267 lines
36 KiB
XML
267 lines
36 KiB
XML
<document id="26857C12B292EA4D051334F8A2C4C675" ID-CLB-Dataset="300538" ID-DOI="10.17161/jom.i103.15655" ID-GBIF-Dataset="9597d08b-2c12-466b-a7fe-efe79569be3b" ID-ISSN="2325-4467" ID-Zenodo-Dep="13146053" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1722455600613" checkinUser="felipe" docAuthor="Engel, Michael S. & Davis, Steven R." docDate="2021" docId="B82B87C2635EF06CF1E2FD8A5F6354CB" docLanguage="en" docName="JMelittology.2021.103.1-52.pdf" docOrigin="Journal of Melittology 2021 (103)" docSource="http://dx.doi.org/10.17161/jom.i103.15655" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Haidomelikertes uraeus Engel 2021, new species" docType="treatment" docVersion="4" lastPageNumber="25" masterDocId="4412FFBA6349F076F07BFE0B5C595119" masterDocTitle="New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="22" updateTime="1722605887797" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="4007B4C2089BBAAFFBA8994F9AD61DE4">New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)</mods:title>
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<mods:name id="E16B19FBAA883C5E3D9EA0D62437FA65" type="personal">
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<mods:namePart id="B60868316DFF5800A2A461D20704C560">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="518BCD7FF25A542F607D24EB7A705A44">Division of Entomology, Natural History Museum and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). & Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA (mengel @ amnh. org, sdavis @ amnh. org).</mods:affiliation>
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<mods:namePart id="009873392E1AF1D99FB6CA45C47A0BF1">Davis, Steven R.</mods:namePart>
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<mods:affiliation id="3E11B2C203A18B40F82EDFB894BFA01E">Division of Entomology, Natural History Museum and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu).</mods:affiliation>
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<mods:title id="D7D277912B61D6F0C5128269BC4631F3">Journal of Melittology</mods:title>
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<mods:part id="AFF050CE478E10E62C41ADCEB4838F70">
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<mods:date id="70E75F588226A23C2F3BA51FE9396A30">2021</mods:date>
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<mods:number id="1FF3C3241CF50754E04DE47D6391934B">2021-07-06</mods:number>
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<mods:number id="40D8A233474D932AFEF0B6AD7E520FEE">2021</mods:number>
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<mods:number id="5C65D291DAA3130D60C5A37090973259">103</mods:number>
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<treatment id="B82B87C2635EF06CF1E2FD8A5F6354CB" ID-DOI="http://doi.org/10.5281/zenodo.13170225" ID-Zenodo-Dep="13170225" LSID="urn:lsid:plazi:treatment:B82B87C2635EF06CF1E2FD8A5F6354CB" httpUri="http://treatment.plazi.org/id/B82B87C2635EF06CF1E2FD8A5F6354CB" lastPageId="26" lastPageNumber="25" pageId="23" pageNumber="22">
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<heading id="6B7581B8635EF061F1E2FD8A5FCE5280" box="[409,919,897,921]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="22" reason="2">
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<taxonomicName id="F7824D57635EF061F1E2FD8A5F595280" authority="Engel" authorityName="Engel" authorityYear="2021" box="[409,768,897,921]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="uraeus">
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<emphasis id="02F6EAC6635EF061F1E2FD8A5EEA5280" bold="true" box="[409,691,897,921]" italics="true" pageId="23" pageNumber="22">Haidomelikertes uraeus</emphasis>
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Engel
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</taxonomicName>
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,
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<taxonomicNameLabel id="19C557BD635EF061F371FD895FCE5280" box="[778,919,897,921]" pageId="23" pageNumber="22" rank="species">new species</taxonomicNameLabel>
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</heading>
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<subSubSection id="7898655F635EF061F17AFDA85F6852C0" pageId="23" pageNumber="22" type="description">
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<paragraph id="303D36D4635EF061F17AFDA8587652A0" blockId="23.[257,1071,931,985]" box="[257,1071,931,953]" pageId="23" pageNumber="22">
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ZooBank:
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urn:lsid:zoobank.org:act:
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<uuid id="44240C01635EF061F219FDA8587652A0" box="[610,1071,931,953]" pageId="23" pageNumber="22">74265917-E6B4-44B4-8F45-93D269B0D810</uuid>
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</uri>
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<paragraph id="303D36D4635EF061F184FDCA5F6852C0" blockId="23.[257,1071,931,985]" box="[511,817,961,985]" pageId="23" pageNumber="22">
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(
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<figureCitation id="A8B92A51635EF061F27CFDCA5E3A52C0" box="[519,611,961,985]" captionStart="Figures 19–20" captionStartId="19.[161,240,1292,1314]" captionTargetBox="[160,1168,224,1286]" captionTargetId="figure-173@19.[160,1168,224,1286]" captionTargetPageId="19" captionText="Figures 19–20. Inner (retrolateral) surfaces of metatibiae of Haidomelikertes, new genus, and Aethemelikertes, new genus. 19. Haidomelikertes uraeus, new species, holotype worker (SMF Be 14263a). 20. Aethemelikertes emunctorii, new species, holotype worker (SMF Be 14262a)." figureDoi="http://doi.org/10.5281/zenodo.13146073" httpUri="https://zenodo.org/record/13146073/files/figure.png" pageId="23" pageNumber="22">Figs. 19</figureCitation>
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,
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<figureCitation id="A8B92A51635EF061F215FDCA5ED252C0" box="[622,651,961,985]" captionStart="Figure 21" captionStartId="21.[160,229,1400,1422]" captionTargetBox="[161,1168,224,1394]" captionTargetId="figure-132@21.[160,1168,224,1394]" captionTargetPageId="21" captionText="Figure 21. Holotype worker of Haidomelikertes uraeus, new genus and species, in Eocene (Bartonian) Baltic amber (SMF Be 14263a)." figureDoi="http://doi.org/10.5281/zenodo.13146077" httpUri="https://zenodo.org/record/13146077/files/figure.png" pageId="23" pageNumber="22">21</figureCitation>
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,
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<figureCitation id="A8B92A51635EF061F2EDFDCA5EEA52C0" box="[662,691,961,985]" captionStart="Figure 22" captionStartId="22.[160,229,1083,1105]" captionTargetBox="[160,1168,224,1076]" captionTargetId="figure-226@22.[160,1168,224,1076]" captionTargetPageId="22" captionText="Figure 22. X-ray µCT scan of holotype worker of Haidomelikertes uraeus, new genus and species (SMF Be 14263a), oblique frontolateral view. Note that scan resolution was insufficient to resolve most setae." figureDoi="http://doi.org/10.5281/zenodo.13146079" httpUri="https://zenodo.org/record/13146079/files/figure.png" pageId="23" pageNumber="22">22</figureCitation>
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,
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<figureCitation id="A8B92A51635EF061F2C5FDCA5F7052C0" box="[702,809,961,985]" captionStart-0="Figures 23–26" captionStart-1="Figures 27–28" captionStartId-0="24.[160,241,1266,1288]" captionStartId-1="25.[160,241,831,853]" captionTargetBox-0="[160,1168,224,1258]" captionTargetBox-1="[160,1168,224,824]" captionTargetId-0="figure-175@24.[160,1168,224,1258]" captionTargetId-1="figure-366@25.[160,1168,224,824]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="Figures 23–26. X-ray µCT scans of heads of holotype workers of Haidomelikertes proboscidea (Engel), new combination (AMNH B-JH96), and H. uraeus, new genus and species (SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 23. Facial view of H. proboscidea. 24. Facial view of H. uraeus. 25. Oblique profile of H. proboscidea. 26. Oblique profile of H. uraeus. Note that scan resolution was insufficient to resolve most setae." captionText-1="Figures 27–28. Oblique facial views of holotype worker of Haidomelikertes uraeus, new genus and species (SMF Be 14263a). 27. Slightly dorsal left oblique; white arrow points to apex of facial protuberance. 28. Slightly ventral right oblique." figureDoi-0="http://doi.org/10.5281/zenodo.13146081" figureDoi-1="http://doi.org/10.5281/zenodo.13146083" httpUri-0="https://zenodo.org/record/13146081/files/figure.png" httpUri-1="https://zenodo.org/record/13146083/files/figure.png" pageId="23" pageNumber="22">24, 26–28</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="7898655F635EF061F0ABFA0A5FB255A0" pageId="23" pageNumber="22" type="diagnosis">
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<paragraph id="303D36D4635EF061F0ABFA0A5FB255A0" blockId="23.[160,1175,1025,1785]" pageId="23" pageNumber="22">
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DIAGNOSIS: This species is distinguished from its congener by the form of the clypeal protrusion, which extends only slightly above the upper torular tangent, and its forward-projecting surface only extends as a lamella, no longer than a torular diameter (
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<figureCitation id="A8B92A51635EF061F0A1FA6A5DDF5560" box="[218,390,1121,1145]" captionStart-0="Figures 23–26" captionStart-1="Figures 27–28" captionStartId-0="24.[160,241,1266,1288]" captionStartId-1="25.[160,241,831,853]" captionTargetBox-0="[160,1168,224,1258]" captionTargetBox-1="[160,1168,224,824]" captionTargetId-0="figure-175@24.[160,1168,224,1258]" captionTargetId-1="figure-366@25.[160,1168,224,824]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="Figures 23–26. X-ray µCT scans of heads of holotype workers of Haidomelikertes proboscidea (Engel), new combination (AMNH B-JH96), and H. uraeus, new genus and species (SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 23. Facial view of H. proboscidea. 24. Facial view of H. uraeus. 25. Oblique profile of H. proboscidea. 26. Oblique profile of H. uraeus. Note that scan resolution was insufficient to resolve most setae." captionText-1="Figures 27–28. Oblique facial views of holotype worker of Haidomelikertes uraeus, new genus and species (SMF Be 14263a). 27. Slightly dorsal left oblique; white arrow points to apex of facial protuberance. 28. Slightly ventral right oblique." figureDoi-0="http://doi.org/10.5281/zenodo.13146081" figureDoi-1="http://doi.org/10.5281/zenodo.13146083" httpUri-0="https://zenodo.org/record/13146081/files/figure.png" httpUri-1="https://zenodo.org/record/13146083/files/figure.png" pageId="23" pageNumber="22">Figs. 24, 26–28</figureCitation>
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). The neck of the clypeal protrusion is also not narrowed above the supraclypeal area and between the antennal toruli relative to its apical width (
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<figureCitation id="A8B92A51635EF061F448FA8A58D05580" box="[1075,1161,1153,1177]" captionStart="Figures 23–26" captionStartId="24.[160,241,1266,1288]" captionTargetBox="[160,1168,224,1258]" captionTargetId="figure-175@24.[160,1168,224,1258]" captionTargetPageId="24" captionText="Figures 23–26. X-ray µCT scans of heads of holotype workers of Haidomelikertes proboscidea (Engel), new combination (AMNH B-JH96), and H. uraeus, new genus and species (SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum. 23. Facial view of H. proboscidea. 24. Facial view of H. uraeus. 25. Oblique profile of H. proboscidea. 26. Oblique profile of H. uraeus. Note that scan resolution was insufficient to resolve most setae." figureDoi="http://doi.org/10.5281/zenodo.13146081" httpUri="https://zenodo.org/record/13146081/files/figure.png" pageId="23" pageNumber="22">Fig. 24</figureCitation>
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). Additionally, this species is about a full 20% shorter than
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<taxonomicName id="F7824D57635EF061F33DFAA95FBD55A0" baseAuthorityName="Engel" baseAuthorityYear="2001" box="[838,996,1185,1209]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="proboscidea">
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<emphasis id="02F6EAC6635EF061F33DFAA95FBD55A0" box="[838,996,1185,1209]" italics="true" pageId="23" pageNumber="22">H. proboscidea</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="7898655F635EF06CF0ABFACA5D765428" lastPageId="26" lastPageNumber="25" pageId="23" pageNumber="22" type="description">
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<paragraph id="303D36D4635EF06EF0ABFACA5D6D5705" blockId="23.[160,1175,1025,1785]" lastBlockId="24.[160,1169,1476,1756]" lastPageId="24" lastPageNumber="23" pageId="23" pageNumber="22">
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DESCRIPTION: ⚲: Total body length (as preserved)
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<quantity id="F77A9B31635EF061F36BFACA5F3055C0" box="[784,873,1217,1241]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="23" pageNumber="22" unit="mm" value="3.3">3.3 mm</quantity>
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; forewing length (as preserved)
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<quantity id="F77A9B31635EF061F179FAEA5D0655E0" box="[258,351,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="22" unit="mm" value="3.5">3.5 mm</quantity>
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. Head as long as wide, length (summit of vertex to clypeal apical margin)
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<quantity id="F77A9B31635EF061F171FB0A5D2C5400" box="[266,373,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="22" unit="mm" value="1.3">1.30 mm</quantity>
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, width (maximum width across compound eyes)
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<quantity id="F77A9B31635EF061F3A0FB0A581F5400" box="[987,1094,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="22" unit="mm" value="1.3">1.30 mm</quantity>
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; compound eye length
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<quantity id="F77A9B31635EF061F1F8FB2A5DB75420" box="[387,494,1313,1337]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="22" unit="mm" value="0.8">0.80 mm</quantity>
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; upper interorbital distance
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<quantity id="F77A9B31635EF061F336FB2A5FE05420" box="[845,953,1313,1337]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="23" pageNumber="22" unit="mm" value="0.94">0.94 mm</quantity>
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, lower interorbital distance
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<quantity id="F77A9B31635EF061F157FB4A5DCE5440" box="[300,407,1345,1369]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="22" unit="mm" value="0.8">0.80 mm</quantity>
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; inner ocular margins largely straight, with faint arch slightly above midlength, converging below; ocellar triangle broad, interocellar distance
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<quantity id="F77A9B31635EF061F41AFB6A5C8B5480" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="mm" value="0.4">0.40 mm</quantity>
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. Scape length
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<quantity id="F77A9B31635EF061F1F2FB8A5DAA5480" box="[393,499,1409,1433]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="22" unit="mm" value="0.5">0.50 mm</quantity>
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, shorter than torulocellar distance. Mandible elongate, length
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<quantity id="F77A9B31635EF061F088FBAA5D0554A0" box="[243,348,1441,1465]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="23" pageNumber="22" unit="mm" value="0.65">0.65 mm</quantity>
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; labrum broader than long, basal width
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<quantity id="F77A9B31635EF061F344FBAA5FF154A0" box="[831,936,1441,1465]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="23" pageNumber="22" unit="mm" value="0.46">0.46 mm</quantity>
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, medial length
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<quantity id="F77A9B31635EF061F41AFBAA5C8B54C0" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="23" pageNumber="22" unit="mm" value="0.27">0.27 mm</quantity>
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. Pronotum short, declivitous, without defined transverse dorsal ridge, therefore sloping continuously from mesoscutal border; pronotal lateral ridge absent; mesoscutum anterior border broadly rounded, anterior lip gently curving to meet posterior pronotal margin, not overhanging pronotal border; mesoscutum medial length
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<quantity id="F77A9B31635EF061F41AF82A5C8B5740" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="23" pageNumber="22" unit="mm" value="0.77">0.77 mm</quantity>
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; intertegular distance
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<quantity id="F77A9B31635EF061F1A5F84A5E1E5740" box="[478,583,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="23" pageNumber="22" unit="mm" value="0.37">0.37 mm</quantity>
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; mesoscutellum medial length
|
||
<quantity id="F77A9B31635EF061F3C6F84A587F5740" box="[957,1062,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="23" pageNumber="22" unit="mm" value="0.31">0.31 mm</quantity>
|
||
. Metatibia slender, length
|
||
<quantity id="F77A9B31635EF061F1F0F86A5DAF5760" box="[395,502,1633,1657]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="22" unit="mm" value="1.2">1.20 mm</quantity>
|
||
, maximum width
|
||
<quantity id="F77A9B31635EF061F2A3F86A5F1A5760" box="[728,835,1633,1657]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="23" pageNumber="22" unit="mm" value="0.34">0.34 mm</quantity>
|
||
. Forewing with basal vein (1M) confluent with 1cu-a, pterostigma longer than wide, maximum width just basad midlength, margin inside marginal cell sloping to costal margin; marginal cell acutely rounded apically; basal vein (1M) straight; three submarginal cells (1rs-m present), second submarginal cell trapezoidal, posterior border strongly angled posteriorly; 2Rs straight; 3Rs shorter than r-rs, together slightly longer than 4Rs; 1rs-m faintly arched, 2rs-m arched apicad in posterior half; wing membranes hyaline clear; veins dark brown.
|
||
</paragraph>
|
||
<caption id="64FD665C6351F06EF0DBFAF95D4F5481" ID-DOI="http://doi.org/10.5281/zenodo.13146081" ID-Zenodo-Dep="13146081" httpUri="https://zenodo.org/record/13146081/files/figure.png" pageId="24" pageNumber="23" startId="24.[160,241,1266,1288]" targetBox="[160,1168,224,1258]" targetPageId="24" targetType="figure">
|
||
<paragraph id="303D36D46351F06EF0DBFAF95D4F5481" blockId="24.[160,1169,1266,1432]" pageId="24" pageNumber="23">
|
||
<emphasis id="02F6EAC66351F06EF0DBFAF95D64541E" bold="true" box="[160,317,1266,1288]" pageId="24" pageNumber="23">Figures 23–26.</emphasis>
|
||
X-ray µCT scans of heads of holotype workers of
|
||
<taxonomicName id="F7824D576351F06EF300FAF858C85411" baseAuthorityName="Engel" baseAuthorityYear="2001" box="[891,1169,1267,1288]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="proboscidea">
|
||
<emphasis id="02F6EAC66351F06EF300FAF858C85411" box="[891,1169,1267,1288]" italics="true" pageId="24" pageNumber="23">Haidomelikertes proboscidea</emphasis>
|
||
</taxonomicName>
|
||
(Engel),
|
||
<taxonomicNameLabel id="19C557BD6351F06EF087FB1B5DEE543C" box="[252,439,1295,1317]" pageId="24" pageNumber="23" rank="species">new combination</taxonomicNameLabel>
|
||
(AMNH B-JH96), and
|
||
<taxonomicName id="F7824D576351F06EF2CDFB1B5F41543C" authorityName="Engel & Davis" authorityYear="2021" box="[694,792,1296,1317]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="uraeus" status="new genus and species">
|
||
<emphasis id="02F6EAC66351F06EF2CDFB1B5F41543C" box="[694,792,1296,1317]" italics="true" pageId="24" pageNumber="23">H. uraeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="19C557BD6351F06EF353FB1B587A543C" box="[808,1059,1295,1317]" pageId="24" pageNumber="23" rank="species">new genus and species</taxonomicNameLabel>
|
||
(SMF (Be 14263a), in Baltic amber; yellow highlights disc and modification of clypeus, orange highlights labrum.
|
||
<emphasis id="02F6EAC66351F06EF17BFB425D465447" bold="true" box="[256,287,1353,1374]" pageId="24" pageNumber="23">23.</emphasis>
|
||
Facial view of
|
||
<taxonomicName id="F7824D576351F06EF1C5FB415E145447" baseAuthorityName="Engel" baseAuthorityYear="2001" box="[446,589,1353,1375]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="proboscidea">
|
||
<emphasis id="02F6EAC66351F06EF1C5FB415E145447" box="[446,589,1353,1375]" italics="true" pageId="24" pageNumber="23">H. proboscidea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="02F6EAC66351F06EF224FB425E245447" bold="true" box="[607,637,1353,1374]" pageId="24" pageNumber="23">24.</emphasis>
|
||
Facial view of
|
||
<taxonomicName id="F7824D576351F06EF366FB415F245446" authorityName="Engel & Davis" authorityYear="2021" box="[797,893,1354,1375]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="uraeus">
|
||
<emphasis id="02F6EAC66351F06EF366FB415F245446" box="[797,893,1354,1375]" italics="true" pageId="24" pageNumber="23">H. uraeus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="02F6EAC66351F06EF3EBFB425FF65447" bold="true" box="[912,943,1353,1374]" pageId="24" pageNumber="23">25.</emphasis>
|
||
Oblique profile of
|
||
<taxonomicName id="F7824D576351F06EF403FB415D495462" baseAuthorityName="Engel" baseAuthorityYear="2001" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="proboscidea">
|
||
<emphasis id="02F6EAC66351F06EF403FB415D495462" italics="true" pageId="24" pageNumber="23">H. proboscidea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="02F6EAC66351F06EF15BFB6D5D675462" bold="true" box="[288,318,1382,1403]" pageId="24" pageNumber="23">26.</emphasis>
|
||
Oblique profile of
|
||
<taxonomicName id="F7824D576351F06EF278FB6C5E3B5465" authorityName="Engel & Davis" authorityYear="2021" box="[515,610,1383,1404]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="23" phylum="Arthropoda" rank="species" species="uraeus">
|
||
<emphasis id="02F6EAC66351F06EF278FB6C5E3B5465" box="[515,610,1383,1404]" italics="true" pageId="24" pageNumber="23">H. uraeus</emphasis>
|
||
</taxonomicName>
|
||
. Note that scan resolution was insufficient to resolve most setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="303D36D46351F06EF0ABF82F5D3357C5" blockId="24.[160,1169,1476,1756]" pageId="24" pageNumber="23">Mandible smooth and shining; labrum smooth; clypeal disc finely imbricate; face smooth and shining; mesoscutum and mesoscutellum smooth and shining; mespisternum largely smooth to finely imbricate; basal area of propodeum glabrous, smooth, shining; legs generally smooth; metatibial corbicular surface smooth and shining; metasomal terga smooth and shining, with minute punctures at setal bases; sterna faintly imbricate.</paragraph>
|
||
<caption id="64FD665C6350F06FF0DBFD345EC45297" ID-DOI="http://doi.org/10.5281/zenodo.13146083" ID-Zenodo-Dep="13146083" httpUri="https://zenodo.org/record/13146083/files/figure.png" pageId="25" pageNumber="24" startId="25.[160,241,831,853]" targetBox="[160,1168,224,824]" targetPageId="25" targetType="figure">
|
||
<paragraph id="303D36D46350F06FF0DBFD345EC45297" blockId="25.[160,1168,831,910]" pageId="25" pageNumber="24">
|
||
<emphasis id="02F6EAC66350F06FF0DBFD345D62524D" bold="true" box="[160,315,831,853]" pageId="25" pageNumber="24">Figures 27–28.</emphasis>
|
||
Oblique facial views of holotype worker of
|
||
<taxonomicName id="F7824D576350F06FF35CFD345855524C" authorityName="Engel & Davis" authorityYear="2021" box="[807,1036,831,853]" class="Insecta" family="Apidae" genus="Haidomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="24" phylum="Arthropoda" rank="species" species="uraeus" status="new genus and species">
|
||
<emphasis id="02F6EAC66350F06FF35CFD345855524C" box="[807,1036,831,853]" italics="true" pageId="25" pageNumber="24">Haidomelikertes uraeus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="19C557BD6350F06FF460FD345D40526B" pageId="25" pageNumber="24" rank="species">new genus and species</taxonomicNameLabel>
|
||
(SMF Be 14263a).
|
||
<emphasis id="02F6EAC66350F06FF1A0FD575DA15268" bold="true" box="[475,504,860,881]" pageId="25" pageNumber="24">27.</emphasis>
|
||
Slightly dorsal left oblique; white arrow points to apex of facial protuberance.
|
||
<emphasis id="02F6EAC66350F06FF13BFD735D075294" bold="true" box="[320,350,888,909]" pageId="25" pageNumber="24">28.</emphasis>
|
||
Slightly ventral right oblique.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="303D36D46350F06CF0ABFDB05CA155C8" blockId="25.[160,1168,955,1779]" lastBlockId="26.[160,1168,954,1490]" lastPageId="26" lastPageNumber="25" pageId="25" pageNumber="24">
|
||
Pubescence generally sparse; mandible with sparse, minute, suberect, simple setae on outer surface proximally, longer, erect, simple setae scatered along length of lower margin; labrum with sparsely scatered, short, suberect to erect setae on disc, laterally with numerous, elongate, erect, simple setae forming fringe along margins; clypeus largely without setae on disc, some short, fine, suberect setae in apicolateral areas lateral to and in angled area lateral to labrum; clypeal extension some scatered, short, fine, erect, simple setae on the upper and lower surfaces of the anterior extension; face with sparsely scatered, short, erect, simple setae intermixed with some minutely branched setae; scape with minute, appressed, simple setae scatered along length; vertex with similar setae as those on face along with bristle-like setae that are about as long as median ocellar diameter or often slightly shorter; mesoscutum with sparsely scatered, erect, simple bristles intermixed with finer, erect, plumose setae, such plumose setae more numerous and particularly so anteriorly; mesoscutellum as on mesoscutum except lateral and posterior margins with more abundant erect bristles and numerous plumose setae; mespisternum with scatered, short, decumbent, simple setae; pro- and mesocoxae, pro- and mesotrochanters, and pro- and mesofemora with sparse, minute, largely appressed, simple setae; pro- and mesotibiae with some longer, erect, simple or branched setae, particularly apically, and more so on outer surface of mesotibia; pro- and mesotarsi with abundant, longer, erect, simple bristles; metacoxa, metatrochanter, and metafemur similar to corresponding podites on midleg, except inner apical surface of metafemur with small ovoid patch of keirotrichia, finer and shorter than those of metatibia; metatibia with posterior margin with fringe of elongate, plumose (branches minute and along length of setal rachis) setae, setae longer than metatibial width, anterior margin with sparsely scatered simple setae and bristles, corbicular surface with sparsely scatered, erect, fine, simple setae; inner surface with keirotrichiate zone covering most of surface except exceedingly narrow, posterior, slightly depressed glabrate zone and rectangular apical glabrate zone, length of apical glabrate zone slightly less than apical width of metatibia; rastellum composed of stiff bristles along entire inner apical width of metatibia; metabasitarsus with sparse, simple setae on outer surface, inner surface and margins with abundant, elongate, erect to suberect, simple bristles; metasomal terga with sparse, minute, largely appressed, simple setae, such setae especially sparse on basal terga and becoming progressively more numerous on terga
|
||
<collectionCode id="5693AE116353F06CF354FA715F155588" box="[815,844,1146,1169]" pageId="26" pageNumber="25">IV</collectionCode>
|
||
–
|
||
<collectionCode id="5693AE116353F06CF320FA715F215588" box="[859,888,1146,1169]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="26" pageNumber="25">VI</collectionCode>
|
||
; sterna with fine, erect, simple setae on apical half to third of sterna II–
|
||
<collectionCode id="5693AE116353F06CF2B4FA915EB555A8" box="[719,748,1178,1201]" pageId="26" pageNumber="25">IV</collectionCode>
|
||
, and somewhat shorter on sterna
|
||
<collectionCode id="5693AE116353F06CF406FA9158C955A8" box="[1149,1168,1178,1201]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="25" type="Museum">V</collectionCode>
|
||
and
|
||
<collectionCode id="5693AE116353F06CF0AFFAB15CAD55C8" box="[212,244,1210,1233]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="26" pageNumber="25">VI</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<caption id="64FD665C6353F06CF0DBFD515DDD5294" ID-DOI="http://doi.org/10.5281/zenodo.13146085" ID-Zenodo-Dep="13146085" httpUri="https://zenodo.org/record/13146085/files/figure.png" pageId="26" pageNumber="25" startId="26.[160,231,858,880]" targetBox="[160,1162,224,852]" targetPageId="26" targetType="figure">
|
||
<paragraph id="303D36D46353F06CF0DBFD515DDD5294" blockId="26.[160,1168,858,909]" pageId="26" pageNumber="25">
|
||
<emphasis id="02F6EAC66353F06CF0DBFD515D575276" bold="true" box="[160,270,858,880]" pageId="26" pageNumber="25">Figure 29.</emphasis>
|
||
Two workers of
|
||
<taxonomicName id="F7824D576353F06CF1ADFD515EEA5276" baseAuthorityName="Engel" baseAuthorityYear="2001" box="[470,691,858,879]" class="Insecta" family="Apidae" genus="Amelikertotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="clypeata">
|
||
<emphasis id="02F6EAC66353F06CF1ADFD515EEA5276" box="[470,691,858,879]" italics="true" pageId="26" pageNumber="25">Amelikertotes clypeata</emphasis>
|
||
</taxonomicName>
|
||
(Engel),
|
||
<taxonomicNameLabel id="19C557BD6353F06CF363FD505F8C5269" box="[792,981,858,880]" pageId="26" pageNumber="25" rank="species">new combination</taxonomicNameLabel>
|
||
, in Eocene Baltic amber (SEMC B-147).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="303D36D46353F06CF0ABFAD1586355EB" blockId="26.[160,1168,954,1490]" box="[208,1082,1242,1266]" pageId="26" pageNumber="25">Coloration not preserved, where evident seemingly brown to dark brown.</paragraph>
|
||
<paragraph id="303D36D46353F06CF0ABFAF15D765408" blockId="26.[160,1168,954,1490]" box="[208,303,1274,1300]" pageId="26" pageNumber="25">
|
||
<specimenCount id="2684FD5D6353F06CF0ABFAF15CBD540D" box="[208,228,1274,1300]" count="1" pageId="26" pageNumber="25" type="female">♀</specimenCount>
|
||
:
|
||
<emphasis id="02F6EAC66353F06CF08AFAF15D715408" box="[241,296,1274,1297]" italics="true" pageId="26" pageNumber="25">Latet</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="303D36D46353F06CF0ABFB115D765428" blockId="26.[160,1168,954,1490]" box="[208,303,1306,1332]" pageId="26" pageNumber="25">
|
||
<specimenCount id="2684FD5D6353F06CF0ABFB115CBD542D" box="[208,228,1306,1332]" count="1" pageId="26" pageNumber="25" type="male">♂</specimenCount>
|
||
:
|
||
<emphasis id="02F6EAC66353F06CF08AFB115D715428" box="[241,296,1306,1329]" italics="true" pageId="26" pageNumber="25">Latet</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7898655F6353F06CF0ABFB315FFD546B" pageId="26" pageNumber="25" type="materials_examined">
|
||
<paragraph id="303D36D46353F06CF0ABFB315FFD546B" blockId="26.[160,1168,954,1490]" pageId="26" pageNumber="25">
|
||
<materialsCitation id="80EA3C896353F06CF0ABFB315FFD546B" collectionCode="SMF" pageId="26" pageNumber="25" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="EF3988766353F06CF0ABFB315D11544B" box="[208,328,1338,1362]" pageId="26" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
|
||
: ⚲,
|
||
<collectionCode id="5693AE116353F06CF10AFB315DFE544B" box="[369,423,1338,1362]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="25">SMF</collectionCode>
|
||
Be 14263a (
|
||
<collectionCode id="5693AE116353F06CF24DFB315E28544B" box="[566,625,1338,1362]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="25">SMF</collectionCode>
|
||
); Eocene (Bartonian–Priabonian: refer to note for holotype of
|
||
<taxonomicName id="F7824D576353F06CF12DFB515E245468" authorityName="Engel & Davis" authorityYear="2021" box="[342,637,1370,1394]" class="Insecta" family="Apidae" genus="Aethemelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="25" phylum="Arthropoda" rank="species" species="emunctorii">
|
||
<emphasis id="02F6EAC66353F06CF12DFB515E245468" box="[342,637,1370,1394]" italics="true" pageId="26" pageNumber="25">Aethemelikertes emunctorii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="02F6EAC66353F06CF2F1FB515EA35468" box="[650,762,1370,1394]" italics="true" pageId="26" pageNumber="25">vide supra</emphasis>
|
||
) Baltic amber.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7898655F6353F06CF0ABFB715F6354CB" pageId="26" pageNumber="25" type="etymology">
|
||
<paragraph id="303D36D46353F06CF0ABFB715F6354CB" blockId="26.[160,1168,954,1490]" pageId="26" pageNumber="25">
|
||
ETYMOLOGY: The specific epithet is taken from the Ancient Greek
|
||
<emphasis id="02F6EAC66353F06CF3B7FB715842548B" box="[972,1051,1402,1426]" italics="true" pageId="26" pageNumber="25">ouraîos</emphasis>
|
||
(
|
||
<emphasis id="02F6EAC66353F06CF450FB7258D0548B" box="[1067,1161,1401,1426]" italics="true" pageId="26" pageNumber="25">οὐραῖος</emphasis>
|
||
, uraeus), the stylized, upright rearing cobra used as a symbol of divine authority (symbol of the goddess Wadjet,
|
||
<emphasis id="02F6EAC66353F06CF267FBB15E0054C8" box="[540,601,1464,1490]" italics="true" pageId="26" pageNumber="25">wꜢḏyt</emphasis>
|
||
) in Ancient
|
||
<collectingCountry id="489576446353F06CF297FBB15F6F54CB" box="[748,822,1466,1490]" name="Egypt" pageId="26" pageNumber="25">Egypt</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |