" captionTargetId="figure@9.[151,1436,561,1414]" captionTargetPageId="9" captionText="FIGURE 5. A. Holotype (LSUHC 5037) of Liopeltis tiomanica sp. nov. in life. Photograph by LLG. B. Primary hill dipterocarp forest near the type locality of Liopeltis tiomanica sp. nov. on Pulau Tioman, Pahang State, Peninsular Malaysia. Photograph by LLG. C. Unvouchered L. philippina from Salvacion, Busuanga Island, Calamianes Archipelago, Palawan Prov., Philippines (12.14378ºN 119.93884ºE), in life. Photograph by Shalom Pareja. D. Male L. philippina (KU 327731) from Mount Elyan in the Mantalingajan Mountain Range, Municipality of Brooke’s Point, Palawan Island, Palawan Prov., Philippines, coll. 6 December 2001 by Arvin C. Diesmos, G. Gee, N. Antoque, M. L. Leonida, and J. C. Diesmos. Photograph by ACD. Note the distinct black lateral cephalic stripe in L. tiomanica sp. nov. that is absent in L. philippina." figureDoi="http://doi.org/10.5281/zenodo.3765623" httpUri="https://zenodo.org/record/3765623/files/figure.png" pageId="5" pageNumber="477">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306BFCCFFFBBB1860A6F9D1835BF355" pageId="5" pageNumber="477" type="discussion">
|
||
<paragraph id="8BA3EC47FFFBBB1860A6F9D1835BF355" blockId="5.[151,1437,1194,1906]" pageId="5" pageNumber="477">
|
||
<emphasis id="B9683055FFFBBB1860A6F9D18176F251" bold="true" box="[199,365,1626,1653]" pageId="5" pageNumber="477">Comparisons.</emphasis>
|
||
<taxonomicName id="4C1C97C4FFFBBB186115F9D08256F251" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[372,589,1627,1653]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB186115F9D08256F251" box="[372,589,1627,1653]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB186235F9D082B5F251" bold="true" box="[596,686,1627,1653]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB186235F9D082B5F251" box="[596,686,1627,1653]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all other species of
|
||
<taxonomicName id="4C1C97C4FFFBBB18647DF9D08465F251" authorityName="Fitzinger" authorityYear="1843" box="[1052,1150,1627,1653]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9683055FFFBBB18647DF9D08465F251" box="[1052,1150,1627,1653]" italics="true" pageId="5" pageNumber="477">Liopeltis</emphasis>
|
||
</taxonomicName>
|
||
, except its sister taxon
|
||
<taxonomicName id="4C1C97C4FFFBBB1865E7F9D78111F2BD" baseAuthorityName="Boettger" baseAuthorityYear="1897" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB1865E7F9D78111F2BD" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1327F0C2FFFBBB18617BF9F48144F2BD" box="[282,351,1663,1689]" captionStart="FIGURE 4" captionStartId="7.[151,250,1860,1885]" captionTargetBox="[192,1382,184,1824]" captionTargetId="figure@7.[192,1395,181,1835]" captionTargetPageId="7" captionText="FIGURE 4. Fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 1,152 aligned characters of the complete mitochondrial cytochrome b gene and parts of its flanking tRNA genes from colubrid snakes of the genera Liopeltis, Gongylosoma and Sibynophis. Numbers above nodes are Bayesian posterior probabilities. Numbers at terminal tips are GenBank accession numbers. Voucher and locality data for samples are provided in Table 2. The holotype of Liopeltis tiomanica sp. nov. in life is shown in the image." figureDoi="http://doi.org/10.5281/zenodo.3765621" httpUri="https://zenodo.org/record/3765621/files/figure.png" pageId="5" pageNumber="477">Fig. 4</figureCitation>
|
||
), by having the nasal fused with the internasal and four longitudinal stripes on the dorsum (
|
||
<bibRefCitation id="EF8D91B6FFFBBB186505F9F4815AF299" author="Weinell" etAl="et al." firstAuthor="Weinell" pageId="5" pageNumber="477" pagination="1 - 49" refId="ref7555" refString="Weinell, J. L., Hooper, E., Leviton, A. E. & Brown, R. M. (2019) Illustrated key to the snakes of the Philippines. Proceedings of the California Academy of Sciences, 66 (1), 1 - 49." type="journal article" year="2019">
|
||
Weinell
|
||
<emphasis id="B9683055FFFBBB1860A6F92F80E5F299" box="[199,254,1699,1725]" italics="true" pageId="5" pageNumber="477">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1C97C4FFFBBB186133F9288231F299" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[338,554,1699,1725]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB186133F9288231F299" box="[338,554,1699,1725]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB186251F9288292F299" bold="true" box="[560,649,1699,1725]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB186251F9288292F299" box="[560,649,1699,1725]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C1C97C4FFFBBB186378F92F83B2F299" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[793,937,1699,1725]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB186378F92F83B2F299" box="[793,937,1699,1725]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
by having 161 ventral scales (vs.
|
||
<quantity id="4CE441A2FFFBBB18657EF9298587F299" box="[1311,1436,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="3.6703" metricValueMax="3.81" metricValueMin="3.5305999999999997" pageId="5" pageNumber="477" unit="in" value="144.5" valueMax="150.0" valueMin="139.0">139–150 in</quantity>
|
||
<taxonomicName id="4C1C97C4FFFBBB1860F6F9438133F2C5" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[151,296,1735,1761]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB1860F6F9438133F2C5" box="[151,296,1735,1761]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
); two preocular scales (vs. one in
|
||
<taxonomicName id="4C1C97C4FFFBBB1862CBF9438320F2C5" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[682,827,1735,1761]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB1862CBF9438320F2C5" box="[682,827,1735,1761]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
); and a distinct black lateral cephalic stripe (vs. indistinct in
|
||
<taxonomicName id="4C1C97C4FFFBBB18608AF9678160F321" baseAuthorityName="Boettger" baseAuthorityYear="1897" box="[235,379,1771,1797]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B9683055FFFBBB18608AF9678160F321" box="[235,379,1771,1797]" italics="true" pageId="5" pageNumber="477">L. philippina</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="1327F0C2FFFBBB1861E8F96081D7F321" box="[393,460,1770,1797]" captionStart="FIGURE 5" captionStartId="9.[151,250,1439,1464]" captionTargetBox="[151,1436,561,1414]" captionTargetId="figure@9.[151,1436,561,1414]" captionTargetPageId="9" captionText="FIGURE 5. A. Holotype (LSUHC 5037) of Liopeltis tiomanica sp. nov. in life. Photograph by LLG. B. Primary hill dipterocarp forest near the type locality of Liopeltis tiomanica sp. nov. on Pulau Tioman, Pahang State, Peninsular Malaysia. Photograph by LLG. C. Unvouchered L. philippina from Salvacion, Busuanga Island, Calamianes Archipelago, Palawan Prov., Philippines (12.14378ºN 119.93884ºE), in life. Photograph by Shalom Pareja. D. Male L. philippina (KU 327731) from Mount Elyan in the Mantalingajan Mountain Range, Municipality of Brooke’s Point, Palawan Island, Palawan Prov., Philippines, coll. 6 December 2001 by Arvin C. Diesmos, G. Gee, N. Antoque, M. L. Leonida, and J. C. Diesmos. Photograph by ACD. Note the distinct black lateral cephalic stripe in L. tiomanica sp. nov. that is absent in L. philippina." figureDoi="http://doi.org/10.5281/zenodo.3765623" httpUri="https://zenodo.org/record/3765623/files/figure.png" pageId="5" pageNumber="477">Fig. 5</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1C97C4FFFBBB186181F96082A3F321" authority="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart, 2020" authorityName="Som & Grismer & Grismer & Wood & Quah & Brown & Diesmos & Weinell & Stuart" authorityYear="2020" box="[480,696,1771,1797]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tiomanica" status="sp. nov.">
|
||
<emphasis id="B9683055FFFBBB186181F96082A3F321" box="[480,696,1771,1797]" italics="true" pageId="5" pageNumber="477">Liopeltis tiomanica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9683055FFFBBB1862DEF960830CF321" bold="true" box="[703,791,1771,1797]" pageId="5" pageNumber="477">
|
||
<taxonomicNameLabel id="A25B8D2EFFFBBB1862DEF960830CF321" box="[703,791,1771,1797]" pageId="5" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
further differs from the sympatric (and possibly syntopic)
|
||
<taxonomicName id="4C1C97C4FFFBBB1860F6F89B8110F30D" box="[151,267,1807,1833]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB1860F6F89B8110F30D" box="[151,267,1807,1833]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
by having the prefrontal in contact only with the second supralabial (vs. second and third supralabials in
|
||
<taxonomicName id="4C1C97C4FFFBBB1860F6F8BF8110F369" box="[151,267,1843,1869]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB1860F6F8BF8110F369" box="[151,267,1843,1869]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
); and immaculate ventral scales (vs. three gray ventral stripes in most
|
||
<taxonomicName id="4C1C97C4FFFBBB186447F8BF8481F369" box="[1062,1178,1843,1869]" class="Reptilia" family="Colubridae" genus="Liopeltis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="477" phylum="Chordata" rank="species" species="tricolor">
|
||
<emphasis id="B9683055FFFBBB186447F8BF8481F369" box="[1062,1178,1843,1869]" italics="true" pageId="5" pageNumber="477">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
). Scalation differences among these three species are summarized in
|
||
<tableCitation id="C69ED9FCFFFBBB1862F6F8DC82EEF355" box="[663,757,1879,1905]" captionStart="TABLE 2" captionStartId="4.[151,239,730,754]" captionTargetPageId="4" captionText="TABLE 2. Morphological characters of Liopeltis tiomanica sp. nov. from Pulau Tioman, Peninsular Malaysia, and its sister taxon L. philippina from the Palawan Pleistocene Aggregate Island Complex (PAIC) of the western Philippines." httpUri="http://table.plazi.org/id/DF63BCCFFFFABB1960F6FD51859BF733" pageId="5" pageNumber="477" tableUuid="DF63BCCFFFFABB1960F6FD51859BF733">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C69ED9FCFFFBBB18634AF8DC8320F355" box="[811,827,1879,1905]" captionStart="TABLE 3" captionStartId="8.[151,239,664,690]" captionTargetPageId="8" captionText="TABLE 3. Morphological characters of Liopeltis tricolor, a species that occurs in sympatry with L. tiomanica sp. nov." httpUri="http://table.plazi.org/id/DF63BCCFFFF6BB1560F6FD138593F696" pageId="5" pageNumber="477" tableUuid="DF63BCCFFFF6BB1560F6FD138593F696">3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |