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<document id="3F19FE62CFCC4B7F1F0BE032293D448A" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFDCFFDE6194FA57003CF486" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus rectus Perkins 2021, new species" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="33" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="01DF5BC61DE3CA8C9C68689F93EDE1A6">
<mods:title id="34B0F33779F76B5FF86C32336E225BCD">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="E96E07854AECDE291297CD20F1B0D999">Perkins, Philip D.</mods:namePart>
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<mods:title id="40ECE1D6DFC29BCF288D5CD27E23E329">Zootaxa</mods:title>
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<mods:part id="B30EC542CF50F2FE7E6BE354BF94BC8F">
<mods:date id="8C162557810A1BB0E5050D43337EEF64">2021</mods:date>
<mods:detail id="2EEEB22540237CB0C338ADC8E6DB151E" type="pubDate">
<mods:number id="C9EB5E6467E76A49C398B68459A1A4F5">2021-06-28</mods:number>
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<mods:number id="E47D6B6E1C42F032B8E4A6D4525C1345">4994</mods:number>
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<mods:number id="F7F597D4525D5B85AF81DEAF78E39422">1</mods:number>
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<mods:classification id="BDA914B77A118BB4AF764CABD7357FFA">journal article</mods:classification>
<mods:identifier id="8C6580854ACF14E719C9FABFA32D60B6" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
<mods:identifier id="DB9475B70C9E1AC28A74F9EEDE6E00E0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="38D24A68B2CC8253982E37B87B7A357F" type="Zenodo-Dep">5043261</mods:identifier>
<mods:identifier id="1406DE10873670A2EFFBA754536D445B" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFDCFFDE6194FA57003CF486" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDE6194FA57003CF486" httpUri="http://treatment.plazi.org/id/03A187DEFFDCFFDE6194FA57003CF486" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="C3126543FFDCFFDF6194FA570653F2C6" box="[151,524,1447,1474]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BB736C8FFDCFFDF6194FA570653F2C6" blockId="32.[151,524,1447,1510]" box="[151,524,1447,1474]" pageId="32" pageNumber="33">
<heading id="D0FF81A4FFDCFFDF6194FA570653F2C6" bold="true" box="[151,524,1447,1474]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
<emphasis id="B97CEADAFFDCFFDF6194FA570653F2C6" bold="true" box="[151,524,1447,1474]" pageId="32" pageNumber="33">
<taxonomicName id="4C084D4BFFDCFFDF6194FA570532F2C6" ID-CoL="8Y4R8" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,365,1447,1474]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rectus" status="sp. nov.">
<emphasis id="B97CEADAFFDCFFDF6194FA570532F2C6" bold="true" box="[151,365,1447,1474]" italics="true" pageId="32" pageNumber="33">Hydrochus rectus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24F57A1FFDCFFDF607FFA570653F2C6" box="[380,524,1447,1473]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDCFFDF6194FA3B053EF2E2" box="[151,353,1483,1510]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BB736C8FFDCFFDF6194FA3B053EF2E2" blockId="32.[151,524,1447,1510]" box="[151,353,1483,1510]" pageId="32" pageNumber="33">
<figureCitation id="13332A4DFFDCFFDF6194FA3B04BAF2E2" box="[151,229,1483,1509]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="32" pageNumber="33">Figs. 2</figureCitation>
,
<figureCitation id="13332A4DFFDCFFDF61F2FA3B0550F2E2" box="[241,271,1483,1509]" captionStart="FIGURES 2223" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 2223. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="32" pageNumber="33">23</figureCitation>
,
<figureCitation id="13332A4DFFDCFFDF6018FA3B0566F2E1" box="[283,313,1483,1510]" captionStart="FIGURES 5960" captionStartId="82.[151,264,1677,1701]" captionTargetBox="[157,1431,181,1654]" captionTargetId="figure-28@82.[157,1431,181,1654]" captionTargetPageId="82" captionText="FIGURES 5960. Geographical distributions. 59. Endemics and near endemics. 60. H. formosus." figureDoi="http://doi.org/10.5281/zenodo.5043360" httpUri="https://zenodo.org/record/5043360/files/figure.png" pageId="32" pageNumber="33">59</figureCitation>
,
<figureCitation id="13332A4DFFDCFFDF6046FA3B053EF2E2" box="[325,353,1483,1509]" captionStart="FIGURE 71" captionStartId="90.[151,250,1796,1820]" captionTargetBox="[166,1425,227,1738]" captionTargetId="figure-28@90.[151,1435,181,1773]" captionTargetPageId="90" captionText="FIGURE 71. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043384" httpUri="https://zenodo.org/record/5043384/files/figure.png" pageId="32" pageNumber="33">71</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDCFFDF6194F9E30090F1B9" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BB736C8FFDCFFDF6194F9E30090F1B9" blockId="32.[151,1437,1555,2014]" pageId="32" pageNumber="33">
<emphasis id="B97CEADAFFDCFFDF6194F9E30512F12A" bold="true" box="[151,333,1555,1581]" pageId="32" pageNumber="33">Type Material.</emphasis>
<materialsCitation id="3B603C95FFDCFFDF605AF9E30681F156" collectionCode="NMNH" collectorName="M. Grosso &amp; Fritz Plaumann" country="Brazil" latitude="-21.8" location="Rio Caraguata" longLatPrecision="1262" longitude="-52.45" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="B97CEADAFFDCFFDF605AF9E30623F12A" bold="true" box="[345,636,1555,1581]" pageId="32" pageNumber="33">
<typeStatus id="54B3886AFFDCFFDF605AF9E30599F12A" box="[345,454,1555,1581]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="F31F7658FFDCFFDF6330F9E30623F12A" box="[563,636,1555,1581]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
</emphasis>
:
<collectingRegion id="49CCF82AFFDCFFDF6393F9E40776F12A" box="[656,809,1555,1582]" country="Brazil" name="Mato Grosso" pageId="32" pageNumber="33">Mato Grosso</collectingRegion>
, “
<location id="8ED76013FFDCFFDF6240F9E307B1F12A" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDE6194FA57003CF486:8ED76013FFDCFFDF6240F9E307B1F12A" box="[835,1006,1555,1581]" country="Brazil" latitude="-21.8" longLatPrecision="1262" longitude="-52.45" name="Rio Caraguata" pageId="32" pageNumber="33" stateProvince="Mato Grosso">Rio Caraguata</location>
,
<collectorName id="26FD531EFFDCFFDF62FDF9E40023F12A" box="[1022,1148,1555,1582]" pageId="32" pageNumber="33">M. Grosso</collectorName>
, Brazil, IX-53,
<geoCoordinate id="EE3C500FFFDCFFDF643AF9E301C8F12A" box="[1337,1431,1555,1581]" degrees="21" direction="south" minutes="48" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-21.8">21°48S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFDCFFDF6194F9C704A3F155" box="[151,252,1591,1618]" degrees="52" direction="west" minutes="27" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="-52.45">52°27W</geoCoordinate>
, 400M alt.,
<collectorName id="26FD531EFFDCFFDF6080F9C70670F156" box="[387,559,1591,1617]" pageId="32" pageNumber="33">Fritz Plaumann</collectorName>
leg.” (
<collectionCode id="ED19AE0DFFDCFFDF637AF9C7068AF156" box="[633,725,1591,1617]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="33" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B603C95FFDCFFDF63E4F9C701C6F155" box="[743,1433,1591,1618]" collectionCode="NMNH" collectorName="Same" country="Brazil" latitude="-21.8" location="Rio Caraguata" longLatPrecision="1262" longitude="-52.45" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis id="B97CEADAFFDCFFDF63E4F9C707A4F156" bold="true" box="[743,1019,1591,1617]" pageId="32" pageNumber="33">
<typeStatus id="54B3886AFFDCFFDF63E4F9C70700F156" box="[743,863,1591,1617]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
(121):
<collectingCountry id="F31F7658FFDCFFDF62B1F9C707A4F156" box="[946,1019,1591,1617]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
</emphasis>
:
<collectorName id="26FD531EFFDCFFDF6509F9C70017F156" box="[1034,1096,1591,1617]" pageId="32" pageNumber="33">Same</collectorName>
data as holotype (77
<collectionCode id="ED19AE0DFFDCFFDF6435F9C801CEF155" box="[1334,1425,1592,1618]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="33" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B603C95FFDCFFDF6194F9AB010BF171" box="[151,1364,1627,1654]" collectingDate="2004-11-24" collectingDateMax="2004-11-25" collectingDateMin="2004-11-24" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.55" location="Estancia Dimas" longLatPrecision="1246" longitude="-55.216667" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Alto Parana" typeStatus="paratype">
<emphasis id="B97CEADAFFDCFFDF6194F9AB0555F172" bold="true" box="[151,266,1627,1653]" pageId="32" pageNumber="33">
<collectingCountry id="F31F7658FFDCFFDF6194F9AB0555F172" box="[151,266,1627,1653]" name="Paraguay" pageId="32" pageNumber="33">Paraguay</collectingCountry>
</emphasis>
:
<collectingRegion id="49CCF82AFFDCFFDF601AF9AB05FDF171" box="[281,418,1627,1654]" country="Paraguay" name="Alto Parana" pageId="32" pageNumber="33">Alto Parana</collectingRegion>
,
<location id="8ED76013FFDCFFDF60ADF9AB063DF172" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDE6194FA57003CF486:8ED76013FFDCFFDF60ADF9AB063DF172" box="[430,610,1627,1653]" country="Paraguay" latitude="-25.55" longLatPrecision="1246" longitude="-55.216667" name="Estancia Dimas" pageId="32" pageNumber="33" stateProvince="Alto Parana">Estancia Dimas</location>
, (
<geoCoordinate id="EE3C500FFFDCFFDF6374F9AB0689F171" box="[631,726,1627,1654]" degrees="25" direction="south" minutes="33" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-25.55">25°33S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFDCFFDF63E0F9AB0716F171" box="[739,841,1627,1654]" degrees="55" direction="west" minutes="13" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="-55.216667">55°13W</geoCoordinate>
),
<date id="FFB61008FFDCFFDF6261F9AB0058F171" box="[866,1031,1627,1654]" pageId="32" pageNumber="33" value="2004-11-24" valueMax="2004-11-25" valueMin="2004-11-24">
<collectingDate id="EFF2E9E0FFDCFFDF6261F9AB0058F171" box="[866,1031,1627,1654]" pageId="32" pageNumber="33" value="2004-11-24" valueMax="2004-11-25" valueMin="2004-11-24">2425.xi.2004</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDCFFDF6546F9AC0093F172" box="[1093,1228,1627,1654]" pageId="32" pageNumber="33">U. Drechsel</collectorName>
(21
<collectionCode id="ED19AE0DFFDCFFDF6401F9AC0115F171" box="[1282,1354,1628,1654]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFDCFFDF6460F9AC06ADF19D" collectingDate="2005-02-17" collectingDateMax="2005-02-22" collectingDateMin="2005-02-17" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.55" location="Estancia Dimas" longLatPrecision="1246" longitude="-55.216667" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Alto Parana" typeStatus="paratype">
same locality,
<date id="FFB61008FFDCFFDF61F8F98F05C6F19D" box="[251,409,1663,1690]" pageId="32" pageNumber="33" value="2005-02-17" valueMax="2005-02-22" valueMin="2005-02-17">
<collectingDate id="EFF2E9E0FFDCFFDF61F8F98F05C6F19D" box="[251,409,1663,1690]" pageId="32" pageNumber="33" value="2005-02-17" valueMax="2005-02-22" valueMin="2005-02-17">1722.ii.2005</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDCFFDF60DFF9700639F19E" box="[476,614,1663,1690]" pageId="32" pageNumber="33">U. Drechsel</collectorName>
(17
<collectionCode id="ED19AE0DFFDCFFDF63A3F97006B7F19D" box="[672,744,1664,1690]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFDCFFDF6200F9700605F1B9" collectingDate="2005-08-27" collectingDateMax="2005-08-29" collectingDateMin="2005-08-27" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.133333" location="Puerto Galileo" longLatPrecision="1247" longitude="-57.833332" municipality="Dep. Pte. Hayes" pageId="32" pageNumber="33" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BD3ACB2FFDCFFDF6200F970079EF19D" box="[771,961,1664,1690]" pageId="32" pageNumber="33">Dep. Pte. Hayes</collectingMunicipality>
,
<location id="8ED76013FFDCFFDF62D3F9700027F19E" LSID="urn:lsid:plazi:treatment:03A187DEFFDCFFDE6194FA57003CF486:8ED76013FFDCFFDF62D3F9700027F19E" box="[976,1144,1663,1690]" country="Paraguay" latitude="-25.133333" longLatPrecision="1247" longitude="-57.833332" municipality="Dep. Pte. Hayes" name="Puerto Galileo" pageId="32" pageNumber="33">Puerto Galileo</location>
, (
<geoCoordinate id="EE3C500FFFDCFFDF6592F98F00BEF19D" box="[1169,1249,1663,1690]" degrees="25" direction="south" minutes="8" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="-25.133333">25°8S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFDCFFDF65F3F98F0108F19D" box="[1264,1367,1663,1690]" degrees="57" direction="west" minutes="50" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="-57.833332">57°50W</geoCoordinate>
),
<date id="FFB61008FFDCFFDF6471F98F057EF1B9" pageId="32" pageNumber="33" value="2005-08-27" valueMax="2005-08-29" valueMin="2005-08-27">
<collectingDate id="EFF2E9E0FFDCFFDF6471F98F057EF1B9" pageId="32" pageNumber="33" value="2005-08-27" valueMax="2005-08-29" valueMin="2005-08-27">27 29.viii.2005</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDCFFDF605EF95405BCF1BA" box="[349,483,1699,1726]" pageId="32" pageNumber="33">U. Drechsel</collectorName>
(4
<collectionCode id="ED19AE0DFFDCFFDF630BF954060FF1B9" box="[520,592,1700,1726]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFDCFFDF6364F9540095F1B9" box="[615,1226,1699,1726]" collectingDate="2005-11-01" collectingDateMax="2005-11-03" collectingDateMin="2005-11-01" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.133333" location="Puerto Galileo" longLatPrecision="1247" longitude="-57.833332" municipality="Dep. Pte. Hayes" pageId="32" pageNumber="33" specimenCount="1" typeStatus="paratype">
same locality,
<date id="FFB61008FFDCFFDF620BF95307CFF1B9" box="[776,912,1699,1726]" pageId="32" pageNumber="33" value="2005-11-01" valueMax="2005-11-03" valueMin="2005-11-01">
<collectingDate id="EFF2E9E0FFDCFFDF620BF95307CFF1B9" box="[776,912,1699,1726]" pageId="32" pageNumber="33" value="2005-11-01" valueMax="2005-11-03" valueMin="2005-11-01">13.xi.2005</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFDCFFDF62CFF954000DF1BA" box="[972,1106,1699,1726]" pageId="32" pageNumber="33">U. Drechsel</collectorName>
(2
<collectionCode id="ED19AE0DFFDCFFDF6574F95400E0F1B9" box="[1143,1215,1700,1726]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="32" pageNumber="33">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDCFFDF61C4F937076EF04A" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BB736C8FFDCFFDF61C4F937076EF04A" blockId="32.[151,1437,1555,2014]" pageId="32" pageNumber="33">
<emphasis id="B97CEADAFFDCFFDF61C4F937058CF1E6" bold="true" box="[199,467,1735,1761]" pageId="32" pageNumber="33">Differential Diagnosis.</emphasis>
Among Brazilian species, recognized by the combination of small size (ca.
<quantity id="4CF09B2DFFDCFFDF642BF93701CFF1E5" box="[1320,1424,1735,1762]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="32" pageNumber="33" unit="mm" value="2.17">2.17 mm</quantity>
), the pronotum with shallow impressions and rather dense punctuation, the pronotal shape, the brown to black color, and the male genitalia (
<figureCitation id="13332A4DFFDCFFDF60AAF8FF065FF02E" box="[425,512,1807,1833]" captionStart="FIGURES 2223" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 2223. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
; described below). Differentiation from other similarly small sized species with similarly shaped pronotum will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDCFFDE61C4F8A70089F45A" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8BB736C8FFDCFFDE61C4F8A701D7F7B6" blockId="32.[151,1437,1555,2014]" lastBlockId="33.[151,1437,150,897]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B97CEADAFFDCFFDF61C4F8A7050EF076" bold="true" box="[199,337,1879,1905]" pageId="32" pageNumber="33">Description</emphasis>
. Size:
<typeStatus id="54B3886AFFDCFFDF60A7F8A70659F076" box="[420,518,1879,1905]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.17/0.77; head width 0.55; pronotum l/w 0.50/0.54; PA 0.54; PB 0.46; elytra 1.36/0.77. Habitus as illustrated (
<figureCitation id="13332A4DFFDCFFDF654FF88B00FDF092" box="[1100,1186,1915,1941]" captionStart="FIGURES 2223" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 2223. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
). Dorsum dark brown to black, with moderately strong purple iridescence on head, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 23x that of pronotum. Elytra interstriae ca. 0.51x strial puncture diameter. Interstria 9
<superScript id="7C7D9B80FFDDFFDE6083FF6605D2F7A3" attach="left" box="[384,397,150,164]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
more convex than others, overhanging 10
<superScript id="7C7D9B80FFDDFFDE6267FF66072EF7A3" attach="left" box="[868,881,150,164]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C7D9B80FFDDFFDE65FCFF660153F7A3" attach="left" box="[1279,1292,150,164]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
not raised.
</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FF4B0697F61A" blockId="33.[151,1437,150,897]" pageId="33" pageNumber="34">Pronotum slightly wider than long (ratio ca. 31/28), widest at anterior margin, narrowed at base, sides straight, with minute denticulations; anterior margin bisinuate; depressions very shallow; punctation dense, rather uniform over pronotum, but slightly denser in depressions.</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FED70584F68E" blockId="33.[151,1437,150,897]" pageId="33" pageNumber="34">Elytra with spaces between strial punctures narrow ridges to ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation; series 15 each terminating in larger, translucent punctures.</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FE630002F6AA" blockId="33.[151,1437,150,897]" box="[199,1117,403,429]" pageId="33" pageNumber="34">Ventral characters: Mentum with central depression; submentum with two foveae.</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FE47061BF5CA" blockId="33.[151,1437,150,897]" pageId="33" pageNumber="34">
Male genitalia (
<figureCitation id="13332A4DFFDDFFDE607BFE470595F6D6" box="[376,458,439,465]" captionStart="FIGURES 2223" captionStartId="63.[151,264,1944,1968]" captionTargetBox="[185,1415,201,1871]" captionTargetId="figure-15@63.[151,1435,181,1920]" captionTargetPageId="63" captionText="FIGURES 2223. Habitus and male genitalia of holotypes. 22. Hydrochus artus n. sp. 23. Hydrochus rectus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043302" httpUri="https://zenodo.org/record/5043302/files/figure.png" pageId="33" pageNumber="34">Fig. 23</figureCitation>
) general characters: genitalia very small, stout, length ca. 4x greatest width; parameres extending well beyond distal end of aedeagus, tips in dorsal/ventral views wide in proximal part, narrowing only slightly apically, apices blunt, medial margins straight, slightly diverging one from the other, in lateral view tips arcuate slightly toward ventrad, and markedly narrowing apically; basal piece long, longer than parameres (ratio ca. 7/5), sides slightly arcuate, tapering from distal end to base, constricted just before orifice, in lateral view arcuate, orifice markedly asymmetrical, with rim weakly sclerotized and characteristically shaped; aedeagus in dorsal/ventral views narrow, narrowing slightly from base to pointed apex, in lateral view much wider, sides arcuate, orifice markedly asymmetrical, hook-shaped.
</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FD270567F43E" blockId="33.[151,1437,150,897]" pageId="33" pageNumber="34">Dorsal surface: adtl lacking; pdmm distinctively shaped, diverging one from the other at aedeagal apex, then widening to become contiguous, or slightly overlap one another, aedeagal apex visible in “window” created by pdmm shapes.</paragraph>
<paragraph id="8BB736C8FFDDFFDE61C4FCB30089F45A" blockId="33.[151,1437,150,897]" box="[199,1238,835,861]" pageId="33" pageNumber="34">Ventral surface: pvmm more widely separated than pdmm, contiguous with or nearly so alm.</paragraph>
</subSubSection>
<subSubSection id="C3126543FFDDFFDE61C4FC97003CF486" box="[199,1123,871,897]" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BB736C8FFDDFFDE61C4FC97003CF486" blockId="33.[151,1437,150,897]" box="[199,1123,871,897]" pageId="33" pageNumber="34">
<emphasis id="B97CEADAFFDDFFDE61C4FC970517F486" bold="true" box="[199,328,871,897]" pageId="33" pageNumber="34">Etymology</emphasis>
. Named in reference to the straight parameres, in dorsal/ventral views.
</paragraph>
</subSubSection>
</treatment>
</document>