145 lines
18 KiB
XML
145 lines
18 KiB
XML
<document id="1766B8AECD2B659CFE3DFF0AC3360A56" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE0FFE26194F9F707C8F5CA" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus longissimus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="30" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="29" updateTime="1698978858223" updateUser="plazi">
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<mods:title id="98AEE31AC9EFB3E4114F13C6C0D2F7AB">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2996CA4BD3E2305414C479C2B40E94E1" type="personal">
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<mods:roleTerm id="9A8A4008E88A36A98FC93801EA6C0A79">Author</mods:roleTerm>
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<mods:namePart id="D062EE8CBB65F45E6FB6ABD6E4BD902D">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="A06AE4340B75BBF6414E9D90322368DE">text</mods:typeOfResource>
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<mods:title id="211AA138596012CDCE975550210101CF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FCA17EF9CD9557BDE6462B25DE68D61E">
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<mods:date id="2752B76CABCFC73E3563591C0C401DAB">2021</mods:date>
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<mods:detail id="5047BFA69A18058C48E7D8D48918BF88" type="pubDate">
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<mods:number id="E885035E9AFD46E6F704124E5A99B550">2021-06-28</mods:number>
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<mods:detail id="CCFFC822F520B644A991E5BB47C41752" type="volume">
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<mods:number id="428374FF8CF788B303B1A44F5E74A819">4994</mods:number>
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<mods:detail id="6F0EC459F04B2DD4FCE8D7FEC37186FF" type="issue">
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<mods:number id="A3559BFEEC51CE65E7021455197935C9">1</mods:number>
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<mods:start id="2DF808B1A954745A984096BAEFDE32B3">1</mods:start>
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<mods:end id="87DEF2EFF00E8EB41727316B4453669C">93</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="CB50F71EA484921F463024B90644D4E1">journal article</mods:classification>
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<mods:identifier id="AA1EE2A698EC61D182928E4FFBF5C44C" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="9C50152FFF7224C8E34C3B8D172E6347" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9A41D1651B69624E43684282839A55FA" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="8252750590EBBC0E7772E5FBCD064FF9" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFE0FFE26194F9F707C8F5CA" ID-DOI="http://doi.org/10.5281/zenodo.5043968" ID-Zenodo-Dep="5043968" LSID="urn:lsid:plazi:treatment:03A187DEFFE0FFE26194F9F707C8F5CA" httpUri="http://treatment.plazi.org/id/03A187DEFFE0FFE26194F9F707C8F5CA" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
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<subSubSection id="C3126543FFE0FFE36194F9F7060DF125" box="[151,594,1543,1570]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="8BB736C8FFE0FFE36194F9F7060DF125" blockId="28.[151,594,1543,1607]" box="[151,594,1543,1570]" pageId="28" pageNumber="29">
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<heading id="D0FF81A4FFE0FFE36194F9F7060DF125" bold="true" box="[151,594,1543,1570]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
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<emphasis id="B97CEADAFFE0FFE36194F9F7060DF125" bold="true" box="[151,594,1543,1570]" pageId="28" pageNumber="29">
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<taxonomicName id="4C084D4BFFE0FFE36194F9F705ECF125" ID-CoL="8Y4QN" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,435,1543,1570]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longissimus" status="sp. nov.">
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<emphasis id="B97CEADAFFE0FFE36194F9F705ECF125" bold="true" box="[151,435,1543,1570]" italics="true" pageId="28" pageNumber="29">Hydrochus longissimus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE0FFE360C1F9F8060DF125" box="[450,594,1544,1570]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFE0FFE36194F9DC053EF141" box="[151,353,1580,1607]" pageId="28" pageNumber="29" type="description">
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||
<paragraph id="8BB736C8FFE0FFE36194F9DC053EF141" blockId="28.[151,594,1543,1607]" box="[151,353,1580,1607]" pageId="28" pageNumber="29">
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||
<figureCitation id="13332A4DFFE0FFE36194F9DC04BCF141" box="[151,227,1580,1606]" captionStart="FIGURE 1" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[158,1402,188,1903]" captionTargetId="figure-28@6.[152,1435,181,1920]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043263" httpUri="https://zenodo.org/record/5043263/files/figure.png" pageId="28" pageNumber="29">Figs. 1</figureCitation>
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,
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<figureCitation id="13332A4DFFE0FFE361F2F9DC054FF141" box="[241,272,1580,1606]" captionStart="FIGURES 44–45" captionStartId="74.[151,264,1944,1968]" captionTargetBox="[191,1425,216,1871]" captionTargetId="figure-28@74.[151,1435,181,1920]" captionTargetPageId="74" captionText="FIGURES 44–45. Habitus and male genitalia of holotypes. 44. Hydrochus longissimus n. sp. 45. Hydrochus casuarius n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043330" httpUri="https://zenodo.org/record/5043330/files/figure.png" pageId="28" pageNumber="29">44</figureCitation>
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,
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<figureCitation id="13332A4DFFE0FFE36018F9DC0566F140" box="[283,313,1580,1607]" captionStart="FIGURES 57–58" captionStartId="81.[151,264,1928,1952]" captionTargetBox="[151,1435,181,1905]" captionTargetId="figure-15@81.[151,1435,181,1905]" captionTargetPageId="81" captionText="FIGURES 57–58. Geographical distributions. 57. Endemics and near endemics. 58. H. bituberculatus." figureDoi="http://doi.org/10.5281/zenodo.5043356" httpUri="https://zenodo.org/record/5043356/files/figure.png" pageId="28" pageNumber="29">57</figureCitation>
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,
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<figureCitation id="13332A4DFFE0FFE36046F9DC053EF141" box="[325,353,1580,1606]" captionStart="FIGURE 70" captionStartId="89.[151,250,1862,1886]" captionTargetBox="[156,1432,201,1824]" captionTargetId="figure-15@89.[151,1435,181,1837]" captionTargetPageId="89" captionText="FIGURE 70. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043382" httpUri="https://zenodo.org/record/5043382/files/figure.png" pageId="28" pageNumber="29">70</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFE0FFE36194F984059FF1D1" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="8BB736C8FFE0FFE36194F984059FF1D1" blockId="28.[151,1437,1652,2039]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE36194F9840515F189" bold="true" box="[151,330,1652,1678]" pageId="28" pageNumber="29">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE0FFE36051F984059FF1D1" collectingDate="2018-02-21" collectionCode="INPA" country="Brazil" county="Alto Paraiso" elevation="967" latitude="-14.27741" location="Alto Paraiso" longLatPrecision="1" longitude="-47.75075" municipality="Rio Dos Couros" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
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<emphasis id="B97CEADAFFE0FFE36051F9840642F189" bold="true" box="[338,541,1652,1678]" pageId="28" pageNumber="29">
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<typeStatus id="54B3886AFFE0FFE36051F98405E0F189" box="[338,447,1652,1678]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFE0FFE36333F98406C3F189" box="[560,668,1652,1678]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
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:
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<collectingRegion id="49CCF82AFFE0FFE363ABF98406B1F189" box="[680,750,1652,1678]" country="Brazil" name="Goias" pageId="28" pageNumber="29">Goiás</collectingRegion>
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:
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<collectingCounty id="62D64E44FFE0FFE363FAF98407DAF189" box="[761,901,1652,1678]" pageId="28" pageNumber="29">Alto Paraiso</collectingCounty>
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||
/
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<collectingMunicipality id="6BD3ACB2FFE0FFE3629CF9840005F189" box="[927,1114,1652,1679]" pageId="28" pageNumber="29">Rio Dos Couros</collectingMunicipality>
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, nr.
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<location id="8ED76013FFE0FFE36588F9840148F189" LSID="urn:lsid:plazi:treatment:03A187DEFFE0FFE26194F9F707C8F5CA:8ED76013FFE0FFE36588F9840148F189" box="[1163,1303,1652,1678]" country="Brazil" county="Alto Paraiso" latitude="-14.27741" longLatPrecision="1" longitude="-47.75075" municipality="Rio Dos Couros" name="Alto Paraiso" pageId="28" pageNumber="29" stateProvince="Goias">Alto Paraiso</location>
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(
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<geoCoordinate id="EE3C500FFFE0FFE3642BF98401C3F189" box="[1320,1436,1652,1678]" degrees="14.27741" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="-14.27741">-14.27741</geoCoordinate>
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<geoCoordinate id="EE3C500FFFE0FFE36194F9680550F1B4" box="[151,271,1688,1715]" degrees="47.75075" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="-47.75075">-47.75075</geoCoordinate>
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);
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<quantity id="4CF09B2DFFE0FFE36026F9680530F1B4" box="[293,367,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="9.67" pageId="28" pageNumber="29" unit="m" value="967.0">
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<elevation id="0025D1FBFFE0FFE36026F9680530F1B4" box="[293,367,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="9.67" pageId="28" pageNumber="29" unit="m" value="967.0">967 m</elevation>
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</quantity>
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/; small side stream flowing into river, rock/gravel substrate;
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<date id="FFB61008FFE0FFE36541F96800EBF1B5" box="[1090,1204,1688,1714]" pageId="28" pageNumber="29" value="2018-02-21">
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||
<collectingDate id="EFF2E9E0FFE0FFE36541F96800EBF1B5" box="[1090,1204,1688,1714]" pageId="28" pageNumber="29" value="2018-02-21">21.ii.2018</collectingDate>
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</date>
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/ Benetti and team; BR18-0221-01D” (
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<collectionCode id="ED19AE0DFFE0FFE3606CF94C05EAF1D1" box="[367,437,1724,1750]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="28" pageNumber="29">INPA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE0FFE361C4F9100590F08D" pageId="28" pageNumber="29" type="diagnosis">
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<paragraph id="8BB736C8FFE0FFE361C4F9100590F08D" blockId="28.[151,1437,1652,2039]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE361C4F910058BF1FD" bold="true" box="[199,468,1760,1786]" pageId="28" pageNumber="29">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of moderate size (ca.
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<quantity id="4CF09B2DFFE0FFE36468F9100498F018" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="28" pageNumber="29" unit="mm" value="2.76">2.76 mm</quantity>
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), grey to bronze dorsum, with large black spots, the pronotal depressions moderately large, moderately deep, more densely punctate than almost impunctate reliefs, the pronotal shape, the raised 3
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<superScript id="7C7D9B80FFE0FFE36569F8D80027F031" attach="left" box="[1130,1144,1832,1846]" fontSize="6" pageId="28" pageNumber="29">rd</superScript>
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and 5
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<superScript id="7C7D9B80FFE0FFE365C3F8D80092F031" attach="left" box="[1216,1229,1832,1846]" fontSize="6" pageId="28" pageNumber="29">th</superScript>
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elytral interstriae, without usual callus, and the male genitalia (
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<figureCitation id="13332A4DFFE0FFE3639DF8BC06ABF061" box="[670,756,1868,1894]" captionStart="FIGURES 44–45" captionStartId="74.[151,264,1944,1968]" captionTargetBox="[191,1425,216,1871]" captionTargetId="figure-28@74.[151,1435,181,1920]" captionTargetPageId="74" captionText="FIGURES 44–45. Habitus and male genitalia of holotypes. 44. Hydrochus longissimus n. sp. 45. Hydrochus casuarius n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043330" httpUri="https://zenodo.org/record/5043330/files/figure.png" pageId="28" pageNumber="29">Fig. 44</figureCitation>
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; described below). Completely reliable determinations will include dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE0FFE261C4F8640481F5AE" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
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<paragraph id="8BB736C8FFE0FFE261C4F8640647F7FE" blockId="28.[151,1437,1652,2039]" lastBlockId="29.[151,1437,151,717]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE361C4F864050EF0A9" bold="true" box="[199,337,1940,1966]" pageId="28" pageNumber="29">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFE0FFE360A7F8640659F0A9" box="[420,518,1940,1966]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.76/1.03; head width 0.67; pronotum l/w 0.64/0.71; PA 0.64; PB 0.61; elytra 1.83/1.03. Habitus as illustrated (
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<figureCitation id="13332A4DFFE0FFE36548F84800FFF0D5" box="[1099,1184,1976,2002]" captionStart="FIGURES 44–45" captionStartId="74.[151,264,1944,1968]" captionTargetBox="[191,1425,216,1871]" captionTargetId="figure-28@74.[151,1435,181,1920]" captionTargetPageId="74" captionText="FIGURES 44–45. Habitus and male genitalia of holotypes. 44. Hydrochus longissimus n. sp. 45. Hydrochus casuarius n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043330" httpUri="https://zenodo.org/record/5043330/files/figure.png" pageId="28" pageNumber="29">Fig. 44</figureCitation>
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). Dorsum grey to dull silver, with little or no iridescence, elytra with black spots. Legs brown, lateral margin of tibiae with row of spines. Head with frons medially very slightly tumid, with longitudinal punctate groove on each side of tumidity. Punctation on elytra ca. 2–3x that of pronotum. 5
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<superScript id="7C7D9B80FFE1FFE2637EFF4A06D5F7CF" attach="left" box="[637,650,186,200]" fontSize="6" pageId="29" pageNumber="30">th</superScript>
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interstria raised, without usual callus. 9
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<superScript id="7C7D9B80FFE1FFE26553FF4A0002F7CF" attach="left" box="[1104,1117,186,200]" fontSize="6" pageId="29" pageNumber="30">th</superScript>
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interstria more convex than others, overhanging 10
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<superScript id="7C7D9B80FFE1FFE2609AFF2E05F9F7EB" attach="left" box="[409,422,222,236]" fontSize="6" pageId="29" pageNumber="30">th</superScript>
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interstria.
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</paragraph>
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<paragraph id="8BB736C8FFE1FFE261C4FEF30608F662" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="30">Pronotum slightly wider than long (ratio ca. 38/35), widest at anterior margin, narrowed at base, sides weakly sinuate, non-denticulate; anterior margin bisinuate; depressions moderately large, moderately deep, more densely punctate than almost impunctate reliefs.</paragraph>
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<paragraph id="8BB736C8FFE1FFE261C4FE9F054AF6D6" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="30">Elytra with intervals 3, 5, 7, and 9 slightly raised, not costate; punctures large, spaces between strial punctures ca. 1xpd; apices very slightly acuminate, separately rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
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<paragraph id="8BB736C8FFE1FFE261C4FE2B00A9F51E" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="30">Ventral characters: Mentum moderately coarsely and moderately densely punctate, with shallow median fovea; submentum narrow, with transverse groove from side to side, posterior margin carinate, emarginate.</paragraph>
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<paragraph id="8BB736C8FFE1FFE261C4FDD30481F5AE" blockId="29.[151,1437,151,717]" pageId="29" pageNumber="30">
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Male genitalia (
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<figureCitation id="13332A4DFFE1FFE2607AFDD30594F53A" box="[377,459,547,573]" captionStart="FIGURES 44–45" captionStartId="74.[151,264,1944,1968]" captionTargetBox="[191,1425,216,1871]" captionTargetId="figure-28@74.[151,1435,181,1920]" captionTargetPageId="74" captionText="FIGURES 44–45. Habitus and male genitalia of holotypes. 44. Hydrochus longissimus n. sp. 45. Hydrochus casuarius n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043330" httpUri="https://zenodo.org/record/5043330/files/figure.png" pageId="29" pageNumber="30">Fig. 44</figureCitation>
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) general characters: genitalia without parameres; basal piece very elongate, straight in dorsal/ventral views, arcuate in lateral view; distal piece elongate, straight in dorsal/ventral views and lateral view, very slightly widened at apex, with central concavity, without processes; length ratios of basal piece/distal piece ca. 57/25.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE1FFE261C4FD4307C8F5CA" box="[199,919,691,717]" pageId="29" pageNumber="30" type="etymology">
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<paragraph id="8BB736C8FFE1FFE261C4FD4307C8F5CA" blockId="29.[151,1437,151,717]" box="[199,919,691,717]" pageId="29" pageNumber="30">
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<emphasis id="B97CEADAFFE1FFE261C4FD430517F5CA" bold="true" box="[199,328,691,717]" pageId="29" pageNumber="30">Etymology</emphasis>
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. Named in reference to the very long male genitalia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |