treatments-xml/data/03/A1/87/03A187CAFFC894251FEAFDE390D7FBF4.xml
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<mods:title id="E29E28D0656DEC28D1A6EF288EE8925A">On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:date id="70D4DCDDF61F389AFB198E4F580A232D">2018</mods:date>
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<subSubSection id="C3126557FFC894251FEAFDE390E5FDFA" box="[178,674,555,577]" pageId="16" pageNumber="165" type="nomenclature">
<paragraph id="8BB736DCFFC894251FEAFDE390E5FDFA" blockId="16.[178,674,555,577]" box="[178,674,555,577]" pageId="16" pageNumber="165">
<heading id="D0FF81B0FFC894251FEAFDE390E5FDFA" box="[178,674,555,577]" fontSize="8" level="2" pageId="16" pageNumber="165" reason="2">
<emphasis id="B97CEACEFFC894251FEAFDE39024FDFA" bold="true" box="[178,611,555,577]" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251FEAFDE393B5FDFB" ID-CoL="4R35H" authorityName="Stephens" authorityYear="1832" box="[178,498,555,577]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="humeralis" subGenus="Raphirus">
<emphasis id="B97CEACEFFC894251FEAFDE3934EFDFB" bold="true" box="[178,265,555,577]" italics="true" pageId="16" pageNumber="165">Quedius</emphasis>
(
<emphasis id="B97CEACEFFC894251E40FDE3933EFDFB" bold="true" box="[280,377,555,577]" italics="true" pageId="16" pageNumber="165">Raphirus</emphasis>
)
<emphasis id="B97CEACEFFC894251ED0FDE393B5FDFB" bold="true" box="[392,498,555,577]" italics="true" pageId="16" pageNumber="165">humeralis</emphasis>
</taxonomicName>
<bibRefCitation id="EF994B2DFFC894251EA0FDE39024FDFA" author="STEPHENS J" box="[504,611,555,577]" pageId="16" pageNumber="165" pagination="1 - 240" refId="ref21827" refString="STEPHENS J. F. (1832): Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V. - Baldwin &amp; Cradock: 1 - 240." type="book chapter" year="1832">STEPHENS</bibRefCitation>
</emphasis>
,
<emphasis id="B97CEACEFFC894251D37FDE390E5FDFA" bold="true" box="[623,674,555,576]" pageId="16" pageNumber="165">1832</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFC894251FEAFD9E908AFC82" pageId="16" pageNumber="165" type="reference_group">
<paragraph id="8BB736DCFFC894251FEAFD9E906AFDD6" blockId="16.[178,1086,598,824]" box="[178,557,598,620]" pageId="16" pageNumber="165">
<treatmentCitationGroup id="AB1811F2FFC894251FEAFD9E906AFDD6" box="[178,557,598,620]" pageId="16" pageNumber="165">
<treatmentCitation id="0AA910CDFFC894251FEAFD9E906DFDD6" author="STEPHENS J" box="[178,554,598,620]" page="220" pageId="16" pageNumber="165" year="1832">
<taxonomicName id="4C084D5FFFC894251FEAFD9E906DFDD6" ID-CoL="4R35H" authority="STEPHENS, 1832: 220" authorityName="Stephens" authorityYear="1832" box="[178,554,598,620]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B97CEACEFFC894251FEAFD9E9319FDD1" box="[178,350,598,619]" italics="true" pageId="16" pageNumber="165">Quedius humeralis</emphasis>
<bibRefCitation id="EF994B2DFFC894251E3CFD9F906DFDD6" author="STEPHENS J" box="[356,554,599,620]" pageId="16" pageNumber="165" pagination="1 - 240" refId="ref21827" refString="STEPHENS J. F. (1832): Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. V. - Baldwin &amp; Cradock: 1 - 240." type="book chapter" year="1832">STEPHENS, 1832: 220</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFDB8907AFD3C" blockId="16.[178,1086,598,824]" box="[178,573,624,646]" pageId="16" pageNumber="165">
<treatmentCitationGroup id="AB1811F2FFC894251FEAFDB8907AFD3C" box="[178,573,624,646]" pageId="16" pageNumber="165">
<emphasis id="B97CEACEFFC894251FEAFDB8932BFD3F" box="[178,364,624,645]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251FEAFDB892B9FD3F" ID-CoL="8VS2M" box="[178,254,624,645]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
oblitteratus
</emphasis>
<treatmentCitation id="0AA910CDFFC894251E2AFDB9907EFD3C" author="ERICHSON W" box="[370,569,624,646]" page="549" pageId="16" pageNumber="165" year="1840">
<bibRefCitation id="EF994B2DFFC894251E2AFDB9907EFD3C" author="ERICHSON W" box="[370,569,624,646]" pageId="16" pageNumber="165" pagination="401 - 954" refId="ref20470" refString="ERICHSON W. F. (1840): Genera et species Staphylinorum insectorum coleopterorum familiae. - Morin, Berlin: 401 - 954." type="book chapter" year="1840">ERICHSON, 1840: 549</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFD439027FD1A" blockId="16.[178,1086,598,824]" box="[178,608,651,673]" pageId="16" pageNumber="165">
<treatmentCitationGroup id="AB1811F2FFC894251FEAFD439027FD1A" box="[178,608,651,673]" pageId="16" pageNumber="165">
<emphasis id="B97CEACEFFC894251FEAFD439304FD1A" box="[178,323,651,672]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251FEAFD4392B9FD1A" ID-CoL="8VS2M" box="[178,254,651,672]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
coxalis
</emphasis>
<treatmentCitation id="0AA910CDFFC894251E11FD4393B4FD1B" author="KRAATZ G" box="[329,499,651,673]" page="59" pageId="16" pageNumber="165" year="1858">
<bibRefCitation id="EF994B2DFFC894251E11FD4393B4FD1B" author="KRAATZ G" box="[329,499,651,673]" pageId="16" pageNumber="165" pagination="37 - 67" refId="ref21051" refString="KRAATZ G. (1858): Beitrag zur Kaferfauna Griechenlands. Zweites Stuck: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. - Berliner Entomologische Zeitschrift 2: 37 - 67." type="journal article" year="1858">
KRAATZ, 1858:
<specimenCount id="9D0EFD55FFC894251E84FD4393B4FD1B" box="[476,499,651,673]" pageId="16" pageNumber="165" type="generic">59</specimenCount>
</bibRefCitation>
</treatmentCitation>
f.;
<emphasis id="B97CEACEFFC894251D49FD439027FD1A" bold="true" box="[529,608,651,672]" pageId="16" pageNumber="165">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFD6D904CFD01" blockId="16.[178,1086,598,824]" box="[178,523,677,699]" pageId="16" pageNumber="165">
<emphasis id="B97CEACEFFC894251FEAFD6D9305FD00" box="[178,322,677,698]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251FEAFD6D92B9FD00" ID-CoL="8VS2M" box="[178,254,677,698]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
gestroi
</emphasis>
GRIDELLI, 1922: 136.
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFD779028FD6E" blockId="16.[178,1086,598,824]" box="[178,623,703,725]" pageId="16" pageNumber="165">
<treatmentCitationGroup id="AB1811F2FFC894251FEAFD779028FD6E" box="[178,623,703,725]" pageId="16" pageNumber="165">
<emphasis id="B97CEACEFFC894251FEAFD779378FD6F" box="[178,319,703,725]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251FEAFD7792B9FD6E" ID-CoL="8VS2M" box="[178,254,703,724]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
atticus
</emphasis>
<treatmentCitation id="0AA910CDFFC894251E1DFD779045FD6E" author="COIFFAIT H" box="[325,514,703,725]" page="408" pageId="16" pageNumber="165" year="1967">
<bibRefCitation id="EF994B2DFFC894251E1DFD779045FD6E" author="COIFFAIT H" box="[325,514,703,725]" pageId="16" pageNumber="165" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">
COIFFAIT, 1967:
<specimenCount id="9D0EFD55FFC894251EB8FD779045FD6E" box="[480,514,703,724]" pageId="16" pageNumber="165" type="generic">408</specimenCount>
</bibRefCitation>
</treatmentCitation>
f.;
<emphasis id="B97CEACEFFC894251D78FD779028FD6E" bold="true" box="[544,623,703,724]" pageId="16" pageNumber="165">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFD16908AFC82" blockId="16.[178,1086,598,824]" pageId="16" pageNumber="165">
<materialsCitation id="3B603C81FFC894251FEAFD16908AFC82" ID-GBIF-Occurrence="2846796409" collectionCode="SDEI" collectorName="Coll. Kraatz &amp; G. Fagel" determinerName="E. Gridelli" pageId="16" pageNumber="165" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
T y p e m a t e r i a l e x a m i n e d:
<typeStatus id="54B3887EFFC894251D3CFD1690F8FD49" box="[612,703,734,755]" pageId="16" pageNumber="165" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D0EFD55FFC894251D95FD169099FD49" box="[717,734,734,755]" pageId="16" pageNumber="165" type="male"></specimenCount>
[dissected prior to present study]: &quot;Graecia / Typus /
<emphasis id="B97CEACEFFC894251E31FD3C93EEFCB3" box="[361,425,756,777]" italics="true" pageId="16" pageNumber="165">coxalis</emphasis>
mihi /
<collectorName id="26FD530AFFC894251EABFD3C9023FCB0" box="[499,612,756,778]" pageId="16" pageNumber="165">Coll. Kraatz</collectorName>
/
<emphasis id="B97CEACEFFC894251D24FD3D9087FCB0" box="[636,704,757,778]" italics="true" pageId="16" pageNumber="165">Gestroi</emphasis>
mihi, det.
<determinerName id="7E6BDA18FFC894251C72FD3D91CAFCB0" box="[810,909,756,778]" pageId="16" pageNumber="165">E. Gridelli</determinerName>
/ Dtsch. Ent. Inst. Eberswalde /
<collectorName id="26FD530AFFC894251E68FCC39338FC9B" box="[304,383,779,801]" pageId="16" pageNumber="165">G. Fagel</collectorName>
elig. 1968,
<emphasis id="B97CEACEFFC894251EB6FCC390C6FC9A" box="[494,641,779,800]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251EB6FCC3907EFC9A" ID-CoL="8VS2M" box="[494,569,779,800]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
coxalis
</emphasis>
Kr.
<typeStatus id="54B3887EFFC894251DF4FCC49140FC9B" box="[684,775,780,801]" pageId="16" pageNumber="165" type="lectotype">Lectotype</typeStatus>
/ DEI Müncheberg Col - 10021 /
<taxonomicName id="4C084D5FFFC894251FEAFCEA93FFFC82" ID-CoL="4R35H" authority="Stephens" authorityName="Stephens" authorityYear="1832" box="[178,440,802,824]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B97CEACEFFC894251FEAFCEA9319FC8D" box="[178,350,802,823]" italics="true" pageId="16" pageNumber="165">Quedius humeralis</emphasis>
Stephens
</taxonomicName>
, det. V.
<bibRefCitation id="EF994B2DFFC894251D5BFCEA9030FC8D" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[515,631,802,824]" pageId="16" pageNumber="165" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (
<collectionCode id="ED19AE19FFC894251DD3FCEA9085FC82" box="[651,706,802,824]" pageId="16" pageNumber="165">SDEI</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFC894251FEAFC8E90D7FBF4" pageId="16" pageNumber="165" type="description">
<paragraph id="8BB736DCFFC894251FEAFC8E91BBFC55" blockId="16.[178,1085,838,1007]" pageId="16" pageNumber="165">
C o m m e n t: The original description of
<emphasis id="B97CEACEFFC894251DCBFC8E90B8FCE1" box="[659,767,838,859]" italics="true" pageId="16" pageNumber="165">Q. coxalis</emphasis>
is based on several
<typeStatus id="54B3887EFFC894251CBCFC8E967AFCE1" box="[996,1085,838,859]" pageId="16" pageNumber="165" type="syntype">syntypes</typeStatus>
(&quot;nicht gerade selten&quot;) from &quot;Athen&quot; (
<bibRefCitation id="EF994B2DFFC894251D67FCAB9095FCC3" author="KRAATZ G" box="[575,722,867,889]" pageId="16" pageNumber="165" pagination="37 - 67" refId="ref21051" refString="KRAATZ G. (1858): Beitrag zur Kaferfauna Griechenlands. Zweites Stuck: Palpicornia, Silphales, Scydmaenidae, Pselaphidae, Staphylinidae. - Berliner Entomologische Zeitschrift 2: 37 - 67." type="journal article" year="1858">KRAATZ 1858</bibRefCitation>
). There is, however, only one
<typeStatus id="54B3887EFFC894251B4AFCAC9299FC2C" pageId="16" pageNumber="165" type="syntype">syntype</typeStatus>
in the Kraatz collection. This specimen, a male, was examined and designated as the
<typeStatus id="54B3887EFFC894251FEAFC569348FC0E" box="[178,271,926,948]" pageId="16" pageNumber="165" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EF994B2DFFC894251E6EFC569394FC0E" author="FAGEL G" box="[310,467,926,948]" pageId="16" pageNumber="165" pagination="1 - 14" refId="ref20541" refString="FAGEL G. (1968 b): Contribution a la connaissance des Staphylinidae CIII. - Le complexe des Quedius oblitteratus ER. - nemoralis BAUDI. - Bulletin Institut Royale des Sciences Naturelles de Belgique 44 (24): 1 - 14." type="journal article" year="1968">FAGEL (1968b)</bibRefCitation>
.
<bibRefCitation id="EF994B2DFFC894251E85FC5690C2FC0E" author="COIFFAIT H" box="[477,645,926,948]" pageId="16" pageNumber="165" pagination="391 - 424" refId="ref20253" refString="COIFFAIT H. (1967): Quedius nouveaux ou mal connus. - Bulletin de la Societe d'Histoire Naturelle de Toulouse 103 (3 - 4): 391 - 424." type="journal article" year="1967">COIFFAIT (1967)</bibRefCitation>
described
<emphasis id="B97CEACEFFC894251DABFC569110FC0E" box="[755,855,926,948]" italics="true" pageId="16" pageNumber="165">Q. atticus</emphasis>
based on a male
<typeStatus id="54B3887EFFC894251B5FFC569299FC6B" pageId="16" pageNumber="165" type="holotype">holotype</typeStatus>
and seven
<typeStatus id="54B3887EFFC894251E32FC74938CFC6B" box="[362,459,956,977]" pageId="16" pageNumber="165" type="paratype">paratypes</typeStatus>
(a male and six females) from &quot;Mont Pantélique près d'Athénes&quot;.
<emphasis id="B97CEACEFFC894251E75FC11938BFC54" box="[301,460,985,1006]" italics="true" pageId="16" pageNumber="165">
<taxonomicName id="4C084D5FFFC894251E75FC1193C6FC54" box="[301,385,985,1006]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
atticus
</emphasis>
was synonymized with
<emphasis id="B97CEACEFFC894251D98FC119160FC54" box="[704,807,985,1006]" italics="true" pageId="16" pageNumber="165">Q. coxalis</emphasis>
by
<bibRefCitation id="EF994B2DFFC894251C15FC1191BFFC55" author="COIFFAIT H" box="[845,1016,985,1007]" pageId="16" pageNumber="165" pagination="133 - 143" refId="ref20345" refString="COIFFAIT H. (1977): Note sur quelques Quedius et Heterothops nouveaux ou mal connus (Col. Staphylinidae). - Nouvell e Revue d'Entomologie 7 (2): 133 - 143." type="journal article" year="1977">COIFFAIT (1977)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB736DCFFC894251FEAFC3690D7FBF4" blockId="16.[178,1085,1022,1103]" pageId="16" pageNumber="165">
An examination of the
<typeStatus id="54B3887EFFC894251EF3FC36904FFBAE" box="[427,520,1022,1044]" pageId="16" pageNumber="165" type="lectotype">lectotype</typeStatus>
of
<emphasis id="B97CEACEFFC894251D69FC3690DBFBA9" box="[561,668,1022,1043]" italics="true" pageId="16" pageNumber="165">Q. coxalis</emphasis>
revealed that it is conspecific with the widespread and common
<taxonomicName id="4C084D5FFFC894251EEEFBD4907DFB8A" authorityName="Stephens" authorityYear="1832" box="[438,570,1051,1073]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="165" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B97CEACEFFC894251EEEFBD4907DFB8A" box="[438,570,1051,1073]" italics="true" pageId="16" pageNumber="165">Q. humeralis</emphasis>
</taxonomicName>
. Hence, both
<emphasis id="B97CEACEFFC894251D9FFBD49169FB8A" box="[711,814,1051,1073]" italics="true" pageId="16" pageNumber="165">Q. coxalis</emphasis>
and its junior synonym
<emphasis id="B97CEACEFFC894251B7CFBD492B2FBF4" italics="true" pageId="16" pageNumber="165">Q. atticus</emphasis>
are placed in synonymy with this name.
</paragraph>
</subSubSection>
</treatment>
</document>