1156 lines
147 KiB
XML
1156 lines
147 KiB
XML
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<mods:title id="1B0CC9A486233C0FC1431A200329B45A">A new species of mossy frog (Anura: Rhacophoridae) from Northeastern Vietnam</mods:title>
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<mods:namePart id="66E191D93678CD5CD5AF99D4EE07ADCB">Ninh, Hoa Thi</mods:namePart>
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<mods:affiliation id="12B7F895977DB7344C7656EF34162FCD">Thai Nguyen University of Education, Thai Nguyen University, 20 Luong Ngoc Quyen, Thai Nguyen, Vietnam.</mods:affiliation>
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<mods:affiliation id="69B89483A384C73CF7BE5C902081D556">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam. & Ministry of Education and Training, 35 Dai Co Viet Road, Hai Ba Trung District, Vietnam. & Center for Biodiversity Conservation and Endangered Species, 541 Nguyen Duy Trinh Street, District 2, Hochiminh City, Vietnam.</mods:affiliation>
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<mods:affiliation id="E342209E8EFA75597E5D4C25CACA22A7">Ministry of Education and Training, 35 Dai Co Viet Road, Hai Ba Trung District, Vietnam.</mods:affiliation>
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<mods:namePart id="1E8ABD6C830F80880D09B289183C80D3">Le, Quyet Khac</mods:namePart>
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<mods:affiliation id="22F1E62E8476403DCD280D05A5451DD8">Center for Biodiversity Conservation and Endangered Species, 541 Nguyen Duy Trinh Street, District 2, Hochiminh City, Vietnam.</mods:affiliation>
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<mods:namePart id="10CD2A33A06D2D6C6CD5EABA88C72FF6">Ziegler, Thomas</mods:namePart>
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<mods:affiliation id="36679E4E3FD6B8A0AF43092064EE27A3">Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany. & Institute of Zoology, University of Cologne, Zülpicher Strasse 47 B, D- 50674 Cologne, Germany.</mods:affiliation>
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<mods:title id="A3BF225FA2A0E6A82828DD1979057F3B">European Journal of Taxonomy</mods:title>
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<mods:part id="06C05D9F45D94BC5C64F27544CFECD29">
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<mods:date id="076608FE3012928D2A10A8520BAFE979">2022</mods:date>
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<treatment id="03D18798FFDD6862FDC8FA35C1768A25" ID-DOI="http://doi.org/10.5281/zenodo.6312436" ID-GBIF-Taxon="193397475" ID-Zenodo-Dep="6312436" LSID="urn:lsid:zoobank.org:act:491C280C-F0CA-475E-8DC9-377603DCA558" httpUri="http://treatment.plazi.org/id/03D18798FFDD6862FDC8FA35C1768A25" lastPageId="12" lastPageNumber="84" pageId="5" pageNumber="77">
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<taxonomicName id="4C784D0DFFDD686BFDC8FA35C1508B99" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[637,853,1446,1473]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="5" pageNumber="77" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
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<emphasis id="B90CEA9CFFDD686BFDC8FA35C1508B99" bold="true" box="[637,853,1446,1473]" italics="true" pageId="5" pageNumber="77">Theloderma khoii</emphasis>
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<taxonomicNameLabel id="A23F57E7FFDD686BFCE9FA34C1B28B99" box="[860,951,1447,1473]" pageId="5" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BC7368EFFDD686BFD52FA7EC148885F" blockId="5.[365,1223,1446,1543]" box="[743,845,1517,1543]" pageId="5" pageNumber="77">
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<figureCitation id="13432A0BFFDD686BFD52FA7EC148885F" box="[743,845,1517,1543]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1989,2015]" captionStartId-1="8.[189,232,1950,1976]" captionTargetBox-0="[189,1398,1144,1950]" captionTargetBox-1="[200,1384,1250,1906]" captionTargetId-0="figure-335@7.[189,1398,1144,1950]" captionTargetId-1="figure-296@8.[189,1398,1239,1913]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 2. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsolateral view, in life." captionText-1="Fig. 3. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsal (left) and ventral view (right) during fixation." figureDoi-0="http://doi.org/10.5281/zenodo.6130893" figureDoi-1="http://doi.org/10.5281/zenodo.6130897" httpUri-0="https://zenodo.org/record/6130893/files/figure.png" httpUri-1="https://zenodo.org/record/6130897/files/figure.png" pageId="5" pageNumber="77">Figs 2–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3626505FFDD6868FF08F9BDC0008F32" lastPageId="6" lastPageNumber="78" pageId="5" pageNumber="77" type="diagnosis">
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<paragraph id="8BC7368EFFDD686BFF08F9BDC3338810" blockId="5.[189,310,1582,1608]" box="[189,310,1582,1608]" pageId="5" pageNumber="77">
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<emphasis id="B90CEA9CFFDD686BFF08F9BDC3338810" bold="true" box="[189,310,1582,1608]" pageId="5" pageNumber="77">Diagnosis</emphasis>
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<paragraph id="8BC7368EFFDD6868FF08F9CEC0008F32" blockId="5.[187,1401,1629,2007]" lastBlockId="6.[188,1400,265,362]" lastPageId="6" lastPageNumber="78" pageId="5" pageNumber="77">
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<taxonomicName id="4C784D0DFFDD686BFF08F9CEC394882F" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[189,401,1629,1655]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="5" pageNumber="77" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
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<emphasis id="B90CEA9CFFDD686BFF08F9CEC394882F" box="[189,401,1629,1655]" italics="true" pageId="5" pageNumber="77">Theloderma khoii</emphasis>
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shows the diagnostic characters of the genus
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|
||
<emphasis id="B90CEA9CFFDD686BFB85F9CEC6BB882F" box="[1072,1214,1629,1655]" italics="true" pageId="5" pageNumber="77">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
, for instance a distinct tympanum, round canthus rostralis, bony ridges from canthus rostralis to occiput absent, and skin of head not co-ossified to the skull, and also molecular analyses revealed it to be nested within
|
||
<taxonomicName id="4C784D0DFFDD686BFF08F955C34E88B8" authorityName="Tschudi" authorityYear="1838" box="[189,331,1734,1760]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="5" pageNumber="77" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFDD686BFF08F955C34E88B8" box="[189,331,1734,1760]" italics="true" pageId="5" pageNumber="77">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
. The new species is distinguished from its congeners and other small rhacophorid species by a combination of the following characters: 1) large-sized frog of the genus
|
||
<taxonomicName id="4C784D0DFFDD686BFB84F97AC6BA895B" authorityName="Tschudi" authorityYear="1838" box="[1073,1215,1769,1795]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="5" pageNumber="77" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFDD686BFB84F97AC6BA895B" box="[1073,1215,1769,1795]" italics="true" pageId="5" pageNumber="77">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="4C809B6BFFDD686BFABBF97AC772895C" box="[1294,1399,1769,1796]" metricMagnitude="-2" metricUnit="m" metricValue="5.22" pageId="5" pageNumber="77" unit="mm" value="52.2">52.2 mm</quantity>
|
||
in a single male,
|
||
<quantity id="4C809B6BFFDD686BFE34F89EC3ED897F" box="[385,488,1805,1832]" metricMagnitude="-2" metricUnit="m" metricValue="5.9399999999999995" pageId="5" pageNumber="77" unit="mm" value="59.4">59.4 mm</quantity>
|
||
in a single female); 2) head length and width equal; 3) vomerine teeth present; 4) snout pointed and truncated (SNL/SVL 16.3% in male; 16.0% in female); 5) eye large, eye diameter about a half of snout length (ED/SNL 54.60% in male; 58.9% in female), spinules on upper eyelid; 6) tibiotarsal articulation reaching to the posterior border of the eye or the tip of the snout; 7) dorsal skin very rough, with large irregular gland ridges and warts, ventral surface granular; 8) fingers rudimentary webbed, toes almost four-fifths webbed, tips of all digits dilated but all considerably smaller than tympanum; 9) dorsal surface mossy green or olive mottled with dark magenta; 10) lateral and ventral surfaces black with irregular lemon yellow pattern and some white granules; 11) male with large nuptial pads and an inner vocal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFDE6868FF08FE02C3648C1C" pageId="6" pageNumber="78" type="etymology">
|
||
<paragraph id="8BC7368EFFDE6868FF08FE02C3418FF3" blockId="6.[189,324,401,427]" box="[189,324,401,427]" pageId="6" pageNumber="78">
|
||
<heading id="D08F81E2FFDE6868FF08FE02C3418FF3" bold="true" box="[189,324,401,427]" fontSize="11" level="3" pageId="6" pageNumber="78" reason="3">
|
||
<emphasis id="B90CEA9CFFDE6868FF08FE02C3418FF3" bold="true" box="[189,324,401,427]" pageId="6" pageNumber="78">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FE2CC3648C1C" blockId="6.[189,1399,447,580]" pageId="6" pageNumber="78">
|
||
The specific epithet is in honor of Professor Dr Le Vu Khoi of the VNU
|
||
<collectingRegion id="49BCF86CFFDE6868FBAAFE2CC6628F81" box="[1055,1127,447,473]" country="Vietnam" name="Ha Noi" pageId="6" pageNumber="78">Hanoi</collectingRegion>
|
||
University of Science,
|
||
<collectingRegion id="49BCF86CFFDE6868FF08FE70C3138FA4" box="[189,278,482,509]" country="Vietnam" name="Ha Noi" pageId="6" pageNumber="78">Ha Noi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36F761EFFDE6868FE94FE71C3828FA4" box="[289,391,482,508]" name="Vietnam" pageId="6" pageNumber="78">Vietnam</collectingCountry>
|
||
, in recognition of his great support of teaching, mentoring, research and conservation work in
|
||
<collectingCountry id="F36F761EFFDE6868FE97FD95C38C8C78" box="[290,393,518,544]" name="Vietnam" pageId="6" pageNumber="78">Vietnam</collectingCountry>
|
||
. As common names we suggest Khoi’s Mossy Frog (English) and Ếch cây sần khôi (Vietnamese).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFDE6868FF08FDF9C63E8D07" pageId="6" pageNumber="78" type="materials_examined">
|
||
<paragraph id="8BC7368EFFDE6868FF08FDF9C3698CDC" blockId="6.[189,364,618,644]" box="[189,364,618,644]" pageId="6" pageNumber="78">
|
||
<heading id="D08F81E2FFDE6868FF08FDF9C3698CDC" bold="true" box="[189,364,618,644]" fontSize="11" level="3" pageId="6" pageNumber="78" reason="3">
|
||
<emphasis id="B90CEA9CFFDE6868FF08FDF9C3698CDC" bold="true" box="[189,364,618,644]" pageId="6" pageNumber="78">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF5FFD0AC68A8CA2" blockId="6.[189,1398,665,762]" pageId="6" pageNumber="78">
|
||
<materialsCitation id="3B103CD3FFDE6868FF5FFD0AC68A8CA2" ID-GBIF-Occurrence="3467747302" collectingDate="2020-06-08" collectionCode="VNMN" collectorName="H. Q. Nguyen" country="Vietnam" county="northern Vietnam" elevation="1641" location="Quan Ba District" municipality="Quan Ba District" pageId="6" pageNumber="78" specimenCode="VNMN 012757" specimenCount="1" specimenCount-male="1" stateProvince="Ha Giang Province" typeStatus="holotype">
|
||
<emphasis id="B90CEA9CFFDE6868FF5FFD0AC3598CEB" bold="true" box="[234,348,665,691]" pageId="6" pageNumber="78">
|
||
<typeStatus id="54C3882CFFDE6868FF5FFD0AC3598CEB" box="[234,348,665,691]" pageId="6" pageNumber="78" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F36F761EFFDE6868FF08FD2FC3418C8E" box="[189,324,700,726]" name="Vietnam" pageId="6" pageNumber="78">VIETNAM</collectingCountry>
|
||
• adult
|
||
<specimenCount id="9D7EFD07FFDE6868FE14FD2EC3BF8C8F" box="[417,442,701,727]" pageId="6" pageNumber="78" type="male">♂</specimenCount>
|
||
;
|
||
<collectingCounty id="62A64E02FFDE6868FE72FD2FC09D8C8E" box="[455,664,700,726]" pageId="6" pageNumber="78">northern Vietnam</collectingCounty>
|
||
,
|
||
<collectingRegion id="49BCF86CFFDE6868FD10FD2EC18F8C8E" box="[677,906,700,727]" country="Vietnam" name="Ha Giang" pageId="6" pageNumber="78">Ha Giang Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BA3ACF4FFDE6868FC23FD2FC6648C8E" box="[918,1121,700,727]" pageId="6" pageNumber="78">Quan Ba District</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="9A752FACFFDE6868FBDBFD2FC3A88CA1" pageId="6" pageNumber="78">in the karst forest near Tung Vai Commune</locationDeviation>
|
||
;
|
||
<quantity id="4C809B6BFFDE6868FE0CFD4CC0178CA2" box="[441,530,735,762]" metricMagnitude="3" metricUnit="m" metricValue="1.641" pageId="6" pageNumber="78" unit="m" value="1641.0">
|
||
<elevation id="0055D1BDFFDE6868FE0CFD4CC0178CA2" box="[441,530,735,762]" metricMagnitude="3" metricUnit="m" metricValue="1.641" pageId="6" pageNumber="78" unit="m" value="1641.0">1641 m</elevation>
|
||
</quantity>
|
||
a.s.l;
|
||
<date id="FFC6104EFFDE6868FDEDFD4CC0E68CA1" box="[600,739,735,762]" pageId="6" pageNumber="78" value="2020-06-08">
|
||
<collectingDate id="EF82E9A6FFDE6868FDEDFD4CC0E68CA1" box="[600,739,735,762]" pageId="6" pageNumber="78" value="2020-06-08">8 Jun. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="268D5358FFDE6868FD5AFD4CC1888CA2" box="[751,909,735,762]" pageId="6" pageNumber="78">H.Q. Nguyen</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBDE9EF5FFDE6868FC7BFD73C68E8CA2" box="[974,1163,735,762]" pageId="6" pageNumber="78">VNMN 012757</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF5FFCB2C63E8D07" blockId="6.[189,1083,801,863]" pageId="6" pageNumber="78">
|
||
<materialsCitation id="3B103CD3FFDE6868FF5FFCB2C6328D07" ID-GBIF-Occurrence="3467747301" collectingDate="2020-06-08" collectionCode="VNMN" collectorName="H. Q. Nguyen" country="Vietnam" county="northern Vietnam" elevation="1641" location="VIETNAM" municipality="Quan Ba District" pageId="6" pageNumber="78" specimenCode="VNMN 012758" specimenCount="1" specimenCount-female="1" stateProvince="Ha Giang" typeStatus="paratype">
|
||
<emphasis id="B90CEA9CFFDE6868FF5FFCB2C3598D63" bold="true" box="[234,348,801,827]" pageId="6" pageNumber="78">
|
||
<typeStatus id="54C3882CFFDE6868FF5FFCB2C3598D63" box="[234,348,801,827]" pageId="6" pageNumber="78" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F36F761EFFDE6868FF08FCD7C3418D06" box="[189,324,836,862]" name="Vietnam" pageId="6" pageNumber="78">VIETNAM</collectingCountry>
|
||
• adult
|
||
<specimenCount id="9D7EFD07FFDE6868FE2AFCD7C3B38D06" box="[415,438,836,862]" pageId="6" pageNumber="78" type="female">♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="DBDE9EF5FFDE6868FCCFFCD7C6328D07" box="[890,1079,836,863]" pageId="6" pageNumber="78">VNMN 012758</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFDE6866FF08FC16C34D8DE9" lastPageId="8" lastPageNumber="80" pageId="6" pageNumber="78" type="description">
|
||
<paragraph id="8BC7368EFFDE6868FF08FC16C34B8DC7" blockId="6.[189,334,901,927]" box="[189,334,901,927]" pageId="6" pageNumber="78">
|
||
<heading id="D08F81E2FFDE6868FF08FC16C34B8DC7" bold="true" box="[189,334,901,927]" fontSize="11" level="3" pageId="6" pageNumber="78" reason="3">
|
||
<emphasis id="B90CEA9CFFDE6868FF08FC16C34B8DC7" bold="true" box="[189,334,901,927]" pageId="6" pageNumber="78">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF5FFC27C3598D96" blockId="6.[189,819,948,1010]" box="[234,348,948,974]" pageId="6" pageNumber="78">
|
||
<emphasis id="B90CEA9CFFDE6868FF5FFC27C3598D96" bold="true" box="[234,348,948,974]" pageId="6" pageNumber="78">
|
||
<typeStatus id="54C3882CFFDE6868FF5FFC27C3598D96" box="[234,348,948,974]" pageId="6" pageNumber="78" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FC44C1368DAA" blockId="6.[189,819,948,1010]" box="[189,819,983,1010]" pageId="6" pageNumber="78">
|
||
SIZE. Large, body distinctly flattened (SVL
|
||
<quantity id="4C809B6BFFDE6868FD0CFC44C1228DAA" box="[697,807,983,1010]" metricMagnitude="-2" metricUnit="m" metricValue="5.22" pageId="6" pageNumber="78" unit="mm" value="52.2">52.2 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FB8AC65C8A0E" blockId="6.[189,1398,1048,1111]" pageId="6" pageNumber="78">
|
||
HEAD. Very strongly depressed, length and width equal (HL
|
||
<quantity id="4C809B6BFFDE6868FCCFFB8AC1E08A6B" box="[890,997,1049,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="78" unit="mm" value="19.7">19.7 mm</quantity>
|
||
, HW
|
||
<quantity id="4C809B6BFFDE6868FB92FB8AC6968A6B" box="[1063,1171,1049,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="78" unit="mm" value="19.7">19.7 mm</quantity>
|
||
), snout pointed and truncated, snout length double the diameter of eye (SNL
|
||
<quantity id="4C809B6BFFDE6868FCE2FBAFC1B18A0E" box="[855,948,1084,1111]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="78" unit="mm" value="8.5">8.5 mm</quantity>
|
||
, ED
|
||
<quantity id="4C809B6BFFDE6868FC5BFBAFC6488A0E" box="[1006,1101,1084,1110]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="78" unit="mm" value="4.7">4.7 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FBEEC3B88B59" blockId="6.[189,1399,1149,1281]" pageId="6" pageNumber="78">
|
||
CANTHUS ROSTRALIS. Oval, loreal region very oblique, slightly concave; interorbital distance wider than internarial distance and upper eyelid width (IOD 6.0 mm, IN
|
||
<quantity id="4C809B6BFFDE6868FC51FB32C6418AE3" box="[996,1092,1185,1211]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="78" unit="mm" value="3.7">3.7 mm</quantity>
|
||
, UEW
|
||
<quantity id="4C809B6BFFDE6868FB2BFB33C7058AE3" box="[1182,1280,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="78" unit="mm" value="4.5">4.5 mm</quantity>
|
||
); distance between anterior corners of eyes (DAE
|
||
<quantity id="4C809B6BFFDE6868FD2EFB57C10D8A86" box="[667,776,1220,1246]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="6" pageNumber="78" unit="mm" value="10.1">10.1 mm</quantity>
|
||
) about 59.8% distance between posterior corners of eyes (DPE
|
||
<quantity id="4C809B6BFFDE6868FEF6FB74C3B48B59" box="[323,433,1255,1281]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="6" pageNumber="78" unit="mm" value="16.3">16.3 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FABAC3F08B3E" blockId="6.[189,1398,1320,1382]" pageId="6" pageNumber="78">
|
||
NOSTRIL. Oval, inside rather enlarged tubercle on tip of snout, nostrils nearer to tip of snout than to eyes (NS
|
||
<quantity id="4C809B6BFFDE6868FF46FADFC3558B3E" box="[243,336,1356,1382]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="78" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, EN
|
||
<quantity id="4C809B6BFFDE6868FE3FFAD8C3EC8B3E" box="[394,489,1355,1382]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="78" unit="mm" value="6.1">6.1 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FA1EC3748BFF" blockId="6.[189,369,1421,1447]" box="[189,369,1421,1447]" pageId="6" pageNumber="78">PUPIL. Circular.</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08FA5CC76D8854" blockId="6.[189,1398,1486,1548]" pageId="6" pageNumber="78">
|
||
TYMPANUM. Distinct, diameter less than eye diameter and slightly larger than width of third finger disc (TYD
|
||
<quantity id="4C809B6BFFDE6868FEBFFA61C36D8854" box="[266,360,1522,1548]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="78" unit="mm" value="3.7">3.7 mm</quantity>
|
||
), tympanum separated from eye by distance (TYE) 65.0% of tympanum length (TYD).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08F9A0C68B8815" blockId="6.[189,1166,1587,1613]" box="[189,1166,1587,1613]" pageId="6" pageNumber="78">PINEAL OCELLUS. Absent; conical tubercles on eyelid, smaller than those in the back.</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08F9E7C04888E9" blockId="6.[186,1398,1652,1713]" pageId="6" pageNumber="78">SUPRATYMPANIC FOLD. Distinct, interrupted, extending from behind eye to beyond level of axilla, composed of large irregular glandular ridges.</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08F949C338894E" blockId="6.[189,1398,1753,1814]" pageId="6" pageNumber="78">VOMERINE TEETH. Arranged in two small oblique groups between choanae and widely separated from each other.</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08F8ADC3648923" blockId="6.[189,1398,1854,1915]" pageId="6" pageNumber="78">TONGUE. Large, accounting for almost half of mouth, heart-shaped, dorsal surface smooth, notched and free posterior.</paragraph>
|
||
<paragraph id="8BC7368EFFDE6868FF08F830C77D8987" blockId="6.[189,1400,1954,2016]" pageId="6" pageNumber="78">
|
||
FORELIMBS. Moderately strong and long (FLL
|
||
<quantity id="4C809B6BFFDE6868FD6BF831C14F89E5" box="[734,842,1954,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="6" pageNumber="78" unit="mm" value="13.2">13.2 mm</quantity>
|
||
, HAL
|
||
<quantity id="4C809B6BFFDE6868FC2EF831C60D89E5" box="[923,1032,1954,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="6" pageNumber="78" unit="mm" value="15.1">15.1 mm</quantity>
|
||
, 54.3% of SVL), much shorter than hindlimbs (FLL 87.5% of HAL); relative length of fingers: I <II <IV <III; tips of fingers dilated into
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDF6869FF08FE9AC7558F8B" blockId="7.[189,1399,265,468]" pageId="7" pageNumber="79">
|
||
large, rounded and flattened discs, with grooves separating dorsum of discs from venter; disc of finger III approximately two times width of finger III (fd3/fw3 219.2%) but smaller than tympanum diameter (fd3/TYD 83.1%); subarticular tubercles large and distinct, with indistinct supernumerary tubercles; formula of subarticular tubercles: 1, 1, 2, 2; well-developed nuptial pad on lateral side of thumb; outer side of fourth finger fringed, outer edge of fourth finger distinctly serrated; webbing between fingers slightly developed, about 50% between base of finger up to proximal subarticular tubercles (
|
||
<figureCitation id="13432A0BFFDF6869FB4CFE2AC7458F8B" box="[1273,1344,441,467]" captionStart="Fig" captionStartId="8.[189,232,1950,1976]" captionTargetBox="[200,1384,1250,1906]" captionTargetId="figure-296@8.[189,1398,1239,1913]" captionTargetPageId="8" captionText="Fig. 3. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsal (left) and ventral view (right) during fixation." figureDoi="http://doi.org/10.5281/zenodo.6130897" httpUri="https://zenodo.org/record/6130897/files/figure.png" pageId="7" pageNumber="79">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDF6869FF08FE68C0AA8D76" blockId="7.[189,1400,506,814]" pageId="7" pageNumber="79">
|
||
HINDLIMBS. Strong and long, tibia length about five times greater than tibia width (TbL
|
||
<quantity id="4C809B6BFFDF6869FAB7FE68C7778C4D" box="[1282,1394,507,533]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="7" pageNumber="79" unit="mm" value="27.3">27.3 mm</quantity>
|
||
, TbW
|
||
<quantity id="4C809B6BFFDF6869FEB6FD8EC3748C60" box="[259,369,541,568]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" pageId="7" pageNumber="79" unit="mm" value="5.65">5.65 mm</quantity>
|
||
, TbL/TbW 4.6), tibia longer than thigh length and foot length (TbL/FeL 107%; TbL/ FoL 108%), tibio-tarsal articulation reaching posterior edge of snout when hindlimbs pressed forward as opposed to eye; relative length of toes I <II <V <III <IV; toes with rounded discs, tips of toes dilated into flattened discs with grooves that separate dorsum from venter, smaller than discs on fingers; webbing formula I(1/3) – (1)II(0) – (1/2)III(0) – (1)IV(1) – (1/3)V (
|
||
<figureCitation id="13432A0BFFDF6869FC6FFD39C6248C9D" box="[986,1057,682,709]" captionStart="Fig" captionStartId="8.[189,232,1950,1976]" captionTargetBox="[200,1384,1250,1906]" captionTargetId="figure-296@8.[189,1398,1239,1913]" captionTargetPageId="8" captionText="Fig. 3. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsal (left) and ventral view (right) during fixation." figureDoi="http://doi.org/10.5281/zenodo.6130897" httpUri="https://zenodo.org/record/6130897/files/figure.png" pageId="7" pageNumber="79">Fig. 3</figureCitation>
|
||
); large subarticular tubercles roundish: 1, 1, 2, 3, 2; large inner metatarsal tubercle oval and raised (IMT
|
||
<quantity id="4C809B6BFFDF6869FBF1FD5DC6A18CB0" box="[1092,1188,718,744]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="79" unit="mm" value="2.7">2.7 mm</quantity>
|
||
), outer metatarsal tubercle absent; with small warts on inner aspect of metatarsal area; outer side of fifth toe fringed, outer edge of fringe with five distinct serrations.
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFDF6866FF08FCC5C10C8F1E" blockId="7.[188,1398,853,1021]" lastBlockId="8.[189,1399,265,327]" lastPageId="8" lastPageNumber="80" pageId="7" pageNumber="79">SKIN TEXTURE IN LIFE. Dorsal surface of head, dorsum, arms and legs above very rough, with large irregular glandular ridges ordered symmetrically in middle vertebral region; a large, distinct horn gland behind head in X-shape; on top of each supraorbital 3–4 enlarged glands, distinctly conical; lateral parts of body become granular; large conical tubercles on back of thighs near vent; throat and chest with some small blurred pattern; belly and ventral surface and underside of thigh with thickened flat granules or warts on groin; axillary region and underside of tibia smooth; surroundings of tympanum with small tubercles, with 4–5 enlarged tubercles posteriorly.</paragraph>
|
||
<caption id="DF076606FFDF6869FF08F856C6E98987" ID-DOI="http://doi.org/10.5281/zenodo.6130893" ID-Zenodo-Dep="6130893" box="[189,1260,1989,2016]" httpUri="https://zenodo.org/record/6130893/files/figure.png" pageId="7" pageNumber="79" startId="7.[189,232,1989,2015]" targetBox="[189,1398,1144,1950]" targetPageId="7">
|
||
<paragraph id="8BC7368EFFDF6869FF08F856C6E98987" blockId="7.[189,1260,1989,2016]" box="[189,1260,1989,2016]" pageId="7" pageNumber="79">
|
||
<emphasis id="B90CEA9CFFDF6869FF08F856C30E8987" bold="true" box="[189,267,1989,2015]" pageId="7" pageNumber="79">Fig. 2.</emphasis>
|
||
<taxonomicName id="4C784D0DFFDF6869FEA6F856C3E68987" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[275,483,1989,2015]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="7" pageNumber="79" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFDF6869FEA6F856C3E68987" box="[275,483,1989,2015]" italics="true" pageId="7" pageNumber="79">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFDF6869FE5FF855C04289B8" box="[490,583,1990,2016]" pageId="7" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (VNMN 012757). Dorsolateral view, in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC7368EFFD06866FF08FEFDC16D8C03" blockId="8.[188,1400,366,603]" pageId="8" pageNumber="80">
|
||
COLORATION IN LIFE. Dorsal surface mossy-green or light-olive mixed with mossy-green, resembling moss; tips of skin ridges and tubercles reddish-brown; flank with clearly shaped dark brown pattern, ventrally joining a yellowish-green band; dorsal surfaces of forelimbs and hindlimbs with army-green bands, and some irregular patches on webbing; both finger and toe discs green, in center deep olive; nuptial pad ivory; throat and chest dark-brown with yellowish-green warts; belly has black shape of irregular size with yellowish-green pattern and some cream warts; pupil black, iris yellowish-green with irregular black reticulation; tympanum dark-olive (
|
||
<figureCitation id="13432A0BFFD06866FCA7FDD2C15F8C03" box="[786,858,577,603]" captionStart="Fig" captionStartId="7.[189,232,1989,2015]" captionTargetBox="[189,1398,1144,1950]" captionTargetId="figure-335@7.[189,1398,1144,1950]" captionTargetPageId="7" captionText="Fig. 2. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsolateral view, in life." figureDoi="http://doi.org/10.5281/zenodo.6130893" httpUri="https://zenodo.org/record/6130893/files/figure.png" pageId="8" pageNumber="80">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD06866FF08FD11C3808D71" blockId="8.[189,1399,642,810]" pageId="8" pageNumber="80">
|
||
COLORATION IN PRESERVATIVE. In ethanol, dorsal surfaces of head, dorsum, and upper part of flanks dark indigo with brown markings and blotches; tympanum light brown; forelimb, dorsal surface of thigh, tibia and foot black with lights bands, posterior part of thigh below vent dark-grey with large ivory bands; chin and throat light-grey with small ivory spots; chest and belly dark to black, with cream areas on chest (
|
||
<figureCitation id="13432A0BFFD06866FE9BFC9CC3718D71" box="[302,372,783,809]" captionStart="Fig" captionStartId="8.[189,232,1950,1976]" captionTargetBox="[200,1384,1250,1906]" captionTargetId="figure-296@8.[189,1398,1239,1913]" captionTargetPageId="8" captionText="Fig. 3. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). Dorsal (left) and ventral view (right) during fixation." figureDoi="http://doi.org/10.5281/zenodo.6130897" httpUri="https://zenodo.org/record/6130897/files/figure.png" pageId="8" pageNumber="80">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD06866FF5FFCC3C3F28D33" blockId="8.[189,1399,848,945]" box="[234,503,848,875]" pageId="8" pageNumber="80">
|
||
<emphasis id="B90CEA9CFFD06866FF5FFCC3C3F28D33" bold="true" box="[234,503,848,875]" pageId="8" pageNumber="80">
|
||
Variation of
|
||
<typeStatus id="54C3882CFFD06866FE32FCC2C3F28D33" box="[391,503,849,875]" pageId="8" pageNumber="80" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD06866FF08FCE7C34D8DE9" blockId="8.[189,1399,848,945]" pageId="8" pageNumber="80">
|
||
Male smaller than female, with single vocal sac and nuptial pads. For measurements of the
|
||
<typeStatus id="54C3882CFFD06866FB4CFCE7C7298DD6" box="[1273,1324,884,910]" pageId="8" pageNumber="80">type</typeStatus>
|
||
series see
|
||
<tableCitation id="C6FA0335FFD06866FF5FFC04C3468DE9" box="[234,323,919,945]" captionStart="Table 3" captionStartId="9.[188,257,265,291]" captionTargetPageId="9" captionText="Table 3. Measurements (in mm) and proportions of the type series of Theloderma khoii sp. nov. (for abbreviations see Material and methods)." httpUri="http://table.plazi.org/id/DF076606FFD16867FF09FE9AC0A78F1F" pageId="8" pageNumber="80" tableUuid="DF076606FFD16867FF09FE9AC0A78F1F">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFD06865FF08FC4BC1D48D76" lastPageId="11" lastPageNumber="83" pageId="8" pageNumber="80" type="discussion">
|
||
<paragraph id="8BC7368EFFD06866FF08FC4BC0388DAA" blockId="8.[189,573,984,1010]" box="[189,573,984,1010]" pageId="8" pageNumber="80">
|
||
<heading id="D08F81E2FFD06866FF08FC4BC0388DAA" bold="true" box="[189,573,984,1010]" fontSize="11" level="3" pageId="8" pageNumber="80" reason="3">
|
||
<emphasis id="B90CEA9CFFD06866FF08FC4BC0388DAA" bold="true" box="[189,573,984,1010]" pageId="8" pageNumber="80">Comparison with other species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD06867FF08FB94C34E8B4A" blockId="8.[189,1399,1031,1163]" lastBlockId="9.[189,1399,1201,1298]" lastPageId="9" lastPageNumber="81" pageId="8" pageNumber="80">
|
||
We compared
|
||
<taxonomicName id="4C784D0DFFD06866FEC4FB94C04D8A79" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[369,584,1031,1057]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="8" pageNumber="80" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD06866FEC4FB94C04D8A79" box="[369,584,1031,1057]" italics="true" pageId="8" pageNumber="80">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD06866FDE1FB94C0B08A79" box="[596,693,1031,1057]" pageId="8" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with other members of the genus
|
||
<taxonomicName id="4C784D0DFFD06866FBDBFB94C6F98A79" authorityName="Tschudi" authorityYear="1838" box="[1134,1276,1031,1057]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="8" pageNumber="80" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFD06866FBDBFB94C6F98A79" box="[1134,1276,1031,1057]" italics="true" pageId="8" pageNumber="80">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
based on morphological specimen examination (see Appendix 1) and data obtained from the literature (e.g.,
|
||
<bibRefCitation id="EFE94B7FFFD06866FF08FBDEC3678A3F" author="Tschudi J. J. von" box="[189,354,1101,1127]" pageId="8" pageNumber="80" refId="ref10264" refString="Tschudi J. J. von. 1838. Classification der Batrachier mit Berucksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Petitpierre, Neuchatel. https: // doi. org / 10.5962 / bhl. title. 4883" type="book" year="1838">Tschudi 1838</bibRefCitation>
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;
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||
<bibRefCitation id="EFE94B7FFFD06866FEC7FBDEC0308A3F" author="Boulenger G. A." box="[370,565,1101,1127]" pageId="8" pageNumber="80" pagination="186 - 188" refId="ref7573" refString="Boulenger G. A. 1903. Descriptions of three new batrachians from Tonkin. Annals and Magazine of Natural History, Series 7 12: 186 - 188. https: // doi. org / 10.1080 / 00222930308678835" type="journal article" year="1903">Boulenger 1903</bibRefCitation>
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;
|
||
<bibRefCitation id="EFE94B7FFFD06866FDF1FBDEC0D68A3F" author="Smith M. A." box="[580,723,1101,1127]" pageId="8" pageNumber="80" pagination="225 - 234" refId="ref10052" refString="Smith M. A. 1924. New tree-frogs from Indo-China and the Malay Peninsula. Proceedings of the Zoological Society of London 1924: 225 - 234. https: // doi. org / 10.1111 / j. 1096 - 3642.1924. tb 01499. x" type="journal article" year="1924">Smith 1924</bibRefCitation>
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|
||
<bibRefCitation id="EFE94B7FFFD16867FB99FB47C7088AB7" author="Rowley J. J. L. & Le D. T. T. & Hoang H. D. & Dau V. Q. & Cao T. T." box="[1068,1293,1236,1263]" pageId="9" pageNumber="81" pagination="1 - 20" refId="ref9921" refString="Rowley J. J. L., Le D. T. T., Hoang H. D., Dau V. Q. & Cao T. T. 2011. Two new species of Theloderma (Anura: Rhacophoridae) from Vietnam. Zootaxa 3098 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 3098.1.1" type="journal article" year="2011">
|
||
Rowley
|
||
<emphasis id="B90CEA9CFFD16867FB3BFB46C6CC8AB6" box="[1166,1225,1236,1262]" italics="true" pageId="9" pageNumber="81">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE94B7FFFD16867FAAFFB46C33A8B4A" author="Nguyen T. T. & Le D. T. & Nguyen S. H. L. & Matsui M. & Nguyen T. Q." pageId="9" pageNumber="81" pagination="112 - 120" refId="ref9225" refString="Nguyen T. T., Le D. T., Nguyen S. H. L., Matsui M. & Nguyen T. Q. 2014. First record of Philautus petilus Stuart and Heatwole, 2004 (Amphibia: Anura: Rhacophoridae) from Vietnam and its phylogenetic position. Current Herpetology 33: 112 - 120. https: // doi. org / 10.5358 / hsj. 33.112" type="journal article" year="2014">
|
||
Nguyen
|
||
<emphasis id="B90CEA9CFFD16867FF08FB6AC2FD8B4A" box="[189,248,1272,1298]" italics="true" pageId="9" pageNumber="81">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF076606FFD06866FF08F80DC3768984" ID-DOI="http://doi.org/10.5281/zenodo.6130897" ID-Zenodo-Dep="6130897" httpUri="https://zenodo.org/record/6130897/files/figure.png" pageId="8" pageNumber="80" startId="8.[189,232,1950,1976]" targetBox="[200,1384,1250,1906]" targetPageId="8">
|
||
<paragraph id="8BC7368EFFD06866FF08F80DC3768984" blockId="8.[189,1398,1950,2012]" pageId="8" pageNumber="80">
|
||
<emphasis id="B90CEA9CFFD06866FF08F80DC30989E0" bold="true" box="[189,268,1950,1976]" pageId="8" pageNumber="80">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C784D0DFFD06866FEA1F80CC3E089E1" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[276,485,1951,1977]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="8" pageNumber="80" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD06866FEA1F80CC3E089E1" box="[276,485,1951,1977]" italics="true" pageId="8" pageNumber="80">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD06866FE58F80CC04F89E1" box="[493,586,1951,1977]" pageId="8" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (VNMN 012757). Dorsal (left) and ventral view (right) during fixation.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF076606FFD16867FF09FE9AC0A78F1F" ID-Table-UUID="DF076606FFD16867FF09FE9AC0A78F1F" httpUri="http://table.plazi.org/id/DF076606FFD16867FF09FE9AC0A78F1F" pageId="9" pageNumber="81" startId="9.[188,257,265,291]" targetBox="[199,1347,393,1134]" targetIsTable="true" targetPageId="9">
|
||
<paragraph id="8BC7368EFFD16867FF09FE9AC0A78F1F" blockId="9.[188,1401,265,327]" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FF09FE9AC3258F7B" bold="true" box="[188,288,265,291]" pageId="9" pageNumber="81">Table 3.</emphasis>
|
||
Measurements (in mm) and proportions of the type series of
|
||
<taxonomicName id="4C784D0DFFD16867FBB9FE9AC6DB8F7B" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[1036,1246,265,291]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD16867FBB9FE9AC6DB8F7B" box="[1036,1246,265,291]" italics="true" pageId="9" pageNumber="81">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD16867FB5DFE99C7408F7C" box="[1256,1349,266,292]" pageId="9" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(for abbreviations see Material and methods).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC7368EFFD16867FE0AFE1AC7368A35" pageId="9" pageNumber="81">
|
||
<table id="F978C42EFFD19791FF72FE1AC7468A36" box="[199,1347,393,1134]" gridcols="6" gridrows="18" pageId="9" pageNumber="81">
|
||
<tr id="354834CCFFD19791FF72FE1AC7468FF9" box="[199,1347,393,417]" gridrow="0" pageId="9" pageNumber="81" rowspan-0="1" rowspan-3="1">
|
||
<th id="76995DB0FFD19791FE02FE1AC01C8FF9" box="[439,537,393,417]" gridcol="1" gridrow="0" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FE0AFE1AC0178FF9" bold="true" box="[447,530,393,417]" pageId="9" pageNumber="81">VNMN</emphasis>
|
||
</th>
|
||
<th id="76995DB0FFD19791FD35FE1AC0E38FF9" box="[640,742,393,417]" gridcol="2" gridrow="0" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FD32FE1AC0E58FF9" bold="true" box="[647,736,393,417]" pageId="9" pageNumber="81">VNMM</emphasis>
|
||
</th>
|
||
<th id="76995DB0FFD19791FBA0FE1AC6728FF9" box="[1045,1143,393,417]" gridcol="4" gridrow="0" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FBA8FE1AC6758FF9" bold="true" box="[1053,1136,393,417]" pageId="9" pageNumber="81">VNMN</emphasis>
|
||
</th>
|
||
<th id="76995DB0FFD19791FB68FE1AC7468FF9" box="[1245,1347,393,417]" gridcol="5" gridrow="0" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FB56FE1AC7398FF9" bold="true" box="[1251,1340,393,417]" pageId="9" pageNumber="81">VNMM</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FE22C7468F91" box="[199,1347,433,457]" gridrow="1" pageId="9" pageNumber="81" rowspan-0="1" rowspan-3="1">
|
||
<td id="76995DB0FFD19791FE02FE22C01C8F91" box="[439,537,433,457]" gridcol="1" gridrow="1" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FE75FE22C0158F91" bold="true" box="[448,528,433,457]" pageId="9" pageNumber="81">012757</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FD35FE22C0E38F91" box="[640,742,433,457]" gridcol="2" gridrow="1" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FD3EFE22C0DE8F91" bold="true" box="[651,731,433,457]" pageId="9" pageNumber="81">012758</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FBA0FE22C6728F91" box="[1045,1143,433,457]" gridcol="4" gridrow="1" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FBABFE22C66B8F91" bold="true" box="[1054,1134,433,457]" pageId="9" pageNumber="81">012757</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FB68FE22C7468F91" box="[1245,1347,433,457]" gridcol="5" gridrow="1" pageId="9" pageNumber="81">
|
||
<emphasis id="B90CEA9CFFD16867FB5DFE22C73D8F91" bold="true" box="[1256,1336,433,457]" pageId="9" pageNumber="81">012758</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FE4AC7468C40" box="[199,1347,473,536]" gridrow="2" pageId="9" pageNumber="81" rowspan-0="1" rowspan-3="1">
|
||
<td id="76995DB0FFD19791FE02FE4AC01C8C40" box="[439,537,473,536]" gridcol="1" gridrow="2" pageId="9" pageNumber="81">
|
||
♂
|
||
<emphasis id="B90CEA9CFFD16867FE02FD93C01C8C40" bold="true" box="[439,537,512,536]" pageId="9" pageNumber="81">holotype</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FD35FE4AC0E38C40" box="[640,742,473,536]" gridcol="2" gridrow="2" pageId="9" pageNumber="81">
|
||
♀
|
||
<emphasis id="B90CEA9CFFD16867FD35FD93C0E38C40" bold="true" box="[640,742,512,536]" pageId="9" pageNumber="81">paratype</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FBA0FE4AC6728C40" box="[1045,1143,473,536]" gridcol="4" gridrow="2" pageId="9" pageNumber="81">
|
||
♂
|
||
<emphasis id="B90CEA9CFFD16867FBA0FD93C6728C40" bold="true" box="[1045,1143,512,536]" pageId="9" pageNumber="81">holotype</emphasis>
|
||
</td>
|
||
<td id="76995DB0FFD19791FB68FE4AC7468C40" box="[1245,1347,473,536]" gridcol="5" gridrow="2" pageId="9" pageNumber="81">
|
||
♀
|
||
<emphasis id="B90CEA9CFFD16867FB68FD93C7468C40" bold="true" box="[1245,1347,512,536]" pageId="9" pageNumber="81">paratype</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FDBBC7468C18" box="[199,1347,552,576]" gridrow="3" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FDBBC3018C18" box="[199,260,552,576]" gridcol="0" gridrow="3" pageId="9" pageNumber="81">SVL</th>
|
||
<td id="76995DB0FFD19791FE02FDBBC01C8C18" box="[439,537,552,576]" gridcol="1" gridrow="3" pageId="9" pageNumber="81">52.2</td>
|
||
<td id="76995DB0FFD19791FD35FDBBC0E38C18" box="[640,742,552,576]" gridcol="2" gridrow="3" pageId="9" pageNumber="81">59.4</td>
|
||
<td id="76995DB0FFD19791FCF0FDBBC1B28C18" box="[837,951,552,576]" gridcol="3" gridrow="3" pageId="9" pageNumber="81">TYD</td>
|
||
<td id="76995DB0FFD19791FBA0FDBBC6728C18" box="[1045,1143,552,576]" gridcol="4" gridrow="3" pageId="9" pageNumber="81">3.7</td>
|
||
<td id="76995DB0FFD19791FB68FDBBC7468C18" box="[1245,1347,552,576]" gridcol="5" gridrow="3" pageId="9" pageNumber="81">4.0</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FDC3C7468C30" box="[199,1347,592,616]" gridrow="4" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FDC3C3018C30" box="[199,260,592,616]" gridcol="0" gridrow="4" pageId="9" pageNumber="81">HW</th>
|
||
<td id="76995DB0FFD19791FE02FDC3C01C8C30" box="[439,537,592,616]" gridcol="1" gridrow="4" pageId="9" pageNumber="81">19.7</td>
|
||
<td id="76995DB0FFD19791FD35FDC3C0E38C30" box="[640,742,592,616]" gridcol="2" gridrow="4" pageId="9" pageNumber="81">20.7</td>
|
||
<td id="76995DB0FFD19791FCF0FDC3C1B28C30" box="[837,951,592,616]" gridcol="3" gridrow="4" pageId="9" pageNumber="81">TYE</td>
|
||
<td id="76995DB0FFD19791FBA0FDC3C6728C30" box="[1045,1143,592,616]" gridcol="4" gridrow="4" pageId="9" pageNumber="81">2.4</td>
|
||
<td id="76995DB0FFD19791FB68FDC3C7468C30" box="[1245,1347,592,616]" gridcol="5" gridrow="4" pageId="9" pageNumber="81">2.6</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FDE4C7468CC8" box="[199,1347,631,656]" gridrow="5" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FDE4C3018CC8" box="[199,260,631,656]" gridcol="0" gridrow="5" pageId="9" pageNumber="81">HL</th>
|
||
<td id="76995DB0FFD19791FE02FDE4C01C8CC8" box="[439,537,631,656]" gridcol="1" gridrow="5" pageId="9" pageNumber="81">19.7</td>
|
||
<td id="76995DB0FFD19791FD35FDE4C0E38CC8" box="[640,742,631,656]" gridcol="2" gridrow="5" pageId="9" pageNumber="81">21.5</td>
|
||
<td id="76995DB0FFD19791FCF0FDE4C1B28CC8" box="[837,951,631,656]" gridcol="3" gridrow="5" pageId="9" pageNumber="81">FLL</td>
|
||
<td id="76995DB0FFD19791FBA0FDE4C6728CC8" box="[1045,1143,631,656]" gridcol="4" gridrow="5" pageId="9" pageNumber="81">13.2</td>
|
||
<td id="76995DB0FFD19791FB68FDE4C7468CC8" box="[1245,1347,631,656]" gridcol="5" gridrow="5" pageId="9" pageNumber="81">14.7</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FD0CC7468CE0" box="[199,1347,671,696]" gridrow="6" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FD0CC3018CE0" box="[199,260,671,696]" gridcol="0" gridrow="6" pageId="9" pageNumber="81">MN</th>
|
||
<td id="76995DB0FFD19791FE02FD0CC01C8CE0" box="[439,537,671,696]" gridcol="1" gridrow="6" pageId="9" pageNumber="81">18.5</td>
|
||
<td id="76995DB0FFD19791FD35FD0CC0E38CE0" box="[640,742,671,696]" gridcol="2" gridrow="6" pageId="9" pageNumber="81">19.4</td>
|
||
<td id="76995DB0FFD19791FCF0FD0CC1B28CE0" box="[837,951,671,696]" gridcol="3" gridrow="6" pageId="9" pageNumber="81">HAL</td>
|
||
<td id="76995DB0FFD19791FBA0FD0CC6728CE0" box="[1045,1143,671,696]" gridcol="4" gridrow="6" pageId="9" pageNumber="81">15.1</td>
|
||
<td id="76995DB0FFD19791FB68FD0CC7468CE0" box="[1245,1347,671,696]" gridcol="5" gridrow="6" pageId="9" pageNumber="81">19.6</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FD54C7468C87" box="[199,1347,711,735]" gridrow="7" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FD54C3018C87" box="[199,260,711,735]" gridcol="0" gridrow="7" pageId="9" pageNumber="81">MFE</th>
|
||
<td id="76995DB0FFD19791FE02FD54C01C8C87" box="[439,537,711,735]" gridcol="1" gridrow="7" pageId="9" pageNumber="81">13.1</td>
|
||
<td id="76995DB0FFD19791FD35FD54C0E38C87" box="[640,742,711,735]" gridcol="2" gridrow="7" pageId="9" pageNumber="81">14.0</td>
|
||
<td id="76995DB0FFD19791FCF0FD54C1B28C87" box="[837,951,711,735]" gridcol="3" gridrow="7" pageId="9" pageNumber="81">TFL</td>
|
||
<td id="76995DB0FFD19791FBA0FD54C6728C87" box="[1045,1143,711,735]" gridcol="4" gridrow="7" pageId="9" pageNumber="81">3.54</td>
|
||
<td id="76995DB0FFD19791FB68FD54C7468C87" box="[1245,1347,711,735]" gridcol="5" gridrow="7" pageId="9" pageNumber="81">5.54</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FD7CC7468D5F" box="[199,1347,751,775]" gridrow="8" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FD7CC3018D5F" box="[199,260,751,775]" gridcol="0" gridrow="8" pageId="9" pageNumber="81">MBE</th>
|
||
<td id="76995DB0FFD19791FE02FD7CC01C8D5F" box="[439,537,751,775]" gridcol="1" gridrow="8" pageId="9" pageNumber="81">8.8</td>
|
||
<td id="76995DB0FFD19791FD35FD7CC0E38D5F" box="[640,742,751,775]" gridcol="2" gridrow="8" pageId="9" pageNumber="81">8.7</td>
|
||
<td id="76995DB0FFD19791FCF0FD7CC1B28D5F" box="[837,951,751,775]" gridcol="3" gridrow="8" pageId="9" pageNumber="81">fd3</td>
|
||
<td id="76995DB0FFD19791FBA0FD7CC6728D5F" box="[1045,1143,751,775]" gridcol="4" gridrow="8" pageId="9" pageNumber="81">3.2</td>
|
||
<td id="76995DB0FFD19791FB68FD7CC7468D5F" box="[1245,1347,751,775]" gridcol="5" gridrow="8" pageId="9" pageNumber="81">4.2</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FC84C7468D77" box="[199,1347,791,815]" gridrow="9" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FC84C3018D77" box="[199,260,791,815]" gridcol="0" gridrow="9" pageId="9" pageNumber="81">SNL</th>
|
||
<td id="76995DB0FFD19791FE02FC84C01C8D77" box="[439,537,791,815]" gridcol="1" gridrow="9" pageId="9" pageNumber="81">8.5</td>
|
||
<td id="76995DB0FFD19791FD35FC84C0E38D77" box="[640,742,791,815]" gridcol="2" gridrow="9" pageId="9" pageNumber="81">9.5</td>
|
||
<td id="76995DB0FFD19791FCF0FC84C1B28D77" box="[837,951,791,815]" gridcol="3" gridrow="9" pageId="9" pageNumber="81">FeL</td>
|
||
<td id="76995DB0FFD19791FBA0FC84C6728D77" box="[1045,1143,791,815]" gridcol="4" gridrow="9" pageId="9" pageNumber="81">25.6</td>
|
||
<td id="76995DB0FFD19791FB68FC84C7468D77" box="[1245,1347,791,815]" gridcol="5" gridrow="9" pageId="9" pageNumber="81">27.1</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FCADC7468D0F" box="[199,1347,830,855]" gridrow="10" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FCADC3018D0F" box="[199,260,830,855]" gridcol="0" gridrow="10" pageId="9" pageNumber="81">ED</th>
|
||
<td id="76995DB0FFD19791FE02FCADC01C8D0F" box="[439,537,830,855]" gridcol="1" gridrow="10" pageId="9" pageNumber="81">4.6</td>
|
||
<td id="76995DB0FFD19791FD35FCADC0E38D0F" box="[640,742,830,855]" gridcol="2" gridrow="10" pageId="9" pageNumber="81">5.6</td>
|
||
<td id="76995DB0FFD19791FCF0FCADC1B28D0F" box="[837,951,830,855]" gridcol="3" gridrow="10" pageId="9" pageNumber="81">TbL</td>
|
||
<td id="76995DB0FFD19791FBA0FCADC6728D0F" box="[1045,1143,830,855]" gridcol="4" gridrow="10" pageId="9" pageNumber="81">27.3</td>
|
||
<td id="76995DB0FFD19791FB68FCADC7468D0F" box="[1245,1347,830,855]" gridcol="5" gridrow="10" pageId="9" pageNumber="81">29.6</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FCF5C7468D27" box="[199,1347,870,895]" gridrow="11" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FCF5C3018D27" box="[199,260,870,895]" gridcol="0" gridrow="11" pageId="9" pageNumber="81">UEW</th>
|
||
<td id="76995DB0FFD19791FE02FCF5C01C8D27" box="[439,537,870,895]" gridcol="1" gridrow="11" pageId="9" pageNumber="81">4.5</td>
|
||
<td id="76995DB0FFD19791FD35FCF5C0E38D27" box="[640,742,870,895]" gridcol="2" gridrow="11" pageId="9" pageNumber="81">5.7</td>
|
||
<td id="76995DB0FFD19791FCF0FCF5C1B28D27" box="[837,951,870,895]" gridcol="3" gridrow="11" pageId="9" pageNumber="81">TbW</td>
|
||
<td id="76995DB0FFD19791FBA0FCF5C6728D27" box="[1045,1143,870,895]" gridcol="4" gridrow="11" pageId="9" pageNumber="81">6.0</td>
|
||
<td id="76995DB0FFD19791FB68FCF5C7468D27" box="[1245,1347,870,895]" gridcol="5" gridrow="11" pageId="9" pageNumber="81">7.0</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FC1DC7468DFE" box="[199,1347,910,934]" gridrow="12" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FC1DC3018DFE" box="[199,260,910,934]" gridcol="0" gridrow="12" pageId="9" pageNumber="81">IN</th>
|
||
<td id="76995DB0FFD19791FE02FC1DC01C8DFE" box="[439,537,910,934]" gridcol="1" gridrow="12" pageId="9" pageNumber="81">3.7</td>
|
||
<td id="76995DB0FFD19791FD35FC1DC0E38DFE" box="[640,742,910,934]" gridcol="2" gridrow="12" pageId="9" pageNumber="81">3.6</td>
|
||
<td id="76995DB0FFD19791FCF0FC1DC1B28DFE" box="[837,951,910,934]" gridcol="3" gridrow="12" pageId="9" pageNumber="81">FoL</td>
|
||
<td id="76995DB0FFD19791FBA0FC1DC6728DFE" box="[1045,1143,910,934]" gridcol="4" gridrow="12" pageId="9" pageNumber="81">25.3</td>
|
||
<td id="76995DB0FFD19791FB68FC1DC7468DFE" box="[1245,1347,910,934]" gridcol="5" gridrow="12" pageId="9" pageNumber="81">27.2</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FC25C7468D96" box="[199,1347,950,974]" gridrow="13" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FC25C3018D96" box="[199,260,950,974]" gridcol="0" gridrow="13" pageId="9" pageNumber="81">IOD</th>
|
||
<td id="76995DB0FFD19791FE02FC25C01C8D96" box="[439,537,950,974]" gridcol="1" gridrow="13" pageId="9" pageNumber="81">6.1</td>
|
||
<td id="76995DB0FFD19791FD35FC25C0E38D96" box="[640,742,950,974]" gridcol="2" gridrow="13" pageId="9" pageNumber="81">5.5</td>
|
||
<td id="76995DB0FFD19791FCF0FC25C1B28D96" box="[837,951,950,974]" gridcol="3" gridrow="13" pageId="9" pageNumber="81">FTL</td>
|
||
<td id="76995DB0FFD19791FBA0FC25C6728D96" box="[1045,1143,950,974]" gridcol="4" gridrow="13" pageId="9" pageNumber="81">25.1</td>
|
||
<td id="76995DB0FFD19791FB68FC25C7468D96" box="[1245,1347,950,974]" gridcol="5" gridrow="13" pageId="9" pageNumber="81">26.9</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FC4DC7468DAE" box="[199,1347,990,1014]" gridrow="14" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FC4DC3018DAE" box="[199,260,990,1014]" gridcol="0" gridrow="14" pageId="9" pageNumber="81">DAE</th>
|
||
<td id="76995DB0FFD19791FE02FC4DC01C8DAE" box="[439,537,990,1014]" gridcol="1" gridrow="14" pageId="9" pageNumber="81">10.1</td>
|
||
<td id="76995DB0FFD19791FD35FC4DC0E38DAE" box="[640,742,990,1014]" gridcol="2" gridrow="14" pageId="9" pageNumber="81">11.8</td>
|
||
<td id="76995DB0FFD19791FCF0FC4DC1B28DAE" box="[837,951,990,1014]" gridcol="3" gridrow="14" pageId="9" pageNumber="81">SNL/SVL</td>
|
||
<td id="76995DB0FFD19791FBA0FC4DC6728DAE" box="[1045,1143,990,1014]" gridcol="4" gridrow="14" pageId="9" pageNumber="81">16.3%</td>
|
||
<td id="76995DB0FFD19791FB68FC4DC7468DAE" box="[1245,1347,990,1014]" gridcol="5" gridrow="14" pageId="9" pageNumber="81">16.0%</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FB95C7468A46" box="[199,1347,1030,1054]" gridrow="15" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FB95C3018A46" box="[199,260,1030,1054]" gridcol="0" gridrow="15" pageId="9" pageNumber="81">DPE</th>
|
||
<td id="76995DB0FFD19791FE02FB95C01C8A46" box="[439,537,1030,1054]" gridcol="1" gridrow="15" pageId="9" pageNumber="81">16.3</td>
|
||
<td id="76995DB0FFD19791FD35FB95C0E38A46" box="[640,742,1030,1054]" gridcol="2" gridrow="15" pageId="9" pageNumber="81">17.7</td>
|
||
<td id="76995DB0FFD19791FCF0FB95C1B28A46" box="[837,951,1030,1054]" gridcol="3" gridrow="15" pageId="9" pageNumber="81">ED/SNL</td>
|
||
<td id="76995DB0FFD19791FBA0FB95C6728A46" box="[1045,1143,1030,1054]" gridcol="4" gridrow="15" pageId="9" pageNumber="81">54.6%</td>
|
||
<td id="76995DB0FFD19791FB68FB95C7468A46" box="[1245,1347,1030,1054]" gridcol="5" gridrow="15" pageId="9" pageNumber="81">58.9%</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FBBEC7468A1E" box="[199,1347,1069,1094]" gridrow="16" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FBBEC3018A1E" box="[199,260,1069,1094]" gridcol="0" gridrow="16" pageId="9" pageNumber="81">NS</th>
|
||
<td id="76995DB0FFD19791FE02FBBEC01C8A1E" box="[439,537,1069,1094]" gridcol="1" gridrow="16" pageId="9" pageNumber="81">3.0</td>
|
||
<td id="76995DB0FFD19791FD35FBBEC0E38A1E" box="[640,742,1069,1094]" gridcol="2" gridrow="16" pageId="9" pageNumber="81">2.9</td>
|
||
<td id="76995DB0FFD19791FCF0FBBEC1B28A1E" box="[837,951,1069,1094]" gridcol="3" gridrow="16" pageId="9" pageNumber="81">TYE/TYD</td>
|
||
<td id="76995DB0FFD19791FBA0FBBEC6728A1E" box="[1045,1143,1069,1094]" gridcol="4" gridrow="16" pageId="9" pageNumber="81">65.1%</td>
|
||
<td id="76995DB0FFD19791FB68FBBEC7468A1E" box="[1245,1347,1069,1094]" gridcol="5" gridrow="16" pageId="9" pageNumber="81">63.6%</td>
|
||
</tr>
|
||
<tr id="354834CCFFD19791FF72FBC6C7468A36" box="[199,1347,1109,1134]" gridrow="17" pageId="9" pageNumber="81">
|
||
<th id="76995DB0FFD19791FF72FBC6C3018A36" box="[199,260,1109,1134]" gridcol="0" gridrow="17" pageId="9" pageNumber="81">EN</th>
|
||
<td id="76995DB0FFD19791FE02FBC6C01C8A36" box="[439,537,1109,1134]" gridcol="1" gridrow="17" pageId="9" pageNumber="81">6.1</td>
|
||
<td id="76995DB0FFD19791FD35FBC6C0E38A36" box="[640,742,1109,1134]" gridcol="2" gridrow="17" pageId="9" pageNumber="81">6.2</td>
|
||
<td id="76995DB0FFD19791FCF0FBC6C1B28A36" box="[837,951,1109,1134]" gridcol="3" gridrow="17" pageId="9" pageNumber="81">fd3/TYD</td>
|
||
<td id="76995DB0FFD19791FBA0FBC6C6728A36" box="[1045,1143,1109,1134]" gridcol="4" gridrow="17" pageId="9" pageNumber="81">83.1%</td>
|
||
<td id="76995DB0FFD19791FB68FBC6C7468A36" box="[1245,1347,1109,1134]" gridcol="5" gridrow="17" pageId="9" pageNumber="81">76.5%</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD16867FF08FAAAC69D8945" blockId="9.[187,1401,1337,1821]" pageId="9" pageNumber="81">
|
||
Species of
|
||
<taxonomicName id="4C784D0DFFD16867FE8FFAAAC3CD8B0B" authorityName="Tschudi" authorityYear="1838" box="[314,456,1337,1363]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFD16867FE8FFAAAC3CD8B0B" box="[314,456,1337,1363]" italics="true" pageId="9" pageNumber="81">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
inhabiting Indochina, Sumatra, southern
|
||
<collectingCountry id="F36F761EFFD16867FC1CFAAAC1EA8B0B" box="[937,1007,1337,1363]" name="China" pageId="9" pageNumber="81">China</collectingCountry>
|
||
and eastern
|
||
<collectingCountry id="F36F761EFFD16867FBCAFAAAC6BE8B0B" box="[1151,1211,1337,1363]" name="India" pageId="9" pageNumber="81">India</collectingCountry>
|
||
can be assigned to three different size groups (
|
||
<bibRefCitation id="EFE94B7FFFD16867FD9DFACEC0BE8B2F" author="Dever J. A." box="[552,699,1373,1399]" pageId="9" pageNumber="81" pagination="425 - 436" refId="ref7760" refString="Dever J. A. 2017. A new cryptic species of the Theloderma asperum complex (Anura: Rhacophoridae) from Myanmar. Journal of Herpetology 51 (3): 425 - 436. https: // doi. org / 10.1670 / 17 - 026" type="journal article" year="2017">Dever 2017</bibRefCitation>
|
||
): small species (SVL<
|
||
<quantity id="4C809B6BFFD16867FC60FACFC62B8B2F" box="[981,1070,1372,1399]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="81" unit="mm" value="35.0">35 mm</quantity>
|
||
), such as
|
||
<taxonomicName id="4C784D0DFFD16867FB1FFACEC39C8BC2" authority="(Liu & Hu, 1962)" baseAuthorityName="Liu & Hu" baseAuthorityYear="1962" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="albopunctatum">
|
||
<emphasis id="B90CEA9CFFD16867FB1FFACEC77C8B2E" box="[1194,1401,1372,1398]" italics="true" pageId="9" pageNumber="81">T. albopunctatum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FF70FA13C3958BC2" author="Liu C. C. & Hu S. Q." box="[197,400,1407,1434]" pageId="9" pageNumber="81" pagination="73 - 104" refId="ref8927" refString="Liu C. C. & Hu S. Q. 1962. A herpetological report of Kwangsi. Acta Zoologica Sinica 14 (Supplement): 73 - 104." type="journal article" year="1962">Liu & Hu, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FE1DFA13C64B8BC1" authority="Nguyen, Pham, Nguyen, Ngo & Ziegler, 2016" authorityName="Nguyen, Pham, Nguyen, Ngo & Ziegler" authorityYear="2016" box="[424,1102,1407,1434]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="annae">
|
||
<emphasis id="B90CEA9CFFD16867FE1DFA13C0158BC1" box="[424,528,1408,1433]" italics="true" pageId="9" pageNumber="81">T. annae</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FDAFFA13C64B8BC1" author="Nguyen T. Q. & Pham C. T. & Nguyen T. T. & Ngo N. H. & Ziegler T." box="[538,1102,1407,1434]" pageId="9" pageNumber="81" pagination="171 - 186" refId="ref9159" refString="Nguyen T. Q., Pham C. T., Nguyen T. T., Ngo N. H. & Ziegler T. 2016. A new species of Theloderma (Amphibia: Anura: Rhacophoridae) from Vietnam. Zootaxa 4168 (1): 171 - 186. https: // doi. org / 10.11646 / zootaxa. 4168.1.10" type="journal article" year="2016">Nguyen, Pham, Nguyen, Ngo & Ziegler, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FBE8FA13C3008BE6" authority="(Boulenger, 1886)" baseAuthorityName="Boulenger" baseAuthorityYear="1886" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="asperum">
|
||
<emphasis id="B90CEA9CFFD16867FBE8FA13C6E78BC1" box="[1117,1250,1408,1433]" italics="true" pageId="9" pageNumber="81">T. asperum</emphasis>
|
||
(Boulenger, 1886)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FEA4FA37C1C28BE5" authority="Poyarkov, Kropachev, Gogoleva & Orlov, 2018" authorityName="Poyarkov, Kropachev, Gogoleva & Orlov" authorityYear="2018" box="[273,967,1443,1469]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="auratum">
|
||
<emphasis id="B90CEA9CFFD16867FEA4FA37C3958BE5" box="[273,400,1444,1469]" italics="true" pageId="9" pageNumber="81">T. auratum</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FE22FA30C1C28BE5" author="Poyarkov N. A. & Kropachev I. I. & Gogoleva S. I. & Orlov N. L." box="[407,967,1443,1469]" pageId="9" pageNumber="81" pagination="156 - 180" refId="ref9714" refString="Poyarkov N. A., Kropachev I. I., Gogoleva S. I. & Orlov N. L. 2018. A new species of the genus Theloderma Tschudi, 1838 (Amphibia: Anura: Rhacophoridae) from Tay Nguyen Plateau, central Vietnam. Zoological Research 39: 156 - 180. https: // doi. org / 10.24272 / j. issn. 2095 - 8137.2018.018" type="journal article" year="2018">Poyarkov, Kropachev, Gogoleva & Orlov, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FC66FA37C2FE8BB8" authority="Jiang, Fei & Huang, 2009" authorityName="Jiang, Fei & Huang" authorityYear="2009" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="baibungense">
|
||
<emphasis id="B90CEA9CFFD16867FC66FA37C6868BE5" box="[979,1155,1443,1469]" italics="true" pageId="9" pageNumber="81">T. baibungense</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FB3FFA30C2FE8BB8" author="Jiang J. & Fei L. & Huang Y." pageId="9" pageNumber="81" pagination="708 - 711" refId="ref8415" refString="Jiang J., Fei L. & Huang Y. 2009. Aquixalus beibengensis Jiang, Fei & Huang, sp. nov. In: Fei L., Hu S., Ye C. & Huang Y. (eds) Fauna Sinica. Amphibia. Vol. 2. Anura: 708 - 711. Chinese Academy of Science, Science Press, Beijing." type="book chapter" year="2009">Jiang, Fei & Huang, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FEB3FA54C6B38BB8" authority="Sivongxay, Davankham, Phimmachak, Phoumixay & Stuart, 2016" authorityName="Sivongxay, Davankham, Phimmachak, Phoumixay & Stuart" authorityYear="2016" box="[262,1206,1478,1504]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="lacustrinum">
|
||
<emphasis id="B90CEA9CFFD16867FEB3FA54C3A88BB8" box="[262,429,1478,1504]" italics="true" pageId="9" pageNumber="81">T. lacustrinum</emphasis>
|
||
Sivongxay, Davankham, Phimmachak, Phoumixay & Stuart, 2016
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FB0AFA54C301885B" authority="(Smith, 1924)" baseAuthorityName="Smith" baseAuthorityYear="1924" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="laeve">
|
||
<emphasis id="B90CEA9CFFD16867FB0AFA54C71D8BB8" box="[1215,1304,1478,1504]" italics="true" pageId="9" pageNumber="81">T. laeve</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FA92FA55C2FF885B" author="Smith M. A." pageId="9" pageNumber="81" pagination="225 - 234" refId="ref10052" refString="Smith M. A. 1924. New tree-frogs from Indo-China and the Malay Peninsula. Proceedings of the Zoological Society of London 1924: 225 - 234. https: // doi. org / 10.1111 / j. 1096 - 3642.1924. tb 01499. x" type="journal article" year="1924">Smith, 1924</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FEA4FA79C109885B" authority="Bain, Nguyen & Doan, 2009" authorityName="Bain, Nguyen & Doan" authorityYear="2009" box="[273,780,1513,1540]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="lateriticum">
|
||
<emphasis id="B90CEA9CFFD16867FEA4FA79C3AA885B" box="[273,431,1513,1539]" italics="true" pageId="9" pageNumber="81">T. lateriticum</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FE02FA7AC109885B" author="Bain R. H. & Nguyen T. Q. & Doan K. V." box="[439,780,1513,1540]" pageId="9" pageNumber="81" pagination="58 - 68" refId="ref7513" refString="Bain R. H., Nguyen T. Q. & Doan K. V. 2009. A new species of the genus Theloderma Tschudi, 1838 (Anura: Rhacophoridae) from northwestern Vietnam. Zootaxa 2191 (1): 58 - 68. https: // doi. org / 10.5281 / zenodo. 189385" type="journal article" year="2009">Bain, Nguyen & Doan, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FCADFA79C699885B" authority="McLeod & Ahmad, 2007" authorityName="McLeod & Ahmad" authorityYear="2007" box="[792,1180,1513,1539]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="licin">
|
||
<emphasis id="B90CEA9CFFD16867FCADFA79C16C885B" box="[792,873,1513,1539]" italics="true" pageId="9" pageNumber="81">T. licin</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FCC4FA7AC699885B" author="McLeod D. S. & Ahmad N." box="[881,1180,1513,1539]" pageId="9" pageNumber="81" pagination="65 - 72" refId="ref9013" refString="McLeod D. S. & Ahmad N. 2007. A new species of Theloderma (Anura: Rhacophoridae) from southern Thailand and peninsular Malaysia. Russian Journal of Herpetology 14: 65 - 72." type="journal article" year="2007">McLeod & Ahmad, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FB1CFA79C740885B" authorityName="Rowley, Le, Hoang, Dau & Cao" authorityYear="2011" box="[1193,1349,1513,1539]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="nebulosum">
|
||
<emphasis id="B90CEA9CFFD16867FB1CFA79C740885B" box="[1193,1349,1513,1539]" italics="true" pageId="9" pageNumber="81">T. nebulosum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C784D0DFFD16867FF08F99EC11D887F" authority="Rowley, Le, Hoang, Dau & Cao, 2011" authorityName="Rowley, Le, Hoang, Dau & Cao" authorityYear="2011" box="[189,792,1548,1575]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="palliatum">
|
||
<emphasis id="B90CEA9CFFD16867FF08F99EC34F887E" box="[189,330,1548,1574]" italics="true" pageId="9" pageNumber="81">T. palliatum</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FEE6F99FC11D887F" author="Rowley J. J. L. & Le D. T. T. & Hoang H. D. & Dau V. Q. & Cao T. T." box="[339,792,1548,1575]" pageId="9" pageNumber="81" pagination="1 - 20" refId="ref9921" refString="Rowley J. J. L., Le D. T. T., Hoang H. D., Dau V. Q. & Cao T. T. 2011. Two new species of Theloderma (Anura: Rhacophoridae) from Vietnam. Zootaxa 3098 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 3098.1.1" type="journal article" year="2011">Rowley, Le, Hoang, Dau & Cao, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FC91F99EC6DB887F" authority="(Stuart & Heatwole, 2004)" baseAuthorityName="Stuart & Heatwole" baseAuthorityYear="2004" box="[804,1246,1548,1575]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="petilum">
|
||
<emphasis id="B90CEA9CFFD16867FC91F99EC19C887E" box="[804,921,1548,1574]" italics="true" pageId="9" pageNumber="81">T. petilum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FC1CF99EC6D3887F" author="Stuart B. L. & Heatwole H. F." box="[937,1238,1548,1575]" pageId="9" pageNumber="81" pagination="17 - 21" refId="ref10100" refString="Stuart B. L. & Heatwole H. F. 2004. A new Philautus (Amphibia: Rhacophoridae) from northern Laos. Asiatic Herpetological Research 10: 17 - 21." type="journal article" year="2004">Stuart & Heatwole, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FB5EF99EC3518812" authority="Dever, 2017" authorityName="Dever" authorityYear="2017" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="pyaukkya">
|
||
<emphasis id="B90CEA9CFFD16867FB5EF99EC772887E" box="[1259,1399,1548,1574]" italics="true" pageId="9" pageNumber="81">T. pyaukkya</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FF08F9A3C3518812" author="Dever J. A." box="[189,340,1584,1610]" pageId="9" pageNumber="81" pagination="425 - 436" refId="ref7760" refString="Dever J. A. 2017. A new cryptic species of the Theloderma asperum complex (Anura: Rhacophoridae) from Myanmar. Journal of Herpetology 51 (3): 425 - 436. https: // doi. org / 10.1670 / 17 - 026" type="journal article" year="2017">Dever, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FED0F9A3C0FA8812" authority="(Liu & Hu, 1926)" baseAuthorityName="Liu & Hu" baseAuthorityYear="1926" box="[357,767,1583,1610]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="rhododiscus">
|
||
<emphasis id="B90CEA9CFFD16867FED0F9A3C0118812" box="[357,532,1584,1610]" italics="true" pageId="9" pageNumber="81">T. rhododiscus</emphasis>
|
||
(Liu & Hu, 1926)
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B90CEA9CFFD16867FCBAF9A3C1228811" box="[783,807,1584,1609]" italics="true" pageId="9" pageNumber="81">T.</emphasis>
|
||
(
|
||
<taxonomicName id="4C784D0DFFD16867FC8CF9A3C1C28812" box="[825,967,1584,1610]" class="Amphibia" family="Rhacophoridae" genus="Stelladerma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFD16867FC8CF9A3C1C28812" box="[825,967,1584,1610]" italics="true" pageId="9" pageNumber="81">Stelladerma</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFE94B7FFFD16867FC40F9A3C6F28812" author="Poyarkov N. A. & Orlov N. L. & Moiseeva A. V. & Pawangkhanant P. & Ruwangsuwan T. & Vassilieva A. B. & Galoyan E. A. & Nguyen T. T. & Gogoleva S. S." box="[1013,1271,1583,1610]" pageId="9" pageNumber="81" pagination="241 - 280" refId="ref9623" refString="Poyarkov N. A., Orlov N. L., Moiseeva A. V., Pawangkhanant P., Ruwangsuwan T., Vassilieva A. B., Galoyan E. A., Nguyen T. T. & Gogoleva S. S. 2015. Sorting out moss frogs: mtDNA data on taxonomic diversity and phylogenetic relationships of the Indochinese species of the genus Theloderma (Anura, Rhacophoridae). Russian Journal of Herpetology 22 (4): 241 - 280." type="journal article" year="2015">
|
||
Poyarkov
|
||
<emphasis id="B90CEA9CFFD16867FBC4F9A3C6B58812" box="[1137,1200,1584,1610]" italics="true" pageId="9" pageNumber="81">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
)
|
||
<taxonomicName id="4C784D0DFFD16867FABEF9A3C35E8835" authority="Taylor, 1926" authorityName="Taylor" authorityYear="1926" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="stellatum">
|
||
<emphasis id="B90CEA9CFFD16867FABEF9A3C7728812" box="[1291,1399,1584,1610]" italics="true" pageId="9" pageNumber="81">stellatum</emphasis>
|
||
Taylor, 1926
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B90CEA9CFFD16867FED8F9C7C3868835" box="[365,387,1620,1645]" italics="true" pageId="9" pageNumber="81">T.</emphasis>
|
||
(
|
||
<taxonomicName id="4C784D0DFFD16867FE2FF9C0C02D8835" box="[410,552,1619,1645]" class="Amphibia" family="Rhacophoridae" genus="Stelladerma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFD16867FE2FF9C0C02D8835" box="[410,552,1619,1645]" italics="true" pageId="9" pageNumber="81">Stelladerma</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFE94B7FFFD16867FDECF9C0C1648836" author="Poyarkov N. A. & Orlov N. L. & Moiseeva A. V. & Pawangkhanant P. & Ruwangsuwan T. & Vassilieva A. B. & Galoyan E. A. & Nguyen T. T. & Gogoleva S. S." box="[601,865,1619,1646]" pageId="9" pageNumber="81" pagination="241 - 280" refId="ref9623" refString="Poyarkov N. A., Orlov N. L., Moiseeva A. V., Pawangkhanant P., Ruwangsuwan T., Vassilieva A. B., Galoyan E. A., Nguyen T. T. & Gogoleva S. S. 2015. Sorting out moss frogs: mtDNA data on taxonomic diversity and phylogenetic relationships of the Indochinese species of the genus Theloderma (Anura, Rhacophoridae). Russian Journal of Herpetology 22 (4): 241 - 280." type="journal article" year="2015">
|
||
Poyarkov
|
||
<emphasis id="B90CEA9CFFD16867FD62F9C7C11D8835" box="[727,792,1619,1645]" italics="true" pageId="9" pageNumber="81">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
)
|
||
<taxonomicName id="4C784D0DFFD16867FCC2F9C7C73E8836" authority="(Poyarkov et al., 2015)" baseAuthorityName="Poyarkov" baseAuthorityYear="2015" box="[887,1339,1619,1646]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="vietnamense">
|
||
<emphasis id="B90CEA9CFFD16867FCC2F9C7C60D8835" box="[887,1032,1620,1645]" italics="true" pageId="9" pageNumber="81">vietnamense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FBA8F9C0C7348836" author="Poyarkov N. A. & Orlov N. L. & Moiseeva A. V. & Pawangkhanant P. & Ruwangsuwan T. & Vassilieva A. B. & Galoyan E. A. & Nguyen T. T. & Gogoleva S. S." box="[1053,1329,1619,1646]" pageId="9" pageNumber="81" pagination="241 - 280" refId="ref9623" refString="Poyarkov N. A., Orlov N. L., Moiseeva A. V., Pawangkhanant P., Ruwangsuwan T., Vassilieva A. B., Galoyan E. A., Nguyen T. T. & Gogoleva S. S. 2015. Sorting out moss frogs: mtDNA data on taxonomic diversity and phylogenetic relationships of the Indochinese species of the genus Theloderma (Anura, Rhacophoridae). Russian Journal of Herpetology 22 (4): 241 - 280." type="journal article" year="2015">
|
||
Poyarkov
|
||
<emphasis id="B90CEA9CFFD16867FB28F9C7C6D88835" box="[1181,1245,1619,1645]" italics="true" pageId="9" pageNumber="81">et al.</emphasis>
|
||
, 2015
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C784D0DFFD16867FF08F9E4C08B88C9" authority="(Orlov & Ho, 2005)" baseAuthorityName="Orlov & Ho" baseAuthorityYear="2005" box="[189,654,1654,1681]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="truongsonense">
|
||
<emphasis id="B90CEA9CFFD16867FF08F9E4C38C88C8" box="[189,393,1655,1680]" italics="true" pageId="9" pageNumber="81">T. truongsonense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FE29F9E5C08188C9" author="Orlov N. L. & Ho C. T." box="[412,644,1654,1681]" pageId="9" pageNumber="81" pagination="135 - 142" refId="ref9448" refString="Orlov N. L. & Ho C. T. 2005. A new species of Philautus from Vietnam (Anura: Rhacophoridae). Russian Journal of Herpetology 12: 135 - 142." type="journal article" year="2005">Orlov & Ho, 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; medium-sized species with SVL of
|
||
<quantity id="4C809B6BFFD16867FBE7F9E5C6DD88C9" box="[1106,1240,1654,1681]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="9" pageNumber="81" unit="mm" value="42.5" valueMax="45.0" valueMin="40.0">40–45 mm</quantity>
|
||
,
|
||
<taxonomicName id="4C784D0DFFD16867FB52F9E4C39C88EB" authority="(Boulenger, 1903)" baseAuthorityName="Boulenger" baseAuthorityYear="1903" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="horridum">
|
||
<emphasis id="B90CEA9CFFD16867FB52F9E4C77288C8" box="[1255,1399,1654,1680]" italics="true" pageId="9" pageNumber="81">T. horridum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FF70F90AC39588EB" author="Boulenger G. A." box="[197,400,1689,1715]" pageId="9" pageNumber="81" pagination="186 - 188" refId="ref7573" refString="Boulenger G. A. 1903. Descriptions of three new batrachians from Tonkin. Annals and Magazine of Natural History, Series 7 12: 186 - 188. https: // doi. org / 10.1080 / 00222930308678835" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FE1DF909C10E88EB" authority="(Annandale, 1912)" baseAuthorityName="Annandale" baseAuthorityYear="1912" box="[424,779,1689,1715]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="moloch">
|
||
<emphasis id="B90CEA9CFFD16867FE1DF909C01B88EB" box="[424,542,1689,1715]" italics="true" pageId="9" pageNumber="81">T. moloch</emphasis>
|
||
(Annandale, 1912)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FCACF909C67488EB" authority="(Ahl, 1927)" baseAuthorityName="Ahl" baseAuthorityYear="1927" box="[793,1137,1689,1715]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="phrynoderma">
|
||
<emphasis id="B90CEA9CFFD16867FCACF909C1D288EB" box="[793,983,1689,1715]" italics="true" pageId="9" pageNumber="81">T. phrynoderma</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FC5FF90AC66D88EB" author="Ahl E." box="[1002,1128,1689,1715]" pageId="9" pageNumber="81" pagination="35 - 47" refId="ref7391" refString="Ahl E. 1927. Zur Systematik der asiatischen Arten der Froschgattung Rhacophorus. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1927: 35 - 47." type="journal article" year="1927">Ahl, 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FBCAF909C74788EB" authorityName="Orlov, Dutta, Ghate & Kent" authorityYear="2006" box="[1151,1346,1689,1715]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="nagalandense">
|
||
<emphasis id="B90CEA9CFFD16867FBCAF909C74788EB" box="[1151,1346,1689,1715]" italics="true" pageId="9" pageNumber="81">T. nagalandense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C784D0DFFD16867FF08F92EC0DC888E" authority="Orlov, Dutta, Ghate & Kent, 2006" authorityName="Orlov, Dutta, Ghate & Kent" authorityYear="2006" box="[189,729,1724,1751]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="ryabovi">
|
||
<emphasis id="B90CEA9CFFD16867FF08F92EC333888E" box="[189,310,1724,1750]" italics="true" pageId="9" pageNumber="81">T. ryabovi</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FE8BF92FC0DC888E" author="Orlov N. L. & Dutta S. K. & Ghate H. V. & Kent Y." box="[318,729,1724,1751]" pageId="9" pageNumber="81" pagination="165 - 175" refId="ref9484" refString="Orlov N. L., Dutta S. K., Ghate H. V. & Kent Y. 2006. New species of Theloderma from Kon Tum Province (Vietnam) and Nagaland State (India) (Anura: Rhacophoridae). Russian Journal of Herpetology 13: 165 - 175." type="journal article" year="2006">Orlov, Dutta, Ghate & Kent, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C784D0DFFD16867FCA9F92EC68E888F" authority="(Kelaart, 1853)" baseAuthorityName="Kelaart" baseAuthorityYear="1853" box="[796,1163,1724,1751]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="schmardana">
|
||
<emphasis id="B90CEA9CFFD16867FCA9F92EC1CE888E" box="[796,971,1724,1750]" italics="true" pageId="9" pageNumber="81">T. schmardana</emphasis>
|
||
(Kelaart, 1853)
|
||
</taxonomicName>
|
||
; large-sized species with SVL
|
||
<quantity id="4C809B6BFFD16867FE83F94CC3B188A2" box="[310,436,1759,1786]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" metricValueMax="7.5" metricValueMin="4.8" pageId="9" pageNumber="81" unit="mm" value="61.5" valueMax="75.0" valueMin="48.0">48–75 mm</quantity>
|
||
(Orlov 1997), including
|
||
<taxonomicName id="4C784D0DFFD16867FD66F973C60188A2" authority="(Bourret, 1937)" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[723,1028,1760,1786]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B90CEA9CFFD16867FD66F973C14188A2" box="[723,836,1760,1786]" italics="true" pageId="9" pageNumber="81">T. bicolor</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FCE6F973C1FE88A2" author="Bourret R." box="[851,1019,1760,1786]" pageId="9" pageNumber="81" pagination="5 - 56" refId="ref7613" refString="Bourret R. 1937. Notes herpetologiques sur l'Indochine francaise. XIV. Les batraciens de la collection du Laboratoire des Sciences Naturelles de l'Universite. Descriptions de quinze especes ou varietes nouvelles. Annexe au Bulletin General de l'Instruction Publique, Hanoi 4: 5 - 56." type="journal article" year="1937">Bourret, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FBBBF973C77788A2" authority="(Boulenger, 1903)" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[1038,1394,1760,1786]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="corticale">
|
||
<emphasis id="B90CEA9CFFD16867FBBBF973C69788A2" box="[1038,1170,1760,1786]" italics="true" pageId="9" pageNumber="81">T. corticale</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD16867FB14F973C76C88A2" author="Boulenger G. A." box="[1185,1385,1760,1786]" pageId="9" pageNumber="81" pagination="186 - 188" refId="ref7573" refString="Boulenger G. A. 1903. Descriptions of three new batrachians from Tonkin. Annals and Magazine of Natural History, Series 7 12: 186 - 188. https: // doi. org / 10.1080 / 00222930308678835" type="journal article" year="1903">Boulenger, 1903</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FF09F897C3D08945" authority="Taylor, 1962" authorityName="Taylor" authorityYear="1962" box="[188,469,1795,1821]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD16867FF09F897C3308945" box="[188,309,1795,1821]" italics="true" pageId="9" pageNumber="81">T. gordoni</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FE88F890C3D08945" author="Taylor E. H." box="[317,469,1795,1821]" pageId="9" pageNumber="81" pagination="265 - 599" refId="ref10226" refString="Taylor E. H. 1962. The amphibian fauna of Thailand. University of Kansas Science Bulletin 43: 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor, 1962</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C784D0DFFD16867FE54F897C12D8945" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" box="[481,808,1795,1821]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="leporosum">
|
||
<emphasis id="B90CEA9CFFD16867FE54F897C07C8945" box="[481,633,1795,1821]" italics="true" pageId="9" pageNumber="81">T. leporosum</emphasis>
|
||
<bibRefCitation id="EFE94B7FFFD16867FDCAF890C12D8945" author="Tschudi J. J. von" box="[639,808,1795,1821]" pageId="9" pageNumber="81" refId="ref10264" refString="Tschudi J. J. von. 1838. Classification der Batrachier mit Berucksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Petitpierre, Neuchatel. https: // doi. org / 10.5962 / bhl. title. 4883" type="book" year="1838">Tschudi, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C784D0DFFD16867FCD3F890C6338945" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[870,1078,1795,1821]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD16867FCD3F890C6338945" box="[870,1078,1795,1821]" italics="true" pageId="9" pageNumber="81">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD16867FB88F890C69D8945" box="[1085,1176,1795,1821]" pageId="9" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD16865FF08F8D7C7138C42" blockId="9.[188,1399,1860,1992]" lastBlockId="11.[188,1401,265,538]" lastPageId="11" lastPageNumber="83" pageId="9" pageNumber="81">
|
||
Compared to
|
||
<taxonomicName id="4C784D0DFFD16867FEECF8D6C3CC8906" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[345,457,1860,1886]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B90CEA9CFFD16867FEECF8D6C3CC8906" box="[345,457,1860,1886]" italics="true" pageId="9" pageNumber="81">T. bicolor</emphasis>
|
||
</taxonomicName>
|
||
, another large-sized group member in
|
||
<collectingCountry id="F36F761EFFD16867FC3DF8D7C1EB8906" box="[904,1006,1860,1886]" name="Vietnam" pageId="9" pageNumber="81">Vietnam</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C784D0DFFD16867FC4DF8D7C6C38906" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[1016,1222,1860,1886]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD16867FC4DF8D7C6C38906" box="[1016,1222,1860,1886]" italics="true" pageId="9" pageNumber="81">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD16867FB78F8D6C7228907" box="[1229,1319,1861,1887]" pageId="9" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished by its coloration pattern: the lateral part from the axilla to groin is black with an irregular yellowish-green pattern in
|
||
<taxonomicName id="4C784D0DFFD16867FE4EF818C06889FD" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[507,621,1931,1957]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B90CEA9CFFD16867FE4EF818C06889FD" box="[507,621,1931,1957]" italics="true" pageId="9" pageNumber="81">T. bicolor</emphasis>
|
||
</taxonomicName>
|
||
vs flank pattern with clear brown-dark speckles, ventrally joining the yellowish-green band in
|
||
<taxonomicName id="4C784D0DFFD16867FDBCF83DC0DD8990" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[521,728,1966,1992]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="9" pageNumber="81" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD16867FDBCF83DC0DD8990" box="[521,728,1966,1992]" italics="true" pageId="9" pageNumber="81">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD16867FD6AF83DC1388990" box="[735,829,1966,1992]" pageId="9" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; a comparatively lighter belly pattern, consisting of irregular blackish brown speckles with whitish pattern in
|
||
<taxonomicName id="4C784D0DFFD26864FC2BFE99C6168F7B" baseAuthorityName="Bourret" baseAuthorityYear="1937" box="[926,1043,265,291]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B90CEA9CFFD26864FC2BFE99C6168F7B" box="[926,1043,265,291]" italics="true" pageId="10" pageNumber="82">T. bicolor</emphasis>
|
||
</taxonomicName>
|
||
vs belly irregular black with yellowish-green pattern and some cream warts, and the presence of few smaller bluish spots on the throat, in
|
||
<taxonomicName id="4C784D0DFFD26864FE80FEDCC00D8F31" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[309,520,335,361]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD26864FE80FEDCC00D8F31" box="[309,520,335,361]" italics="true" pageId="10" pageNumber="82">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD26864FDBAFEC3C0698F32" box="[527,620,336,362]" pageId="10" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13432A0BFFD26864FDCBFEDCC0CC8F32" box="[638,713,335,362]" captionStart="Fig" captionStartId="10.[188,231,1953,1979]" captionTargetBox="[198,1388,621,1890]" captionTargetId="figure-150@10.[187,1398,583,1913]" captionTargetPageId="10" captionText="Fig. 4. A, C, E. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). B, D, F. Theloderma bicolor (Bourret, 1937) (VNMN 010811). Dorsal (A–B) and ventral views (C–D) as well as lateral head (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6130901" httpUri="https://zenodo.org/record/6130901/files/figure.png" pageId="10" pageNumber="82">Fig. 4</figureCitation>
|
||
); by the comparatively dark-colored posterior part of the thighs, consisting of black blotches and irregular light-green or olive reticulations (vs black with yellowgreenish bands and a few red-brown warts in
|
||
<taxonomicName id="4C784D0DFFD26864FD5AFE05C1C68FE8" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[751,963,406,432]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD26864FD5AFE05C1C68FE8" box="[751,963,406,432]" italics="true" pageId="10" pageNumber="82">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD26864FC78FE05C62B8FE8" box="[973,1070,406,432]" pageId="10" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
;
|
||
<figureCitation id="13432A0BFFD26864FB8BFE05C68C8FE8" box="[1086,1161,406,432]" captionStart="Fig" captionStartId="10.[188,231,1953,1979]" captionTargetBox="[198,1388,621,1890]" captionTargetId="figure-150@10.[187,1398,583,1913]" captionTargetPageId="10" captionText="Fig. 4. A, C, E. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). B, D, F. Theloderma bicolor (Bourret, 1937) (VNMN 010811). Dorsal (A–B) and ventral views (C–D) as well as lateral head (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6130901" httpUri="https://zenodo.org/record/6130901/files/figure.png" pageId="10" pageNumber="82">Fig. 4</figureCitation>
|
||
); and foot webbing well-developed: I(0) – (1)II(0) – (1/2)III(0) – (1)IV(1) – (0)V (vs I(1/3) – (1)II(0) – (1/2)III(0) – (1) IV(1) – (1/3)V in
|
||
<taxonomicName id="4C784D0DFFD26864FE3DFE4FC0528FAE" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[392,599,476,502]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD26864FE3DFE4FC0528FAE" box="[392,599,476,502]" italics="true" pageId="10" pageNumber="82">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD26864FDEBFE4EC0BF8FAF" box="[606,698,477,503]" pageId="10" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
;
|
||
<figureCitation id="13432A0BFFD26864FD70FE4FC10E8FAE" box="[709,779,476,502]" captionStart="Fig" captionStartId="10.[188,231,1953,1979]" captionTargetBox="[198,1388,621,1890]" captionTargetId="figure-150@10.[187,1398,583,1913]" captionTargetPageId="10" captionText="Fig. 4. A, C, E. Theloderma khoii sp. nov., holotype, ♂ (VNMN 012757). B, D, F. Theloderma bicolor (Bourret, 1937) (VNMN 010811). Dorsal (A–B) and ventral views (C–D) as well as lateral head (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6130901" httpUri="https://zenodo.org/record/6130901/files/figure.png" pageId="10" pageNumber="82">Fig. 4</figureCitation>
|
||
).
|
||
<taxonomicName id="4C784D0DFFD26864FCABFE4FC1E88FAE" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[798,1005,476,502]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD26864FCABFE4FC1E88FAE" box="[798,1005,476,502]" italics="true" pageId="10" pageNumber="82">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD26864FC47FE4EC64E8FAF" box="[1010,1099,477,503]" pageId="10" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD26864FBECFE4FC6FD8FAE" author="Bourret R." box="[1113,1272,476,502]" pageId="10" pageNumber="82" pagination="5 - 56" refId="ref7613" refString="Bourret R. 1937. Notes herpetologiques sur l'Indochine francaise. XIV. Les batraciens de la collection du Laboratoire des Sciences Naturelles de l'Universite. Descriptions de quinze especes ou varietes nouvelles. Annexe au Bulletin General de l'Instruction Publique, Hanoi 4: 5 - 56." type="journal article" year="1937">Bourret 1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE94B7FFFD26864FAB6FE4EC2FC8C42" author="Hou M. & Yu G. & Chen H. & Liao C. & Zhang L. & Chen J. & Li P. & Orlov N. L." pageId="10" pageNumber="82" pagination="99 - 127" refId="ref8203" refString="Hou M., Yu G., Chen H., Liao C., Zhang L., Chen J., Li P. & Orlov N. L. 2017. The taxonomic status and distribution range of six Theloderma species (Anura: Rhacophoridae) with a new record in China. Russian Journal of Herpetology 24: 99 - 127. https: // doi. org / 10.30906 / 1026 - 2296 - 2019 - 24 - 2 - 99 - 127" type="journal article" year="2017">
|
||
Hou
|
||
<emphasis id="B90CEA9CFFD26864FA89FE4EC76B8FAE" box="[1340,1390,476,502]" italics="true" pageId="10" pageNumber="82">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
) differs from
|
||
<taxonomicName id="4C784D0DFFD26864FE15FD93C0238C41" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[416,550,511,537]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="corticale">
|
||
<emphasis id="B90CEA9CFFD26864FE15FD93C0238C41" box="[416,550,511,537]" italics="true" pageId="10" pageNumber="82">T. corticale</emphasis>
|
||
</taxonomicName>
|
||
by its shorter body size: SVL
|
||
<quantity id="4C809B6BFFD26864FC27FE6CC1F98C42" box="[914,1020,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.22" pageId="10" pageNumber="82" unit="mm" value="52.2">52.2 mm</quantity>
|
||
in male and
|
||
<quantity id="4C809B6BFFD26864FB22FE6CC7048C42" box="[1175,1281,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="5.9399999999999995" pageId="10" pageNumber="82" unit="mm" value="59.4">59.4 mm</quantity>
|
||
in female in
|
||
<taxonomicName id="4C784D0DFFD36865FF6BFE9AC3B48F7B" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[222,433,265,291]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FF6BFE9AC3B48F7B" box="[222,433,265,291]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FE0FFE99C01D8F7C" box="[442,536,266,292]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs SVL up to
|
||
<quantity id="4C809B6BFFD36865FD7BFE9AC1268F7C" box="[718,803,265,292]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="83" unit="mm" value="67.0">67 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C784D0DFFD36865FCFBFE99C1D08F7B" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[846,981,265,291]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="corticale">
|
||
<emphasis id="B90CEA9CFFD36865FCFBFE99C1D08F7B" box="[846,981,265,291]" italics="true" pageId="11" pageNumber="83">T. corticale</emphasis>
|
||
</taxonomicName>
|
||
; by colour pattern: a much lighter belly pattern, yellow-lemon with irregular small black patterns and creamy warts (vs dark belly pattern, blackish ground color with thin yellowish-green pattern and some cream warts in
|
||
<taxonomicName id="4C784D0DFFD36865FB16FEDCC7728F31" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[1187,1399,335,361]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FB16FEDCC7728F31" box="[1187,1399,335,361]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FF08FEE0C31F8FD5" box="[189,282,371,397]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); by skin texture: the absence of enlarged warts in the scapular area (vs large irregular prominent warts on dorsum, forming large groups at the base of the head and scapular area in
|
||
<taxonomicName id="4C784D0DFFD36865FB12FE05C77D8FE8" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[1191,1400,406,432]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FB12FE05C77D8FE8" box="[1191,1400,406,432]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FF08FE29C31B8F8C" box="[189,286,442,468]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); by the head being slightly wider than long (vs head length and width equal in
|
||
<taxonomicName id="4C784D0DFFD36865FB5EFE2AC2FD8FAE" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FB5EFE2AC2FD8FAE" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FF4AFE4EC3588FAF" box="[255,349,477,503]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); and foot webbing well developed: webbing formula: I(0) – (1 + 12)II(0) – (1)III(0) – (1) IV(1) – (0)V (vs I(1/3) – (1)II(0) – (1/2)III(0) – (1)IV(1) – (1/3)V in
|
||
<taxonomicName id="4C784D0DFFD36865FC51FE6CC6B18C41" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[996,1204,511,537]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FC51FE6CC6B18C41" box="[996,1204,511,537]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FB0EFD93C7138C42" box="[1211,1302,512,538]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF076606FFD26864FF09F832C7728987" ID-DOI="http://doi.org/10.5281/zenodo.6130901" ID-Zenodo-Dep="6130901" httpUri="https://zenodo.org/record/6130901/files/figure.png" pageId="10" pageNumber="82" startId="10.[188,231,1953,1979]" targetBox="[198,1388,621,1890]" targetPageId="10">
|
||
<paragraph id="8BC7368EFFD26864FF09F832C7728987" blockId="10.[187,1399,1953,2015]" pageId="10" pageNumber="82">
|
||
<emphasis id="B90CEA9CFFD26864FF09F832C36E89E4" bold="true" box="[188,363,1953,1980]" pageId="10" pageNumber="82">Fig. 4. A, C, E</emphasis>
|
||
.
|
||
<taxonomicName id="4C784D0DFFD26864FECCF831C04C89E4" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[377,585,1954,1980]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD26864FECCF831C04C89E4" box="[377,585,1954,1980]" italics="true" pageId="10" pageNumber="82">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD26864FDE5F831C0A889E4" box="[592,685,1954,1980]" pageId="10" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (VNMN 012757).
|
||
<emphasis id="B90CEA9CFFD26864FB93F831C67B89E4" bold="true" box="[1062,1150,1954,1980]" pageId="10" pageNumber="82">B, D, F</emphasis>
|
||
.
|
||
<taxonomicName id="4C784D0DFFD26864FB38F831C3748987" authority="(Bourret, 1937)" baseAuthorityName="Bourret" baseAuthorityYear="1937" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="10" pageNumber="82" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis id="B90CEA9CFFD26864FB38F831C77289E4" box="[1165,1399,1954,1980]" italics="true" pageId="10" pageNumber="82">Theloderma bicolor</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD26864FF76F856C36C8987" author="Bourret R." box="[195,361,1989,2015]" pageId="10" pageNumber="82" pagination="5 - 56" refId="ref7613" refString="Bourret R. 1937. Notes herpetologiques sur l'Indochine francaise. XIV. Les batraciens de la collection du Laboratoire des Sciences Naturelles de l'Universite. Descriptions de quinze especes ou varietes nouvelles. Annexe au Bulletin General de l'Instruction Publique, Hanoi 4: 5 - 56." type="journal article" year="1937">Bourret, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(VNMN 010811). Dorsal (A–B) and ventral views (C–D) as well as lateral head (E–F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC7368EFFD36865FF08FDD2C1D48D76" blockId="11.[189,1400,577,815]" pageId="11" pageNumber="83">
|
||
<taxonomicName id="4C784D0DFFD36865FF08FDD2C38B8C03" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[189,398,577,603]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FF08FDD2C38B8C03" box="[189,398,577,603]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FE23FDD2C3EE8C03" box="[406,491,577,603]" pageId="11" pageNumber="83" rank="species">sp. nov</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C784D0DFFD36865FD3CFDD1C1068C03" authorityName="Taylor" authorityYear="1962" box="[649,771,577,603]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD36865FD3CFDD1C1068C03" box="[649,771,577,603]" italics="true" pageId="11" pageNumber="83">T. gordoni</emphasis>
|
||
</taxonomicName>
|
||
by its coloration pattern: dorsal surface mossy-green or light-olive mixed with mossy-green, resembling moss; ventral yellowish-green band in
|
||
<taxonomicName id="4C784D0DFFD36865FB5CFDF7C2FD8CF9" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FB5CFDF7C2FD8CF9" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FEB4FD1BC35B8CFA" box="[257,350,648,674]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs dorsum dark brown or coffee with some clusters light brown or orange on enlarged gland ridges; ventral surface dark blue with numerous irregular grayish white patterns and speckles in
|
||
<taxonomicName id="4C784D0DFFD36865FF6BFD5DC35C8CB0" authorityName="Taylor" authorityYear="1962" box="[222,345,718,744]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD36865FF6BFD5DC35C8CB0" box="[222,345,718,744]" italics="true" pageId="11" pageNumber="83">T. gordoni</emphasis>
|
||
</taxonomicName>
|
||
. Tympanum and tympanic fold distinct in
|
||
<taxonomicName id="4C784D0DFFD36865FCEBFD5DC6348CB0" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[862,1073,718,744]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FCEBFD5DC6348CB0" box="[862,1073,718,744]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FB8FFD5DC6928CB0" box="[1082,1175,718,744]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs tympanum and tympanic fold invisible in
|
||
<taxonomicName id="4C784D0DFFD36865FE42FD61C06A8D53" authorityName="Taylor" authorityYear="1962" box="[503,623,753,779]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD36865FE42FD61C06A8D53" box="[503,623,753,779]" italics="true" pageId="11" pageNumber="83">T. gordoni</emphasis>
|
||
</taxonomicName>
|
||
; head width equal to head length in
|
||
<taxonomicName id="4C784D0DFFD36865FB94FD62C6F78D53" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[1057,1266,753,779]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FB94FD62C6F78D53" box="[1057,1266,753,779]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FB4CFD62C7508D53" box="[1273,1365,753,779]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs head width greater than head length in
|
||
<taxonomicName id="4C784D0DFFD36865FD33FC86C1C88D76" authority="(Qui et al. 2018)" baseAuthorityName="Qui" baseAuthorityYear="2018" box="[646,973,788,815]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD36865FD33FC86C0FA8D76" box="[646,767,788,814]" italics="true" pageId="11" pageNumber="83">T. gordoni</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD36865FCBBFC87C1C08D76" author="Qui S. & Yu G. H. & Lei B. & Yi F. & Zhang D. L. & Dong Z. W. & Li P. P. & Orlov N. L. & Mian H." box="[782,965,788,815]" pageId="11" pageNumber="83" pagination="43 - 55" refId="ref9788" refString="Qui S., Yu G. H., Lei B., Yi F., Zhang D. L., Dong Z. W., Li P. P., Orlov N. L. & Mian H. 2018. First record of Theloderma gordoni Taylor, 1962 from Yunnan Province, China. Russian Journal of Herpetology 25 (1): 43 - 55. https: // doi. org / 10.30906 / 1026 - 2296 - 2018 - 25 - 1 - 43 - 55" type="journal article" year="2018">
|
||
Qui
|
||
<emphasis id="B90CEA9CFFD36865FCF6FC86C1738D76" box="[835,886,788,814]" italics="true" pageId="11" pageNumber="83">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFD36862FF08FCC6C38E8CDB" lastPageId="12" lastPageNumber="84" pageId="11" pageNumber="83" type="biology_ecology">
|
||
<paragraph id="8BC7368EFFD36865FF08FCC6C3838D28" blockId="11.[189,390,853,880]" box="[189,390,853,880]" pageId="11" pageNumber="83">
|
||
<heading id="D08F81E2FFD36865FF08FCC6C3838D28" bold="true" box="[189,390,853,880]" fontSize="11" level="3" pageId="11" pageNumber="83" reason="3">
|
||
<emphasis id="B90CEA9CFFD36865FF08FCC6C3838D28" bold="true" box="[189,390,853,880]" pageId="11" pageNumber="83">Ecological notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD36862FF08FC17C38E8CDB" blockId="11.[189,1400,900,1032]" lastBlockId="12.[188,1399,265,643]" lastPageId="12" lastPageNumber="84" pageId="11" pageNumber="83">
|
||
<taxonomicName id="4C784D0DFFD36865FF08FC17C38B8DC6" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[189,398,900,926]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FF08FC17C38B8DC6" box="[189,398,900,926]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FE20FC16C3F58DC7" box="[405,496,901,927]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
appears to be closely associated with karstic environments. Specimens were found at night between 19:00 and 23:30 h near cave entrances and in valleys surrounded by limestone cliffs, about
|
||
<quantity id="4C809B6BFFD36865FEE1FC59C3A58DBD" box="[340,416,970,997]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="11" pageNumber="83" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
|
||
from water sources (
|
||
<figureCitation id="13432A0BFFD36865FD2BFC59C0E28DBD" box="[670,743,970,997]" captionStart="Fig" captionStartId="11.[187,230,1954,1980]" captionTargetBox="[189,1398,1109,1916]" captionTargetId="figure-332@11.[189,1400,1109,1916]" captionTargetPageId="11" captionText="Fig. 5. Habitat of Theloderma khoii sp. nov. at the type locality in Quan Ba District, Ha Giang Province, Northern Vietnam." figureDoi="http://doi.org/10.5281/zenodo.6130905" httpUri="https://zenodo.org/record/6130905/files/figure.png" pageId="11" pageNumber="83">Fig. 5</figureCitation>
|
||
). Advertisement calls, eggs and tadpoles of the species have not been recorded during our field surveys. The main habitat at the
|
||
<typeStatus id="54C3882CFFD36865FB98FC7DC6658A50" box="[1069,1120,1006,1032]" pageId="11" pageNumber="83">type</typeStatus>
|
||
locality was secondary karst forest of medium and small hardwoods mixed with shrubs and vines. One specimen was found on a leaf, about
|
||
<quantity id="4C809B6BFFD46862FEE3FEBFC3CB8F1F" box="[342,462,300,327]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="12" pageNumber="84" unit="m" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 m</quantity>
|
||
above the ground, the other specimen was collected on a limestone cliff, about
|
||
<quantity id="4C809B6BFFD46862FF0BFEDCC3308F32" box="[190,309,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="12" pageNumber="84" unit="m" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 m</quantity>
|
||
above the ground. The air temperature at the time of collection ranged from 18.3 to 23.2
|
||
<superScript id="7C0D9BC6FFD46862FAEFFEDCC7728F31" attach="right" box="[1370,1399,335,361]" fontSize="6" pageId="12" pageNumber="84">oC</superScript>
|
||
and relative humidity from 91 to 100%. In Tung Vai, several species of
|
||
<taxonomicName id="4C784D0DFFD46862FC4DFEE1C6838FD4" authorityName="Tschudi" authorityYear="1838" box="[1016,1158,370,396]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEA9CFFD46862FC4DFEE1C6838FD4" box="[1016,1158,370,396]" italics="true" pageId="12" pageNumber="84">Theloderma</emphasis>
|
||
</taxonomicName>
|
||
have been recorded, viz.
|
||
<taxonomicName id="4C784D0DFFD46862FF46FE04C0AD8FE9" authority="(Liu & Hu, 1962)" baseAuthorityName="Liu & Hu" baseAuthorityYear="1962" box="[243,680,406,433]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="albopunctatum">
|
||
<emphasis id="B90CEA9CFFD46862FF46FE04C3C68FE8" box="[243,451,406,432]" italics="true" pageId="12" pageNumber="84">T. albopunctatum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD46862FE60FE05C0A58FE9" author="Liu C. C. & Hu S. Q." box="[469,672,406,433]" pageId="12" pageNumber="84" pagination="73 - 104" refId="ref8927" refString="Liu C. C. & Hu S. Q. 1962. A herpetological report of Kwangsi. Acta Zoologica Sinica 14 (Supplement): 73 - 104." type="journal article" year="1962">Liu & Hu, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D0DFFD46862FD02FE04C1EE8FE9" authority="(Taylor, 1962)" baseAuthorityName="Taylor" baseAuthorityYear="1962" box="[695,1003,406,433]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="gordoni">
|
||
<emphasis id="B90CEA9CFFD46862FD02FE04C1368FE8" box="[695,819,406,432]" italics="true" pageId="12" pageNumber="84">T. gordoni</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD46862FCF0FE05C1E68FE9" author="Taylor E. H." box="[837,995,406,433]" pageId="12" pageNumber="84" pagination="265 - 599" refId="ref10226" refString="Taylor E. H. 1962. The amphibian fauna of Thailand. University of Kansas Science Bulletin 43: 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D0DFFD46862FC4FFE04C3518F8B" authority="(Bain, Nguyen & Doan, 2009)" baseAuthorityName="Bain, Nguyen & Doan" baseAuthorityYear="2009" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="lateriticum">
|
||
<emphasis id="B90CEA9CFFD46862FC4FFE04C69E8FE8" box="[1018,1179,406,432]" italics="true" pageId="12" pageNumber="84">T. lateriticum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD46862FB1BFE05C3498F8B" author="Bain R. H. & Nguyen T. Q. & Doan K. V." pageId="12" pageNumber="84" pagination="58 - 68" refId="ref7513" refString="Bain R. H., Nguyen T. Q. & Doan K. V. 2009. A new species of the genus Theloderma Tschudi, 1838 (Anura: Rhacophoridae) from northwestern Vietnam. Zootaxa 2191 (1): 58 - 68. https: // doi. org / 10.5281 / zenodo. 189385" type="journal article" year="2009">Bain, Nguyen & Doan, 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C784D0DFFD46862FE22FE29C1218F8C" authority="(Liu & Hu, 1962)" baseAuthorityName="Liu & Hu" baseAuthorityYear="1962" box="[407,804,441,468]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="rhododiscus">
|
||
<emphasis id="B90CEA9CFFD46862FE22FE29C0408F8B" box="[407,581,441,467]" italics="true" pageId="12" pageNumber="84">T. rhododiscus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B7FFFD46862FDE1FE2AC11E8F8C" author="Liu C. C. & Hu S. Q." box="[596,795,441,468]" pageId="12" pageNumber="84" pagination="73 - 104" refId="ref8927" refString="Liu C. C. & Hu S. Q. 1962. A herpetological report of Kwangsi. Acta Zoologica Sinica 14 (Supplement): 73 - 104." type="journal article" year="1962">Liu & Hu, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but all were found in tree holes filled with water or on tree leaves near streams in the valleys. Other tree frogs that were found at the same site were
|
||
<taxonomicName id="4C784D0DFFD46862FF08FE6CC3718C42" box="[189,372,511,538]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B90CEA9CFFD46862FF08FE6CC3498C41" box="[189,332,511,537]" italics="true" pageId="12" pageNumber="84">Polypedates</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(of the
|
||
<taxonomicName id="4C784D0DFFD46862FE79FD93C1568C41" authority="Hallowell, 1861" authorityName="Hallowell" authorityYear="1861" box="[460,851,511,537]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B90CEA9CFFD46862FE79FD93C08A8C41" box="[460,655,511,537]" italics="true" pageId="12" pageNumber="84">P. megacephalus</emphasis>
|
||
Hallowell, 1861
|
||
</taxonomicName>
|
||
species complex),
|
||
<taxonomicName id="4C784D0DFFD46862FB9AFE6CC0528C65" authority="(Ohler, Marquis, Swan & Grosjean, 2000)" baseAuthorityName="Ohler, Marquis, Swan & Grosjean" baseAuthorityYear="2000" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="duboisi">
|
||
<emphasis id="B90CEA9CFFD46862FB9AFE6CC7198C41" box="[1071,1308,511,537]" italics="true" pageId="12" pageNumber="84">Zhangixalus duboisi</emphasis>
|
||
(Ohler, Marquis, Swan & Grosjean, 2000)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90CEA9CFFD46862FDD0FDB0C0CB8C64" box="[613,718,546,572]" italics="true" pageId="12" pageNumber="84">Z. jodiae</emphasis>
|
||
Nguyen, Ninh, Orlov, Nguyen & Ziegler, 2020,
|
||
<emphasis id="B90CEA9CFFD46862FAA4FDB0C7728C64" box="[1297,1399,546,572]" italics="true" pageId="12" pageNumber="84">Z. franki</emphasis>
|
||
Ninh, Nguyen, Orlov, Nguyen & Ziegler, 2020,
|
||
<emphasis id="B90CEA9CFFD46862FD40FDD4C1B48C38" box="[757,945,582,608]" italics="true" pageId="12" pageNumber="84">Z. pachyproctus</emphasis>
|
||
Yu, Hui, Hou, Wu, Rao & Yang, 2019 and
|
||
<taxonomicName id="4C784D0DFFD46862FF5AFDFAC38E8CDB" box="[239,395,617,643]" class="Amphibia" family="Rhacophoridae" genus="Kurixalus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B90CEA9CFFD46862FF5AFDFAC3668CDB" box="[239,355,617,643]" italics="true" pageId="12" pageNumber="84">Kurixalus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF076606FFD36865FF0EF831C39389B8" ID-DOI="http://doi.org/10.5281/zenodo.6130905" ID-Zenodo-Dep="6130905" httpUri="https://zenodo.org/record/6130905/files/figure.png" pageId="11" pageNumber="83" startId="11.[187,230,1954,1980]" targetBox="[189,1398,1109,1916]" targetPageId="11">
|
||
<paragraph id="8BC7368EFFD36865FF0EF831C39389B8" blockId="11.[186,1398,1954,2016]" pageId="11" pageNumber="83">
|
||
<emphasis id="B90CEA9CFFD36865FF0EF831C30C89E5" bold="true" box="[187,265,1954,1981]" pageId="11" pageNumber="83">Fig. 5.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="4C784D0DFFD36865FE3EF831C05F89E4" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[395,602,1954,1980]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="11" pageNumber="83" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD36865FE3EF831C05F89E4" box="[395,602,1954,1980]" italics="true" pageId="11" pageNumber="83">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD36865FDD5F830C0BF89E5" box="[608,698,1955,1981]" pageId="11" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
||
at the type locality in Quan Ba District, Ha Giang Province, Northern Vietnam.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3626505FFD46862FF08FD39C76D8D4E" pageId="12" pageNumber="84" type="distribution">
|
||
<paragraph id="8BC7368EFFD46862FF08FD39C3538C9C" blockId="12.[189,342,682,708]" box="[189,342,682,708]" pageId="12" pageNumber="84">
|
||
<heading id="D08F81E2FFD46862FF08FD39C3538C9C" bold="true" box="[189,342,682,708]" fontSize="11" level="3" pageId="12" pageNumber="84" reason="3">
|
||
<emphasis id="B90CEA9CFFD46862FF08FD39C3538C9C" bold="true" box="[189,342,682,708]" pageId="12" pageNumber="84">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD46862FF08FD4AC76D8D4E" blockId="12.[189,1398,729,791]" pageId="12" pageNumber="84">
|
||
<taxonomicName id="4C784D0DFFD46862FF08FD4AC3968CAB" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" box="[189,403,729,755]" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD46862FF08FD4AC3968CAB" box="[189,403,729,755]" italics="true" pageId="12" pageNumber="84">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD46862FE15FD4AC0058CAB" box="[416,512,729,755]" pageId="12" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is currently known only from the
|
||
<typeStatus id="54C3882CFFD46862FC0FFD4AC1E88CAB" box="[954,1005,729,755]" pageId="12" pageNumber="84">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49BCF86CFFD46862FB36FD4AC7778CAB" box="[1155,1394,729,755]" country="Vietnam" name="Ha Giang" pageId="12" pageNumber="84">Ha Giang Province</collectingRegion>
|
||
, northeastern
|
||
<collectingCountry id="F36F761EFFD46862FEE0FD6FC3BD8D4E" box="[341,440,764,790]" name="Vietnam" pageId="12" pageNumber="84">Vietnam</collectingCountry>
|
||
(
|
||
<figureCitation id="13432A0BFFD46862FE7DFD6FC0158D4F" box="[456,528,764,791]" captionStart="Fig" captionStartId="13.[187,230,1885,1911]" captionTargetBox="[285,1297,265,1842]" captionTargetId="figure-14@13.[285,1300,265,1845]" captionTargetPageId="13" captionText="Fig. 6. Map showing distribution of Theloderma bicolor (Bourret, 1937) (blue circles) in Lao Cai and Lai Chau provinces, Vietnam and Jingdong and Luechun counties, Yunnan Province, China (west of the Red River) and the type locality (red circle) of Theloderma khoii sp. nov. in Ha Giang Province, Vietnam (north of the Red River)." figureDoi="http://doi.org/10.5281/zenodo.6130909" httpUri="https://zenodo.org/record/6130909/files/figure.png" pageId="12" pageNumber="84">Fig. 6</figureCitation>
|
||
). The species was recorded at elevations between 1320 and
|
||
<quantity id="4C809B6BFFD46862FB64FD6FC72F8D4F" box="[1233,1322,764,791]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="12" pageNumber="84" unit="m" value="1750.0">1750 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626505FFD46862FF08FCAEC1768A25" pageId="12" pageNumber="84" type="conservation">
|
||
<paragraph id="8BC7368EFFD46862FF08FCAEC3B08D00" blockId="12.[189,437,829,856]" box="[189,437,829,856]" pageId="12" pageNumber="84">
|
||
<heading id="D08F81E2FFD46862FF08FCAEC3B08D00" bold="true" box="[189,437,829,856]" fontSize="11" level="3" pageId="12" pageNumber="84" reason="3">
|
||
<emphasis id="B90CEA9CFFD46862FF08FCAEC3B08D00" bold="true" box="[189,437,829,856]" pageId="12" pageNumber="84">Conservation status</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC7368EFFD46862FF08FCFFC1768A25" blockId="12.[189,1400,876,1149]" pageId="12" pageNumber="84">
|
||
The range of the new species is not expected to extend outside of Mount Tung Vai and it is expected to be found in evergreen forests of
|
||
<collectingRegion id="49BCF86CFFD46862FDF6FC03C1148DF1" box="[579,785,911,938]" country="China" name="Yunnan" pageId="12" pageNumber="84">Yunnan Province</collectingRegion>
|
||
, southern
|
||
<collectingCountry id="F36F761EFFD46862FC38FC1CC1D28DF1" box="[909,983,911,937]" name="China" pageId="12" pageNumber="84">China</collectingCountry>
|
||
. However, the actual distributional range should be the focus of further studies. Currently, we assume that the new species is restricted geographically, likely having an Extent Of Occurrence (EOO) of only <
|
||
<quantity id="4C809B6BFFD46862FB99FC45C6938DA8" box="[1068,1174,982,1008]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="84" unit="km" value="1000.0">1000 km</quantity>
|
||
<superScript id="7C0D9BC6FFD46862FB23FC46C69A8DBB" attach="left" box="[1174,1183,981,995]" fontSize="6" pageId="12" pageNumber="84">2</superScript>
|
||
. The new species likely occurs in one threat-defined location, which has been characterized as an area with a continuing decline in the quality of its habitat due to deforestation (Meyfroidt
|
||
<emphasis id="B90CEA9CFFD46862FC64FB8EC6008A6E" box="[977,1029,1052,1078]" italics="true" pageId="12" pageNumber="84">et al</emphasis>
|
||
. 2013). Therefore,
|
||
<taxonomicName id="4C784D0DFFD46862FB5DFB8FC2FD8A01" authority="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler, 2022" authorityName="Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler" authorityYear="2022" class="Amphibia" family="Rhacophoridae" genus="Theloderma" kingdom="Animalia" order="Anura" pageId="12" pageNumber="84" phylum="Chordata" rank="species" species="khoii" status="sp. nov.">
|
||
<emphasis id="B90CEA9CFFD46862FB5DFB8FC2FD8A01" italics="true" pageId="12" pageNumber="84">Theloderma khoii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57E7FFD46862FF48FBD3C3538A02" box="[253,342,1088,1114]" pageId="12" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
likely qualifies as Endangered (EN) B1ab(iii) in accordance with the categories and criteria of the IUCN Red List of Threatened Species (
|
||
<bibRefCitation id="EFE94B7FFFD46862FD63FBF0C1628A25" author="IUCN" box="[726,871,1123,1149]" pageId="12" pageNumber="84" refId="ref8384" refString="IUCN 2012. The IUCN Red List of Threatened Species. Version 2012.2. Available from http: // www. iucnredlist. org [accessed Oct. 2012]." type="url" year="2012">IUCN 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |