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<document id="1D25C03ABA3A0A92F5586C7F845AA8B4" ID="10.11646/zootaxa.4184.1.2" ID-DOI="10.11646/zootaxa.4184.1.2" ID-GBIF-Dataset="6a80d4c8-9142-43f3-8e2f-ad72423fb244" ID-ISSN="1175-5326" ID-Zenodo-Dep="164467" ID-ZooBank="396B3FCE-089E-4661-A6D0-FE464D4AAA71" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478157353779" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2016" docId="039E87C6FFFDE467D8BFD57EFC2AF9DA" docLanguage="en" docName="zootaxa.4184.1.2.pdf" docOrigin="Zootaxa 4184 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptenidium gruenbergae Darby, 2016, sp.n." docType="treatment" docUuid="5234B5F3-D1FB-4189-85B6-46CE1E4AD79E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="46" masterDocId="FFA7FFBEFFF8E462D828D61AFFECFFFD" masterDocTitle="New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae)" masterLastPageNumber="51" masterPageNumber="41" pageNumber="46" updateTime="1698391745262" updateUser="plazi">
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<mods:title id="C6CFCBBCC864584AA3DAB2F0C4AD820C">New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae)</mods:title>
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<mods:namePart id="8716FC8EDB05E1FBF355BC7DE31B5FCE">Darby, Michael</mods:namePart>
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<mods:date id="40647C79C637E3AFC3C5ED631B7D1A51">2016</mods:date>
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<treatment id="039E87C6FFFDE467D8BFD57EFC2AF9DA" ID-DOI="http://doi.org/10.5281/zenodo.6074565" ID-GBIF-Taxon="124562727" ID-Zenodo-Dep="6074565" LSID="urn:lsid:zoobank.org:act:5234B5F3-D1FB-4189-85B6-46CE1E4AD79E" httpUri="http://treatment.plazi.org/id/039E87C6FFFDE467D8BFD57EFC2AF9DA" lastPageNumber="46" pageId="5" pageNumber="46">
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<paragraph id="8B8836D0FFFDE467D8BFD57EFE1BFC83" blockId="5.[151,503,868,927]" box="[151,503,868,894]" pageId="5" pageNumber="46">
<heading id="D0C081BCFFFDE467D8BFD57EFE1BFC83" bold="true" box="[151,503,868,894]" fontSize="11" level="1" pageId="5" pageNumber="46" reason="1">
<emphasis id="B943EAC2FFFDE467D8BFD57EFE1BFC83" bold="true" box="[151,503,868,894]" pageId="5" pageNumber="46">
<taxonomicName id="4C374D53FFFDE467D8BFD57EFE59FC83" ID-CoL="6WLD2" box="[151,437,868,894]" class="Insecta" family="Ptiliidae" genus="Ptenidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="gruenbergae" status="sp. nov.">
<emphasis id="B943EAC2FFFDE467D8BFD57EFE59FC83" bold="true" box="[151,437,868,894]" italics="true" pageId="5" pageNumber="46">Ptenidium gruenbergae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27057B9FFFDE467D994D57EFE1BFC83" box="[444,503,868,894]" pageId="5" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836D0FFFDE467D8BFD59CFEC6FC62" blockId="5.[151,503,868,927]" box="[151,298,902,927]" pageId="5" pageNumber="46">
(
<figureCitation id="130C2A55FFFDE467D8B7D59CFECEFC62" box="[159,290,902,927]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Figs. 6 AJ</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D655BFFFDE467D8BFD5D4FDF5FAD6" pageId="5" pageNumber="46" type="description">
<paragraph id="8B8836D0FFFDE467D8BFD5D4FBF9FB1E" blockId="5.[151,1437,974,1575]" pageId="5" pageNumber="46">
Length
<quantity id="4CCF9B35FFFDE467D8DDD5D5FE8FFC1B" box="[245,355,975,999]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="46" unit="mm" value="0.8">0.80 mm</quantity>
. Dark, shining, appearing polished, with very short, barely visible, scattered pubescence. Antennomeres 38 length
<quantity id="4CCF9B35FFFDE467D9EFD5E9FDDCFBF7" box="[455,560,1011,1035]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="46" unit="mm" value="0.33">0.33 mm</quantity>
,
<figureCitation id="130C2A55FFFDE467DA17D5E8FD74FBF6" box="[575,664,1010,1035]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6B</figureCitation>
. Head with longer pubescence and a group of much longer setae medially on the anterior margin, the central stouter ones appearing to form a bifid process,
<figureCitation id="130C2A55FFFDE467DC8ED20CFB11FBD2" box="[1190,1277,1046,1071]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6E</figureCitation>
. Width across eyes
<quantity id="4CCF9B35FFFDE467D8F9D221FED5FBAF" box="[209,313,1083,1107]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="46" unit="mm" value="0.32">0.32 mm</quantity>
. Mentum with rounded lateral margins,
<figureCitation id="130C2A55FFFDE467DB28D220FCBBFBAE" box="[768,855,1082,1107]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6G</figureCitation>
. Pronotum length
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, width
<quantity id="4CCF9B35FFFDE467DCCAD220FAA5FBAF" box="[1250,1353,1082,1107]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="46" unit="mm" value="0.45">0.45 mm</quantity>
, with a small group of setae on the posterior margin at the corners,
<figureCitation id="130C2A55FFFDE467DB47D244FC21FB8A" box="[879,973,1118,1143]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6H</figureCitation>
; without any fovea on the anterior and posterior margins. Scutellum with a depression at the anterior corners and a short median keel with rounded margins,
<figureCitation id="130C2A55FFFDE467D8D6D2BCFEA2FB42" box="[254,334,1190,1215]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6J</figureCitation>
. Elytra length
<quantity id="4CCF9B35FFFDE467D9C7D2BCFDB9FB43" box="[495,597,1190,1215]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="46" unit="mm" value="0.54">0.54 mm</quantity>
, width
<quantity id="4CCF9B35FFFDE467DA8FD2BDFCE2FB43" box="[679,782,1191,1215]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="5" pageNumber="46" unit="mm" value="0.47">0.47 mm</quantity>
. Proventral process parallel sided, rounded posteriorly and with a shallow median depression,
<figureCitation id="130C2A55FFFDE467DA08D2D0FD9EFB1E" box="[544,626,1226,1251]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">
Fig.
<specimenCount id="9D31FD59FFFDE467DA7CD2D0FD9EFB1E" box="[596,626,1226,1251]" pageId="5" pageNumber="46" type="generic">6F</specimenCount>
</figureCitation>
. Mesoventral collar and keel,
<figureCitation id="130C2A55FFFDE467DBEBD2D0FBFDFB1E" box="[963,1041,1226,1251]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">
Fig.
<date id="FF891010FFFDE467DBDED2D0FBFDFB1E" box="[1014,1041,1226,1251]" pageId="5" pageNumber="46">6I</date>
</figureCitation>
.
</paragraph>
<paragraph id="8B8836D0FFFDE467D8EFD2F4FDE9FAFA" blockId="5.[151,1437,974,1575]" box="[199,517,1262,1287]" pageId="5" pageNumber="46">
<emphasis id="B943EAC2FFFDE467D8EFD2F4FEEDFAFB" box="[199,257,1262,1286]" italics="true" pageId="5" pageNumber="46">Male</emphasis>
: aedeagus
<figureCitation id="130C2A55FFFDE467D955D2F4FE25FAFA" box="[381,457,1262,1287]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Figs. 6</figureCitation>
Ca/b.
</paragraph>
<paragraph id="8B8836D0FFFDE467D8EFD309FDF5FAD6" blockId="5.[151,1437,974,1575]" box="[199,537,1298,1323]" pageId="5" pageNumber="46">
<emphasis id="B943EAC2FFFDE467D8EFD309FEF7FAD6" box="[199,283,1299,1323]" italics="true" pageId="5" pageNumber="46">Female</emphasis>
: spermatheca
<figureCitation id="130C2A55FFFDE467D994D308FDF9FAD6" box="[444,533,1298,1323]" captionStart="FIGURE 6. A J" captionStartId="6.[151,250,1661,1683]" captionTargetBox="[150,1436,220,1626]" captionTargetId="figure@6.[126,1460,195,1639]" captionTargetPageId="6" captionText="FIGURE 6. A J. Ptenidium (peruvium) gruenbergae sp. n.. A. habitus; B. antennomeres 3 11; Ca / b. dorsal and side views of aedeagus; D. spermatheca; E. head of male x 910; F. proventral keel x 820; G. mentum and submentum x 920; H. hind angle of pronotum showing tuft of setae x 1220; I, mesoventrum showing mesoventral keel and collar with bifurcate rearwards projecting medial area x 485; J. scutellum x 810." httpUri="https://zenodo.org/record/164472/files/figure.png" pageId="5" pageNumber="46">Fig. 6D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D655BFFFDE467D8EFD32CFC4BFAB2" box="[199,935,1334,1359]" pageId="5" pageNumber="46" type="etymology">
<paragraph id="8B8836D0FFFDE467D8EFD32CFC4BFAB2" blockId="5.[151,1437,974,1575]" box="[199,935,1334,1359]" pageId="5" pageNumber="46">
<emphasis id="B943EAC2FFFDE467D8EFD32CFEAAFAB2" bold="true" box="[199,326,1334,1359]" pageId="5" pageNumber="46">Etymology</emphasis>
. Named after the collector Brigitte Grünberg (Berlin).
</paragraph>
</subSubSection>
<subSubSection id="C32D655BFFFDE467D8EFD340FD00FA46" pageId="5" pageNumber="46" type="diagnosis">
<paragraph id="8B8836D0FFFDE467D8EFD340FD00FA46" blockId="5.[151,1437,974,1575]" pageId="5" pageNumber="46">
<emphasis id="B943EAC2FFFDE467D8EFD340FED6FA8E" bold="true" box="[199,314,1370,1395]" pageId="5" pageNumber="46">Diagnosis</emphasis>
. The only other described species in this subgenus, which is typified by the patches of setae on the pronotal posterior angles and the form of the genitalia, are from
<collectingCountry id="F3207640FFFDE467DB46D365FC49FA6B" box="[878,933,1407,1430]" name="Peru" pageId="5" pageNumber="46">Peru</collectingCountry>
.
<taxonomicName id="4C374D53FFFDE467DB87D365FBB5FA6B" box="[943,1113,1406,1430]" class="Insecta" family="Ptiliidae" genus="Ptenidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="gruenbergae">
<emphasis id="B943EAC2FFFDE467DB87D365FBB5FA6B" box="[943,1113,1406,1430]" italics="true" pageId="5" pageNumber="46">P. gruenbergae</emphasis>
</taxonomicName>
can be separated from those by the group of setae on the front margin of the head.
</paragraph>
</subSubSection>
<subSubSection id="C32D655BFFFDE467D8EFD3DCFC2AF9DA" pageId="5" pageNumber="46" type="materials_examined">
<paragraph id="8B8836D0FFFDE467D8EFD3DCFC2AF9DA" blockId="5.[151,1437,974,1575]" pageId="5" pageNumber="46">
<materialsCitation id="3B5F3C8DFFFDE467D8EFD3DCFF1FF9DA" ID-GBIF-Occurrence="1324562317" collectingDate="2012-10-09" collectionCode="INBio" collectorName="Rincon de Osa" country="Costa Rica" elevation="200" latitude="83.29" location="Costa Rica" pageId="5" pageNumber="46" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B943EAC2FFFDE467D8EFD3DCFEA8FA22" bold="true" box="[199,324,1478,1503]" pageId="5" pageNumber="46">
<typeStatus id="548C8872FFFDE467D8EFD3DCFEEEFA22" box="[199,258,1478,1503]" pageId="5" pageNumber="46">Type</typeStatus>
data
</emphasis>
.
<typeStatus id="548C8872FFFDE467D970D3DCFE29FA22" box="[344,453,1478,1503]" pageId="5" pageNumber="46" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D31FD59FFFDE467D9FDD3DDFE00FA23" box="[469,492,1479,1502]" pageId="5" pageNumber="46" type="male"></specimenCount>
,
<collectingCountry id="F3207640FFFDE467D9D7D3DDFD6EFA22" box="[511,642,1478,1503]" name="Costa Rica" pageId="5" pageNumber="46">Costa Rica</collectingCountry>
, Osa Peninsula, Road, Bahia Drake to
<collectorName id="26C25306FFFDE467DC49D3DCFAF9FA22" box="[1121,1301,1478,1503]" pageId="5" pageNumber="46">Rincon de Osa</collectorName>
, forest and plantations,
<collectingMethod id="52764EC7FFFDE467D90ED3F1FE94F9FF" box="[294,376,1515,1538]" pageId="5" pageNumber="46">car net</collectingMethod>
devart, 8.4142
<geoCoordinate id="EE035017FFFDE467DA6FD3F1FD44F9FE" box="[583,680,1515,1539]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="555" value="83.29">N 83.29</geoCoordinate>
40W,
<quantity id="4CCF9B35FFFDE467DAD7D3F1FCB1F9FE" box="[767,861,1515,1539]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="4.0" metricValueMin="0.0" pageId="5" pageNumber="46" unit="m" value="200.0" valueMax="400.0" valueMin="0.0">
<elevation id="001AD1E3FFFDE467DAD7D3F1FCB1F9FE" box="[767,861,1515,1539]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="4.0" metricValueMin="0.0" pageId="5" pageNumber="46" unit="m" value="200.0" valueMax="400.0" valueMin="0.0">0400m</elevation>
</quantity>
,
<date id="FF891010FFFDE467DB47D3F1FC3FF9FE" box="[879,979,1515,1539]" pageId="5" pageNumber="46" value="2012-10-09">
<collectingDate id="EFCDE9F8FFFDE467DB47D3F1FC3FF9FE" box="[879,979,1515,1539]" pageId="5" pageNumber="46" value="2012-10-09">9.x.2012</collectingDate>
</date>
, 15001700 hrs, Schülke &amp; Grünberg (
<collectionCode id="ED26AE15FFFDE467D8B7D014FF07F9DA" box="[159,235,1550,1575]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="46">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5F3C8DFFFDE467D8D5D015FC2EF9DA" ID-GBIF-Occurrence="1324562327" box="[253,962,1550,1575]" collectingDate="2012-10-09" collectionCode="INBio, BMNH" collectorName="Rincon de Osa" country="Costa Rica" elevation="200" latitude="83.29" location="Costa Rica" pageId="5" pageNumber="46" specimenCount="1" typeStatus="paratype">
<typeStatus id="548C8872FFFDE467D8D5D015FE82F9DA" box="[253,366,1551,1575]" pageId="5" pageNumber="46" type="paratype">Paratypes</typeStatus>
: 6 examples, same data as holotype (
<collectionCode id="ED26AE15FFFDE467DB23D014FCB9F9DA" box="[779,853,1550,1575]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="46">INBio</collectionCode>
,
<collectionCode id="ED26AE15FFFDE467DB49D015FC56F9DA" box="[865,954,1551,1575]" pageId="5" pageNumber="46">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>