138 lines
17 KiB
XML
138 lines
17 KiB
XML
<document id="D163E6E4CFAEB20B9562CA8F803062CF" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFECFFEE6194FA1E04BFF412" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus artus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="17" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="3E3BEF95A35F7B213281A5F06FEB8990">
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<mods:title id="F6DA2E77B8952DF19212ECB4C1B96306">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="9DFD47272E4F600C23A0AFF0193BD442" type="personal">
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<mods:roleTerm id="CCE05D67BE3102BD449619ACF0EE22C8">Author</mods:roleTerm>
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<mods:namePart id="A4CDBD2AA38E6BE62E250149C069C5D8">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="AD031E9ECD5B4CB3A8053F111A7FB27A">text</mods:typeOfResource>
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<mods:title id="1EDC657469E49AC4C2831B0AB8DC4FFC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E800517CF1BB4FA32149450E4EE28D54">
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<mods:date id="8BE4EF5291B899C4BA943E6017B834A6">2021</mods:date>
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<mods:number id="6A646D758C9AF876BF91C45751D3FAA3">2021-06-28</mods:number>
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<mods:detail id="FBDBBF65329A3B1B847AC78339D3B23F" type="volume">
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<mods:number id="0D7A868F0E7F52320E8D85D41573D848">4994</mods:number>
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<mods:detail id="D461E08C9430100C76F3AE2703FCD350" type="issue">
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<mods:number id="7A1D858313C60D22267AE1F2F09B3110">1</mods:number>
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<mods:start id="4042C7351DC431F132D7064115B46F9E">1</mods:start>
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<mods:end id="FD7B68DD52263CC69F241995AE9CE293">93</mods:end>
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<mods:classification id="18286A6FF0AACC3EC9DEF761E2CCB0CB">journal article</mods:classification>
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<mods:identifier id="D95D9A22BD00CC7582DF2D9DACB25996" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="B8E512FACFB88CD4C31A3B05F86C88B4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D45753064518806770006DF8FF8A5637" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="9D18D0C874CBA6A22B794454ECFF6AD0" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFECFFEE6194FA1E04BFF412" ID-DOI="http://doi.org/10.5281/zenodo.5043940" ID-Zenodo-Dep="5043940" LSID="urn:lsid:plazi:treatment:03A187DEFFECFFEE6194FA1E04BFF412" httpUri="http://treatment.plazi.org/id/03A187DEFFECFFEE6194FA1E04BFF412" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="C3126543FFECFFEF6194FA1E065EF10F" box="[151,513,1518,1545]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BB736C8FFECFFEF6194FA1E065EF10F" blockId="16.[151,513,1518,1581]" box="[151,513,1518,1545]" pageId="16" pageNumber="17">
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<heading id="D0FF81A4FFECFFEF6194FA1E065EF10F" bold="true" box="[151,513,1518,1545]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
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<emphasis id="B97CEADAFFECFFEF6194FA1E065EF10F" bold="true" box="[151,513,1518,1545]" pageId="16" pageNumber="17">
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<taxonomicName id="4C084D4BFFECFFEF6194FA1E053DF10E" ID-CoL="8Y4P7" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,354,1518,1545]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="artus" status="sp. nov.">
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<emphasis id="B97CEADAFFECFFEF6194FA1E053DF10E" bold="true" box="[151,354,1518,1545]" italics="true" pageId="16" pageNumber="17">Hydrochus artus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFECFFEF6072FA1F065EF10F" box="[369,513,1518,1545]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFECFFEF6194F9E3053EF12A" box="[151,353,1554,1581]" pageId="16" pageNumber="17" type="description">
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<paragraph id="8BB736C8FFECFFEF6194F9E3053EF12A" blockId="16.[151,513,1518,1581]" box="[151,353,1554,1581]" pageId="16" pageNumber="17">
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<figureCitation id="13332A4DFFECFFEF6194F9E304BBF12A" box="[151,228,1555,1581]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="16" pageNumber="17">Figs. 3</figureCitation>
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,
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<figureCitation id="13332A4DFFECFFEF61F2F9E3054FF12A" box="[241,272,1555,1581]" captionStart="FIGURES 22–23" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 22–23. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="16" pageNumber="17">22</figureCitation>
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,
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<figureCitation id="13332A4DFFECFFEF6018F9E20566F12A" box="[283,313,1554,1581]" captionStart="FIGURES 59–60" captionStartId="82.[151,264,1677,1701]" captionTargetBox="[157,1431,181,1654]" captionTargetId="figure-28@82.[157,1431,181,1654]" captionTargetPageId="82" captionText="FIGURES 59–60. Geographical distributions. 59. Endemics and near endemics. 60. H. formosus." figureDoi="http://doi.org/10.5281/zenodo.5043360" httpUri="https://zenodo.org/record/5043360/files/figure.png" pageId="16" pageNumber="17">59</figureCitation>
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,
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<figureCitation id="13332A4DFFECFFEF6046F9E3053EF12A" box="[325,353,1555,1581]" captionStart="FIGURE 71" captionStartId="90.[151,250,1796,1820]" captionTargetBox="[166,1425,227,1738]" captionTargetId="figure-28@90.[151,1435,181,1773]" captionTargetPageId="90" captionText="FIGURE 71. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043384" httpUri="https://zenodo.org/record/5043384/files/figure.png" pageId="16" pageNumber="17">71</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFECFFEF6194F9AB0612F19E" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8BB736C8FFECFFEF6194F9AB0612F19E" blockId="16.[151,1437,1626,2013]" pageId="16" pageNumber="17">
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<emphasis id="B97CEADAFFECFFEF6194F9AB0517F173" bold="true" box="[151,328,1626,1653]" pageId="16" pageNumber="17">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFECFFEF604CF9AB0612F19E" collectingDate="1972-05-28" collectionCode="NMNH" collectorName="R. Williams" country="Brazil" county="Varzea Grande County" location="Parque de Exposicao" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
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<emphasis id="B97CEADAFFECFFEF604CF9AB0647F173" bold="true" box="[335,536,1626,1653]" pageId="16" pageNumber="17">
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<typeStatus id="54B3886AFFECFFEF604CF9AB05E3F172" box="[335,444,1627,1653]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFECFFEF632AF9AB06CAF172" box="[553,661,1627,1653]" name="Brazil" pageId="16" pageNumber="17">BRAZIL</collectingCountry>
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:
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<collectingRegion id="49CCF82AFFECFFEF63A3F9AB076BF172" box="[672,820,1627,1653]" country="Brazil" name="Mato Grosso" pageId="16" pageNumber="17">Mato Grosso</collectingRegion>
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,
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<collectingCounty id="62D64E44FFECFFEF623CF9AB001DF172" box="[831,1090,1627,1653]" pageId="16" pageNumber="17">Várzea Grande County</collectingCounty>
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, Cuiabá,
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<location id="8ED76013FFECFFEF65A9F9AB01C6F172" LSID="urn:lsid:plazi:treatment:03A187DEFFECFFEE6194FA1E04BFF412:8ED76013FFECFFEF65A9F9AB01C6F172" box="[1194,1433,1627,1653]" country="Brazil" county="Varzea Grande County" name="Parque de Exposicao" pageId="16" pageNumber="17" stateProvince="Mato Grosso">Parque de Exposição</location>
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,
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<date id="FFB61008FFECFFEF6194F98F0557F19E" box="[151,264,1663,1689]" pageId="16" pageNumber="17" value="1972-05-28">
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<collectingDate id="EFF2E9E0FFECFFEF6194F98F0557F19E" box="[151,264,1663,1689]" pageId="16" pageNumber="17" value="1972-05-28">28.v.1972</collectingDate>
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</date>
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, leg.
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<collectorName id="26FD531EFFECFFEF6047F98F0592F19E" box="[324,461,1663,1689]" pageId="16" pageNumber="17">R. Williams</collectorName>
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” (
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<collectionCode id="ED19AE0DFFECFFEF60E6F98F061EF19E" box="[485,577,1663,1689]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFECFFEF61C4F9520170F02E" pageId="16" pageNumber="17" type="diagnosis">
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<paragraph id="8BB736C8FFECFFEF61C4F9520170F02E" blockId="16.[151,1437,1626,2013]" pageId="16" pageNumber="17">
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<emphasis id="B97CEADAFFECFFEF61C4F952058CF1BB" bold="true" box="[199,467,1698,1724]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of small size (ca.
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<quantity id="4CF09B2DFFECFFEF642BF95201CFF1BA" box="[1320,1424,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="16" pageNumber="17" unit="mm" value="2.15">2.15 mm</quantity>
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), the pronotum with shallow impressions and rather dense punctuation, the pronotal shape, the brown to black color, the usual area of callus on 5
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<superScript id="7C7D9B80FFECFFEF60CDF91A0584F1FF" attach="left" box="[462,475,1770,1784]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
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interstria not raised, and the male genitalia (
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<figureCitation id="13332A4DFFECFFEF62C8F91B0041F002" box="[971,1054,1771,1797]" captionStart="FIGURES 22–23" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 22–23. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="16" pageNumber="17">Fig. 22</figureCitation>
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; described below). Differentiation from other similarly small sized species with similarly shaped pronotum will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFECFFEE61C4F8C20706F5CA" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
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<paragraph id="8BB736C8FFECFFEF61C4F8C200D3F0DA" blockId="16.[151,1437,1626,2013]" pageId="16" pageNumber="17">
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<emphasis id="B97CEADAFFECFFEF61C4F8C2050EF04B" bold="true" box="[199,337,1842,1868]" pageId="16" pageNumber="17">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFECFFEF60A7F8C30659F04A" box="[420,518,1843,1869]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.15/0.78; head width 0.55; pronotum l/w 0.48/0.54; PA 0.54; PB 0.45; elytra 1.32/0.78. Habitus as illustrated (
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<figureCitation id="13332A4DFFECFFEF652AF8A70023F076" box="[1065,1148,1879,1905]" captionStart="FIGURES 22–23" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 22–23. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="16" pageNumber="17">Fig. 22</figureCitation>
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). Dorsum grey to silver or black, with weak iridescence, elytra without black spots. Legs testaceous to brown, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 0.5–1x strial puncture diameter. Interstria 9
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<superScript id="7C7D9B80FFECFFEF6493F86E01C2F0AB" attach="none" box="[1424,1437,1950,1964]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFECFFEF636BF832062AF0D7" attach="left" box="[616,629,1986,2000]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFECFFEF6500F832004FF0D7" attach="left" box="[1027,1040,1986,2000]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
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not raised.
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</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FF670596F7FE" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="18">Pronotum slightly wider than long (ratio ca. 30/26), widest at anterior margin, narrowed at base, sides straight or very slightly sinuate, smooth; anterior margin weakly bisinuate; depressions very shallow; punctation denser in depressions than on reliefs.</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FEF30762F646" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="18">Elytra with spaces between strial punctures ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FEBB0095F662" blockId="17.[151,1437,151,789]" box="[199,1226,331,357]" pageId="17" pageNumber="18">Ventral characters: Mentum punctate, with central depression; submentum with two foveae.</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FE9F07DCF582" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="18">
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Male genitalia (
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<figureCitation id="13332A4DFFEDFFEE607AFE9F0594F68E" box="[377,459,367,393]" captionStart="FIGURES 22–23" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 22–23. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="17" pageNumber="18">Fig. 22</figureCitation>
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) general characters: genitalia small, elongate, length ca. 7x greatest width, basal piece slightly longer than parameres (ratio ca. 4/3); paramere tips extending well beyond aedeagus tip, medial margins beyond aedeagus tip diverging slightly one from the other, very slightly emarginate, plsa absent, lateral margins of tips very slightly arcuate; parameres with distinctive sclerotization; in lateral view parameres markedly tapering in distal 1/2, sharply pointed apically; aedeagus short, narrow in dorsal/ventral views, much wider, oval, in lateral view, in dorsal/ventral views agw at ca. distal 2/3; basal piece in dorsal/ventral views narrowing from distal end to just before orifice, then slightly widened, in lateral view arcuate, maximum width in dorsal/ventral views and lateral view ca. equal, orifice markedly asymmetrical, margin sclerotized.
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</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FD7F0097F5AE" blockId="17.[151,1437,151,789]" box="[199,1224,655,681]" pageId="17" pageNumber="18">Dorsal surface: adtl not apparent; pdmm markedly bisinuate, widely separated; adbl absent.</paragraph>
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<paragraph id="8BB736C8FFEDFFEE61C4FD430706F5CA" blockId="17.[151,1437,151,789]" box="[199,857,691,717]" pageId="17" pageNumber="18">Ventral surface: avtl not apparent; pvmm similar to pdmm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFEDFFEE61C4FD2704BFF412" pageId="17" pageNumber="18" type="etymology">
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<paragraph id="8BB736C8FFEDFFEE61C4FD2704BFF412" blockId="17.[151,1437,151,789]" pageId="17" pageNumber="18">
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<emphasis id="B97CEADAFFEDFFEE61C4FD270517F5F6" bold="true" box="[199,328,727,753]" pageId="17" pageNumber="18">Etymology</emphasis>
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. Named in reference to the parameres, which apically narrow in both dorsal/ventral and lateral views.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |