147 lines
18 KiB
XML
147 lines
18 KiB
XML
<document id="CBB95EF359B4877000D393F27C123272" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE4FFE76194FF670123F295" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus distinctus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="25" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="25" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="6E5079270173EBC67806BDD6351503CC">
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<mods:title id="70B212EFBDE53BF0AF94EED79E8D6213">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="68FB9FC1134BF9D77C3CFBC4AF41962C" type="personal">
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<mods:roleTerm id="1D200C22EA1C1B09135AD7D5D7D26A78">Author</mods:roleTerm>
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<mods:namePart id="A45D6A711F733EF3D0DA1C5E7283E55A">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="6B4CB7892D54763ACDC85291A104A257">text</mods:typeOfResource>
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<mods:title id="B0C4D153AFB86F797F3F8D29919A645E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="072527087238D68E3CAE051E49A08376">
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<mods:date id="12669745CCBAFE1C69659EC7CF54BA8A">2021</mods:date>
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<mods:detail id="A75F0E17A0FA73EBC74F288E16D39A7F" type="pubDate">
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<mods:number id="2F07B169CB0550D0A187A6D86605C6B2">2021-06-28</mods:number>
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<mods:detail id="1B2FF7870869FCAFC4AF39AF7A0AF7D0" type="volume">
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<mods:number id="EB4EEFF3067E7CAABC3CC4CFA4699D22">4994</mods:number>
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<mods:detail id="7BF627477E6DD716EDEA4CF856418F01" type="issue">
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<mods:number id="62BD251FBEC4390F67B2B7AB5CEE296C">1</mods:number>
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<mods:start id="83EF343948960755B2F3A2EC14B45FEF">1</mods:start>
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<mods:end id="4F2372DFED62A3523B1DB4D4277B9C53">93</mods:end>
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<mods:classification id="66FA20B9A68ED12DFEAB31E53B2A0E53">journal article</mods:classification>
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<mods:identifier id="724B5B79EBD61633DE23A63984719BC4" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="53116E90063036D7DC577A77B7371414" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3C6948B62D44176F4AC9011A27E69B04" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="A8576495DE49060F60EF402E56FA0C10" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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</mods:mods>
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<treatment id="03A187DEFFE4FFE76194FF670123F295" ID-DOI="http://doi.org/10.5281/zenodo.5043957" ID-Zenodo-Dep="5043957" LSID="urn:lsid:plazi:treatment:03A187DEFFE4FFE76194FF670123F295" httpUri="http://treatment.plazi.org/id/03A187DEFFE4FFE76194FF670123F295" lastPageNumber="25" pageId="24" pageNumber="25">
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<subSubSection id="C3126543FFE4FFE76194FF670669F7B6" box="[151,566,151,178]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BB736C8FFE4FFE76194FF670669F7B6" blockId="24.[151,566,151,214]" box="[151,566,151,178]" pageId="24" pageNumber="25">
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<heading id="D0FF81A4FFE4FFE76194FF670669F7B6" bold="true" box="[151,566,151,178]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="4">
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<emphasis id="B97CEADAFFE4FFE76194FF670669F7B6" bold="true" box="[151,566,151,178]" pageId="24" pageNumber="25">
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<taxonomicName id="4C084D4BFFE4FFE76194FF6705C7F7B5" ID-CoL="8Y4PZ" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,408,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="distinctus" status="sp. nov.">
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<emphasis id="B97CEADAFFE4FFE76194FF6705C7F7B5" bold="true" box="[151,408,151,178]" italics="true" pageId="24" pageNumber="25">Hydrochus distinctus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE4FFE760A4FF670669F7B6" box="[423,566,151,177]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE4FFE76194FF4B0576F7D1" box="[151,297,187,214]" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BB736C8FFE4FFE76194FF4B0576F7D1" blockId="24.[151,566,151,214]" box="[151,297,187,214]" pageId="24" pageNumber="25">
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<figureCitation id="13332A4DFFE4FFE76194FF4B04BAF7D2" box="[151,229,187,213]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="24" pageNumber="25">Figs. 2</figureCitation>
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,
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<figureCitation id="13332A4DFFE4FFE761F2FF4B04A0F7D2" box="[241,255,187,213]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="24" pageNumber="25">6</figureCitation>
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,
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<figureCitation id="13332A4DFFE4FFE7600EFF4B0576F7D1" box="[269,297,187,214]" captionStart="FIGURES 57–58" captionStartId="81.[151,264,1928,1952]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-15@81.[151,1435,181,1905]" captionTargetPageId="81" captionText="FIGURES 57–58. Geographical distributions. 57. Endemics and near endemics. 58. H. bituberculatus." figureDoi="http://doi.org/10.5281/zenodo.5043356" httpUri="https://zenodo.org/record/5043356/files/figure.png" pageId="24" pageNumber="25">57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE4FFE76194FEF106CEF63A" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="8BB736C8FFE4FFE76194FEF106CEF63A" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">
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<emphasis id="B97CEADAFFE4FFE76194FEF10516F61D" bold="true" box="[151,329,256,283]" pageId="24" pageNumber="25">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE4FFE76051FEF106CEF63A" collectingDate="1971-10-23" collectingDateMax="1971-10-27" collectingDateMin="1971-10-23" collectionCode="NMNH" country="Brazil" county="Cabeca do Veado" elevation="1100" location="Distrito Federal" municipality="Estacao Florestal" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97CEADAFFE4FFE76051FEF10623F61D" bold="true" box="[338,636,256,283]" pageId="24" pageNumber="25">
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<typeStatus id="54B3886AFFE4FFE76051FEF105E0F61C" box="[338,447,257,283]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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(male): “
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<collectingCountry id="F31F7658FFE4FFE76331FEF00623F61D" box="[562,636,256,282]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
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</emphasis>
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:
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<location id="8ED76013FFE4FFE7638FFEF0071EF61D" LSID="urn:lsid:plazi:treatment:03A187DEFFE4FFE76194FF670123F295:8ED76013FFE4FFE7638FFEF0071EF61D" box="[652,833,256,282]" country="Brazil" county="Cabeca do Veado" municipality="Estacao Florestal" name="Distrito Federal" pageId="24" pageNumber="25">Distrito Federal</location>
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,
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<collectingMunicipality id="6BD3ACB2FFE4FFE7624EFEF1004DF61D" box="[845,1042,256,283]" pageId="24" pageNumber="25">Estacao Florestal</collectingMunicipality>
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,
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<collectingCounty id="62D64E44FFE4FFE7651DFEF000BAF61D" box="[1054,1253,256,282]" pageId="24" pageNumber="25">Cabeca do Veado</collectingCounty>
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,
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<quantity id="4CF09B2DFFE4FFE765F1FEF10116F61C" box="[1266,1353,257,283]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="25" unit="m" value="1100.0">
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<elevation id="0025D1FBFFE4FFE765F1FEF10116F61C" box="[1266,1353,257,283]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="25" unit="m" value="1100.0">1100 m</elevation>
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</quantity>
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,
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<date id="FFB61008FFE4FFE76455FEF1055EF63A" pageId="24" pageNumber="25" value="1971-10-23" valueMax="1971-10-27" valueMin="1971-10-23">
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<collectingDate id="EFF2E9E0FFE4FFE76455FEF1055EF63A" pageId="24" pageNumber="25" value="1971-10-23" valueMax="1971-10-27" valueMin="1971-10-23">23–27 Oct.1971</collectingDate>
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</date>
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, E.G., I & E.A. Munroe” (
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<collectionCode id="ED19AE0DFFE4FFE7632AFED306DAF63A" box="[553,645,291,317]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="25" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE4FFE761C4FEB5013EF6A1" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8BB736C8FFE4FFE761C4FEB5013EF6A1" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">
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<emphasis id="B97CEADAFFE4FFE761C4FEB5058CF658" bold="true" box="[199,467,325,351]" pageId="24" pageNumber="25">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of large size (ca. 5.00 mm), the dorsal contrasting colors and sheen, dorsum with head and pronotum dull grey, elytra black and shiny, except each elytron with six dull grey, partially elevated interseries, and the male genitalia (
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<figureCitation id="13332A4DFFE4FFE7654BFE7B00D2F6A2" box="[1096,1165,395,421]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="24" pageNumber="25">Fig. 6</figureCitation>
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; described below).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE4FFE761C4FE5D052CF277" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BB736C8FFE4FFE761C4FE5D0503F407" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">
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<emphasis id="B97CEADAFFE4FFE761C4FE5D050EF6C0" bold="true" box="[199,337,429,455]" pageId="24" pageNumber="25">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFE4FFE760A7FE5E0659F6CF" box="[420,518,430,456]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 4.98/1.65; head width 1.09; pronotum l/w 1.09/1.07; PA 1.07; PB 0.91; elytra 3.26/1.65. Habitus as illustrated (
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<figureCitation id="13332A4DFFE4FFE76533FE20002BF6EC" box="[1072,1140,464,491]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="24" pageNumber="25">Fig. 6</figureCitation>
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). Body size large (ca. 5.00 mm). Dorsum with head and pronotum dull grey, elytra black and shiny, except each elytron with six dull grey, partially elevated interseries; with slight iridescence; elytra without black spots. Legs testaceous to brownish orange, with tibiofemoral joints darker. Punctation on elytra ca. 6–8x that of pronotum. Elytra interstriae ca. 0.5–1.5x strial puncture diameter. Interstria 9
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<superScript id="7C7D9B80FFE4FFE76329FDAB0668F56E" attach="left" box="[554,567,603,617]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFE4FFE76510FDAB007FF56E" attach="left" box="[1043,1056,603,617]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFE4FFE761A6FD8D04EDF58C" attach="left" box="[165,178,637,651]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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well raised, short, only ca. 4 punctures on each side. Each elytron also with six long raised interstriae: 3
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<superScript id="7C7D9B80FFE4FFE76449FD8D0107F58C" attach="none" box="[1354,1368,637,651]" fontSize="6" pageId="24" pageNumber="25">rd</superScript>
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, from base to ca. ¼; 4
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<superScript id="7C7D9B80FFE4FFE76040FD50050FF5A9" attach="none" box="[323,336,672,686]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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, from ca midlength to callus on 5
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<superScript id="7C7D9B80FFE4FFE763C5FD50068CF5A9" attach="none" box="[710,723,672,686]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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; 5
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<superScript id="7C7D9B80FFE4FFE763ECFD5006A3F5A9" attach="none" box="[751,764,672,686]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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, from near base to ca. midlength; 7
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<superScript id="7C7D9B80FFE4FFE76586FD5000CDF5A9" attach="none" box="[1157,1170,672,686]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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, from base to ca. ¼; 8
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<superScript id="7C7D9B80FFE4FFE7648AFD5001C9F5A9" attach="none" box="[1417,1430,672,686]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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, at humeral umbo; 9
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<superScript id="7C7D9B80FFE4FFE76077FD3305DEF5D6" attach="none" box="[372,385,707,721]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
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, for entire length. All of these areas grey to silver grey, dull and contrasting with black shiny surrounding area.
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</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FCF906F0F46F" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">Pronotum only very slightly wider than long (ratio ca. 30/29), widest at anterior margin, narrowed at base, sides sinuate, smooth; anterior margin slightly bisinuate, and with row of short setae; depressions deep to moderately deep; punctation rather uniform over pronotum.</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FC8007B6F4AA" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">Elytra with spaces between strial punctures narrow ridges to ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with angulation.</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FC4605E9F4F5" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">Ventral characters: Mentum and submentum densely, coarsely punctate; mentum with central depression; submentum with two foveae.</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FC0B0524F3E2" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">
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Male genitalia (
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<figureCitation id="13332A4DFFE4FFE7607FFC0B059EF311" box="[380,449,1019,1046]" captionStart="FIGURES 6–7" captionStartId="55.[151,264,1944,1968]" captionTargetBox="[163,1427,205,1900]" captionTargetId="figure-15@55.[151,1435,181,1920]" captionTargetPageId="55" captionText="FIGURES 6–7. Habitus and male genitalia of holotypes. 6. Hydrochus distinctus n. sp. 7. Hydrochus triangularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043278" httpUri="https://zenodo.org/record/5043278/files/figure.png" pageId="24" pageNumber="25">Fig. 6</figureCitation>
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) general characters: genitalia large, elongate, length ca. 5x greatest width, basal piece very slightly longer than parameres; paramere tips extending well beyond aedeagus tip, medial margins beyond aedeagus tip parallel in distal part, then diverging, plsa moderately large, lateral margins between plsa and apex arcuate; in lateral view parameres almost straight, sharply pointed apically; aedeagus moderately wide, agw at about midlength, becoming narrower in basal 1/3; basal piece in dorsal/ventral views nearly parallel-sided, wider in dorsal/ventral views than lateral view, in lateral view very slightly arcuate, orifice slightly asymmetrical, margin strongly sclerotized.
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</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FB1E06FBF22D" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">Dorsal surface: adtl not apparent; pdmm well separated, overlapping alm in distal 2/3 of aedeagus, then approximating alm; adbl large, angulate laterally.</paragraph>
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<paragraph id="8BB736C8FFE4FFE761C4FAC3052CF277" blockId="24.[151,1437,256,1427]" pageId="24" pageNumber="25">Ventral surface: avtl at ca. proximal ¼, margin delimiting large gp; pvmm more widely separated than pdmm, approximating alm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE4FFE761C4FA890123F295" box="[199,1404,1400,1427]" pageId="24" pageNumber="25" type="etymology">
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<paragraph id="8BB736C8FFE4FFE761C4FA890123F295" blockId="24.[151,1437,256,1427]" box="[199,1404,1400,1427]" pageId="24" pageNumber="25">
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<emphasis id="B97CEADAFFE4FFE761C4FA890517F294" bold="true" box="[199,328,1401,1427]" pageId="24" pageNumber="25">Etymology</emphasis>
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. Named in reference to the pleasingly contrasting components of the dorsal sculpture and color.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |