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<document id="FF0DD98CD957A95AC04D43DBB05A074C" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A9292FFFE9FF03C26B9FBEFCC4" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cephennomicrus giganteus Jałoszyński &amp; Perkovsky, 2016, sp. n." docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="23" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="BCB1C338E312F1FB0791D2FF7F21CFA8">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="70C6EBC51E1B1CBFF36E1A43E3EA5D94">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="69234BD57EB873A74D6BE4B8A9DF5687">Perkovsky, Evgeny</mods:namePart>
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<mods:date id="1A84E980D552E41E85CBD224424DC919">2016</mods:date>
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<treatment id="03A187A9292FFFE9FF03C26B9FBEFCC4" ID-DOI="http://doi.org/10.5281/zenodo.5681743" ID-GBIF-Taxon="124407844" ID-Zenodo-Dep="5681743" LSID="urn:lsid:plazi:treatment:03A187A9292FFFE9FF03C26B9FBEFCC4" httpUri="http://treatment.plazi.org/id/03A187A9292FFFE9FF03C26B9FBEFCC4" lastPageId="24" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="C3126534292FFFE7FF03C26B9C17F850" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BB736BF292FFFE7FF03C26B9F6EF8B1" blockId="22.[151,570,1862,1922]" box="[151,570,1862,1888]" pageId="22" pageNumber="23">
<heading id="D0FF81D3292FFFE7FF03C26B9F6EF8B1" bold="true" box="[151,570,1862,1888]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B97CEAAD292FFFE7FF03C26B9F6EF8B1" bold="true" box="[151,570,1862,1888]" pageId="22" pageNumber="23">
<taxonomicName id="4C084D3C292FFFE7FF31C26B9CA5F8B1" ID-CoL="SFN7" box="[165,497,1862,1888]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="giganteus" status="sp. nov.">
<emphasis id="B97CEAAD292FFFE7FF31C26B9CA5F8B1" bold="true" box="[165,497,1862,1888]" italics="true" pageId="22" pageNumber="23">Cephennomicrus giganteus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292FFFE7FE6CC26B9F6EF8B1" box="[504,570,1862,1888]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF292FFFE7FF03C2449C17F850" blockId="22.[151,570,1862,1922]" box="[151,323,1897,1922]" pageId="22" pageNumber="23">
(
<figureCitation id="13332A3A292FFFE7FF0BC2449DB3F853" box="[159,231,1897,1922]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="22" pageNumber="23">Figs 5</figureCitation>
,
<figureCitation id="13332A3A292FFFE7FF67C2449C6EF850" box="[243,314,1897,1921]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="22" pageNumber="23">7174</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3126534292FFFE7FF03C29C9E80F83F" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BB736BF292FFFE7FF03C29C9E80F83F" blockId="22.[151,1436,1969,2030]" pageId="22" pageNumber="23">
<materialsCitation id="3B603CE2292FFFE7FF03C29C9E80F83F" ID-GBIF-Occurrence="1300093511" collectionCode="SIZK" location="Europe" municipality="Late Eocene" pageId="22" pageNumber="23" specimenCount="2" stateProvince="Rovno" typeStatus="holotype">
<emphasis id="B97CEAAD292FFFE7FF03C29C9EC3F81B" bold="true" box="[151,919,1969,1994]" pageId="22" pageNumber="23">
<typeStatus id="54B3881D292FFFE7FF03C29C9D86F81B" box="[151,210,1969,1994]" pageId="22" pageNumber="23">Type</typeStatus>
material.
<typeStatus id="54B3881D292FFFE7FED8C29C9CE9F81B" box="[332,445,1969,1994]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="6BD3ACC5292FFFE7FE5EC29C9F09F81B" box="[458,605,1969,1994]" pageId="22" pageNumber="23">Late Eocene</collectingMunicipality>
of
<location id="8ED76064292FFFE7FD10C29C9F8AF81B" LSID="urn:lsid:plazi:treatment:03A187A9292FFFE9FF03C26B9FBEFCC4:8ED76064292FFFE7FD10C29C9F8AF81B" box="[644,734,1969,1994]" municipality="Late Eocene" name="Europe" pageId="22" pageNumber="23" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D292FFFE7FD7FC29C9E6DF81B" box="[747,825,1969,1994]" country="Ukraine" name="Rivne" pageId="22" pageNumber="23">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, irregular prism of amber
<quantity id="4CF09B5A292FFFE7FEDDC2F89CF9F83C" box="[329,429,2005,2030]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="22" pageNumber="23" unit="mm" value="21.5">21.5 mm</quantity>
long (
<figureCitation id="13332A3A292FFFE7FE61C2F89F6CF83F" box="[501,568,2005,2030]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="22" pageNumber="23">Fig. 5</figureCitation>
), collection number K-6011 (
<collectionCode id="ED19AE7A292FFFE7FC10C2FB9E9CF83F" box="[900,968,2006,2030]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="22" pageNumber="23">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF776637292EFFE6FF03C6B89F29FBF6" httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1434,193,896]" targetPageId="23">
<paragraph id="8BB736BF292EFFE6FF03C6B89F29FBF6" blockId="23.[151,1436,917,1063]" pageId="23" pageNumber="24">
<emphasis id="B97CEAAD292EFFE6FF03C6B89C07FC7A" bold="true" box="[151,339,917,940]" pageId="23" pageNumber="24">FIGURES 7174.</emphasis>
Holotype of
<taxonomicName id="4C084D3C292EFFE6FE4CC6BB9FB8FC7D" box="[472,748,918,940]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="giganteus" status="sp. nov.">
<emphasis id="B97CEAAD292EFFE6FE4CC6BB9FB8FC7D" box="[472,748,918,940]" italics="true" pageId="23" pageNumber="24">Cephennomicrus giganteus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292EFFE6FD66C6BA9E7FFC7D" box="[754,811,919,940]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B97CEAAD292EFFE6FD66C6BA9E7FFC7D" bold="true" box="[754,811,919,940]" pageId="23" pageNumber="24">sp. n.</emphasis>
</taxonomicNameLabel>
(K-6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp23, maxillary palpomere IIIII; pg, pygidium; psp, prosternal process; st3 8, abdominal sternite IIIVIII; v3, metaventrite.
</paragraph>
</caption>
<subSubSection id="C3126534292EFFE6FF53C163994BFB5A" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BB736BF292EFFE6FF53C163994BFB5A" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
<emphasis id="B97CEAAD292EFFE6FF53C1639C15FBB6" bold="true" box="[199,321,1102,1127]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
This is an unusually large adult, BL
<quantity id="4CF09B5A292EFFE6FD7AC1639E02FBB7" box="[750,854,1102,1127]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="23" pageNumber="24" unit="mm" value="1.55">1.55 mm</quantity>
, body slender (EI 1.48), densely covered with fine setae; pronotum with two pairs of antebasal pits connected by transverse groove.
</paragraph>
</subSubSection>
<subSubSection id="C3126534292EFFE6FF53C1BB998FF91A" pageId="23" pageNumber="24" type="description">
<paragraph id="8BB736BF292EFFE6FF53C1BB9C79FB03" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
<emphasis id="B97CEAAD292EFFE6FF53C1BB9C0DFB7E" bold="true" box="[199,345,1174,1199]" pageId="23" pageNumber="24">Description.</emphasis>
Body (
<figureCitation id="13332A3A292EFFE6FE3AC1BB9F7BFB7E" box="[430,559,1174,1199]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 7174</figureCitation>
) elongate and strongly convex, light brown with slightly lighter appendages; BL
<quantity id="4CF09B5A292EFFE6FF56C1979C7DFB03" box="[194,297,1210,1235]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="23" pageNumber="24" unit="mm" value="1.55">1.55 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF292EFFE6FF53C1F39CC5FA56" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
Head (
<figureCitation id="13332A3A292EFFE6FE83C1F39CC8FB26" box="[279,412,1246,1271]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 7174</figureCitation>
) broadest at eyes, HL
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, HW about
<quantity id="4CF09B5A292EFFE6FC2FC1F29973FB27" box="[955,1063,1247,1271]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="24" unit="mm" value="0.28">0.28 mm</quantity>
; head structures poorly visible, compound eyes large, slightly bean-shaped and strongly convex, finely faceted. Antennae (
<figureCitation id="13332A3A292EFFE6FB0EC02F984FFACA" box="[1178,1307,1282,1307]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 7174</figureCitation>
) about half BL, AnL about
<quantity id="4CF09B5A292EFFE6FEC4C00B9CEDFAEF" box="[336,441,1318,1343]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="24" unit="mm" value="0.75">0.75 mm</quantity>
, scape and pedicel strongly elongate, antennomeres IIIVIII elongate, IXX distinctly transverse, XI about as long as IXX combined, slightly broader than X, less than twice as long as broad and slightly asymmetrical.
</paragraph>
<paragraph id="8BB736BF292EFFE6FF53C0BE9CFFF98E" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
Pronotum (
<figureCitation id="13332A3A292EFFE6FED0C0BF9C96FA7A" box="[324,450,1426,1451]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 71, 73</figureCitation>
) subrectangular with rounded anterior margin; PL
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, PW
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; pronotal disc broadest near anterior third, sides rounded in anterior half and slightly concave near posterior third; posterior pronotal corners distinct; posterior margin arcuate; base with two pairs of small and shallow but distinct antebasal pits (
<figureCitation id="13332A3A292EFFE6FF5BC0D39C76F9C6" box="[207,290,1534,1559]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Fig. 73</figureCitation>
;
<emphasis id="B97CEAAD292EFFE6FEBAC0D39C0CF9C6" box="[302,344,1534,1559]" italics="true" pageId="23" pageNumber="24">abp</emphasis>
) connected by narrow transverse antebasal groove (
<figureCitation id="13332A3A292EFFE6FC0BC0D39EA6F9C6" box="[927,1010,1534,1559]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Fig. 73</figureCitation>
;
<emphasis id="B97CEAAD292EFFE6FC6AC0D3997CF9C6" box="[1022,1064,1534,1559]" italics="true" pageId="23" pageNumber="24">abg</emphasis>
). Punctures on pronotal disc fine but distinct, separated by spaces subequal to diameters of punctures; setae short, moderately dense, nearly recumbent (
<figureCitation id="13332A3A292EFFE6FE88C36B9CCAF98E" box="[284,414,1606,1631]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 71, 73</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292EFFE6FF53C3479968F976" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
Elytra (
<figureCitation id="13332A3A292EFFE6FE88C3479CCBF952" box="[284,415,1642,1667]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 7174</figureCitation>
) elongate oval, strongly convex, EL 1.00 mm, EW
<quantity id="4CF09B5A292EFFE6FC62C347990AF953" box="[1014,1118,1642,1667]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="23" pageNumber="24" unit="mm" value="0.68">0.68 mm</quantity>
, EI 1.48; punctures slightly larger than those on pronotal disc; setae similar to those on pronotum (
<figureCitation id="13332A3A292EFFE6FC39C3A3997BF976" box="[941,1071,1678,1703]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 71, 73</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292EFFE6FF53C39E998FF91A" blockId="23.[151,1437,1102,2027]" box="[199,1243,1714,1739]" pageId="23" pageNumber="24">
Legs (
<figureCitation id="13332A3A292EFFE6FE98C39F9CD8F91A" box="[268,396,1714,1739]" captionStart="FIGURES 71 74" captionStartId="23.[151,264,917,940]" captionTargetBox="[151,1434,193,896]" captionTargetId="figure@23.[151,1436,193,897]" captionTargetPageId="23" captionText="FIGURES 71 74. Holotype of Cephennomicrus giganteus sp. n. (K- 6011). Habitus in dorsolateral (71, 73) and lateral (72, 74) view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral foveae; bst, basisternal part of prosternum; hy, hypomeron,; hyr, hypomeral ridge; lpc, lateral pronotal carina; mcp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 3, maxillary palpomere II III; pg, pygidium; psp, prosternal process; st 3 8, abdominal sternite III VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262424/files/figure.png" pageId="23" pageNumber="24">Figs 72, 74</figureCitation>
) moderately long, slender, unmodified; all tibiae straight or weakly curved.
</paragraph>
</subSubSection>
<subSubSection id="C3126534292EFFE6FF53C3FB9996F93E" box="[199,1218,1750,1775]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BB736BF292EFFE6FF53C3FB9996F93E" blockId="23.[151,1437,1102,2027]" box="[199,1218,1750,1775]" pageId="23" pageNumber="24">
<emphasis id="B97CEAAD292EFFE6FF53C3FB9C1AF93E" bold="true" box="[199,334,1750,1775]" pageId="23" pageNumber="24">Etymology.</emphasis>
The name
<taxonomicName id="4C084D3C292EFFE6FE5FC3FA9F6CF93F" box="[459,568,1751,1774]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B97CEAAD292EFFE6FE5FC3FA9F6CF93F" box="[459,568,1751,1774]" italics="true" pageId="23" pageNumber="24">giganteus</emphasis>
</taxonomicName>
refers to the exceptionally large body of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C3126534292EFFE6FF53C3D79942F817" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BB736BF292EFFE6FF53C3D79942F817" blockId="23.[151,1437,1102,2027]" pageId="23" pageNumber="24">
<emphasis id="B97CEAAD292EFFE6FF53C3D79C6FF8C2" bold="true" box="[199,315,1786,1811]" pageId="23" pageNumber="24">Remarks.</emphasis>
The cephenniine genus
<taxonomicName id="4C084D3C292EFFE6FDFFC3D79E7DF8C2" box="[619,809,1786,1811]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD292EFFE6FDFFC3D79E7DF8C2" box="[619,809,1786,1811]" italics="true" pageId="23" pageNumber="24">Cephennomicrus</emphasis>
</taxonomicName>
can easily be identified on the basis of characters summarized in
<bibRefCitation id="EF994B4E292EFFE6FED3C2339F7FF8E6" author="Jaloszynski" box="[327,555,1822,1847]" pageId="23" pageNumber="24" refString="Jaloszynski, P. (2011 a) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011" yearSuffix="a">Jałoszyński (2011a)</bibRefCitation>
: the maxillary palpomere IV very short and broad, button-like; the pronotum typically with four antebasal pits and often with a transverse groove; each elytron with two small and asetose basal foveae; the prosternal process narrow, subtriangular and weakly elevated, but distinctly separating procoxae; and the mesoventral intercoxal process moderately broad and not elevated. All these characters can be seen in the studied inclusion, and also the general body form is typical of
<taxonomicName id="4C084D3C292EFFE6FCC5C283995BF816" box="[849,1039,1966,1991]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD292EFFE6FCC5C283995BF816" box="[849,1039,1966,1991]" italics="true" pageId="23" pageNumber="24">Cephennomicrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126534292EFFE9FF53C2FF9FBEFCC4" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BB736BF292EFFE9FF53C2FF9FBEFCC4" blockId="23.[151,1437,1102,2027]" lastBlockId="24.[151,1436,151,789]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<taxonomicName id="4C084D3C292EFFE6FF53C2FF9CD1F83A" ID-CoL="8XKBK" box="[199,389,2002,2027]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD292EFFE6FF53C2FF9CD1F83A" box="[199,389,2002,2027]" italics="true" pageId="23" pageNumber="24">Cephennomicrus</emphasis>
</taxonomicName>
currently comprises 74 extant species distributed in the Afrotropical (
<collectingCountry id="F31F762F292EFFE6FB33C2FF9869F83A" box="[1191,1341,2002,2027]" name="South Africa" pageId="23" pageNumber="24">
<collectingRegion id="49CCF85D292EFFE6FB33C2FF99BEF83A" box="[1191,1258,2002,2027]" country="Tanzania" name="Kusini Unguja" pageId="23" pageNumber="24">South</collectingRegion>
Africa
</collectingCountry>
,
<collectingRegion id="49CCF85D292EFFE6FADEC2FE98C3F83A" box="[1354,1431,2003,2027]" country="Democratic Republic of the Congo" name="Bas-Congo" pageId="23" pageNumber="24">
<collectingCountry id="F31F762F292EFFE6FADEC2FE98C3F83A" box="[1354,1431,2003,2027]" name="Democratic Republic of the Congo" pageId="23" pageNumber="24">Congo</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F31F762F2921FFE9FF03C5BA9DABFF61" box="[151,255,151,176]" name="Tanzania" pageId="24" pageNumber="25">Tanzania</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FE9EC5BA9CC7FF61" box="[266,403,151,176]" name="Madagascar" pageId="24" pageNumber="25">Madagascar</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FE0AC5BA9F5AFF61" box="[414,526,151,176]" name="Mauritius" pageId="24" pageNumber="25">Mauritius</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FD8EC5BA9FC1FF61" box="[538,661,151,176]" name="Seychelles" pageId="24" pageNumber="25">Seychelles</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FD34C5BA9FABFF61" box="[672,767,151,176]" name="Réunion" pageId="24" pageNumber="25">
<collectingRegion id="49CCF85D2921FFE9FD34C5BA9FABFF61" box="[672,767,151,176]" country="France" name="Reunion" pageId="24" pageNumber="25">Reunion</collectingRegion>
</collectingCountry>
Island and Grand
<collectingCountry id="F31F762F2921FFE9FC5BC5B5997DFF61" box="[975,1065,152,176]" name="Comoros" pageId="24" pageNumber="25">Comore</collectingCountry>
Island) and
<collectingRegion id="49CCF85D2921FFE9FB21C5BA9845FF61" box="[1205,1297,151,176]" country="Democratic Republic of the Congo" name="Orientale" pageId="24" pageNumber="25">Oriental</collectingRegion>
regions (
<collectingCountry id="F31F762F2921FFE9FAEEC5BA9DB4FF04" name="Sri Lanka" pageId="24" pageNumber="25">Sri Lanka</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FF65C5919C03FF04" box="[241,343,188,213]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FEFDC5919CEAFF04" box="[361,446,188,213]" name="Taiwan" pageId="24" pageNumber="25">Taiwan</collectingCountry>
, subtropical
<collectingCountry id="F31F762F2921FFE9FDCCC5919FCFFF04" box="[600,667,188,213]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
and
<collectingCountry id="F31F762F2921FFE9FD4AC5909E75FF05" box="[734,801,189,212]" name="Japan" pageId="24" pageNumber="25">Japan</collectingCountry>
,
<collectingRegion id="49CCF85D2921FFE9FCA6C5909E3DFF05" box="[818,873,189,212]" country="Tanzania" name="Mjini Magharibi" pageId="24" pageNumber="25">West</collectingRegion>
<collectingCountry id="F31F762F2921FFE9FCE2C5919E89FF04" box="[886,989,188,213]" name="Malaysia" pageId="24" pageNumber="25">Malaysia</collectingCountry>
and
<collectingRegion id="49CCF85D2921FFE9FB8BC59199D5FF05" box="[1055,1153,188,212]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="25">Sumatra</collectingRegion>
), extending through the
<collectingCountry id="F31F762F2921FFE9FF03C5F29C41FF29" box="[151,277,223,248]" name="Philippines" pageId="24" pageNumber="25">Philippines</collectingCountry>
and
<collectingCountry id="F31F762F2921FFE9FEC4C5CD9F7AFF29" box="[336,558,223,248]" name="Papua New Guinea" pageId="24" pageNumber="25">
<collectingRegion id="49CCF85D2921FFE9FEC4C5CD9CC0FF26" box="[336,404,224,247]" country="Indonesia" name="Papua Barat" pageId="24" pageNumber="25">Papua</collectingRegion>
New Guinea
</collectingCountry>
to
<collectingCountry id="F31F762F2921FFE9FDC2C5F29FE8FF29" box="[598,700,223,248]" name="Australia" pageId="24" pageNumber="25">Australia</collectingCountry>
(including Lord Howe Island),
<collectingCountry id="F31F762F2921FFE9FBB9C5CD99B4FF29" box="[1069,1248,223,248]" name="New Caledonia" pageId="24" pageNumber="25">New Caledonia</collectingCountry>
,
<collectingCountry id="F31F762F2921FFE9FB7AC5F29841FF29" box="[1262,1301,223,248]" name="Fiji" pageId="24" pageNumber="25">Fiji</collectingCountry>
and
<collectingCountry id="F31F762F2921FFE9FAC4C5CD98C8FF29" box="[1360,1436,224,248]" name="Samoa" pageId="24" pageNumber="25">Samoa</collectingCountry>
(
<bibRefCitation id="EF994B4E2921FFE9FF0AC4299C3AFECD" author="Jaloszynski" box="[158,366,260,285]" pageId="24" pageNumber="25" refString="Jaloszynski, P. (2011 a) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011" yearSuffix="a">Jałoszyński 2011a</bibRefCitation>
,
<bibRefCitation id="EF994B4E2921FFE9FEEEC4299C97FECC" author="Jaloszynski" box="[378,451,260,285]" pageId="24" pageNumber="25" refString="Jaloszynski, P. (2015 e) Four new species of Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa, 3911 (2), 273 - 279. http: // dx. doi. org / 10.11646 / zootaxa. 3911.2.8" type="journal article" year="2015" yearSuffix="e">2015e</bibRefCitation>
). Many undescribed species were seen by the first author from various parts of East and
<collectingRegion id="49CCF85D2921FFE9FF03C40A9D8CFE91" box="[151,216,295,320]" country="Tanzania" name="Kusini Unguja" pageId="24" pageNumber="25">South</collectingRegion>
Asia, especially from the Sunda Islands and several from the Himalayas; also in
<collectingCountry id="F31F762F2921FFE9FBE4C40A9983FE91" box="[1136,1239,295,320]" name="Australia" pageId="24" pageNumber="25">Australia</collectingCountry>
<taxonomicName id="4C084D3C2921FFE9FB4AC40A98C8FE91" ID-CoL="8XKBK" box="[1246,1436,295,320]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2921FFE9FB4AC40A98C8FE91" box="[1246,1436,295,320]" italics="true" pageId="24" pageNumber="25">Cephennomicrus</emphasis>
</taxonomicName>
is more speciose than recorded so far, and the poorly studied fauna of
<collectingCountry id="F31F762F2921FFE9FC62C4619929FEB4" box="[1014,1149,332,357]" name="Madagascar" pageId="24" pageNumber="25">Madagascar</collectingCountry>
is exceptionally rich in undescribed species of this interesting genus (Jałoszyński, unpublished obs.). A possibly broader distribution in other parts of Africa, Melanesia,
<collectingCountry id="F31F762F2921FFE9FDB2C4B99FF0FE7C" box="[550,676,404,429]" name="Micronesia" pageId="24" pageNumber="25">Micronesia</collectingCountry>
and Polynesia was also suggested (
<bibRefCitation id="EF994B4E2921FFE9FBC0C4B99878FE7D" author="Jaloszynski" box="[1108,1324,404,429]" pageId="24" pageNumber="25" refString="Jaloszynski, P. (2011 a) Cephenniini with prothoracic glands and internal ' cavities': new taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470 - 496. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2011.00574. x" type="journal article" year="2011" yearSuffix="a">Jałoszyński 2011a</bibRefCitation>
). This is, therefore, a large and taxonomically challenging genus, comprising many species so similar one to another that in some cases only the male genitalia can be used for identification (e.g.,
<bibRefCitation id="EF994B4E2921FFE9FC25C4F199D7FE25" author="Jaloszynski" box="[945,1155,476,501]" pageId="24" pageNumber="25" refString="Jaloszynski, P. (2009 a) Cephennomicrus Reitter (Coleoptera, Staphylinidae, Scydmaeninae) of Japan and Taiwan: taxonomic notes, ten new species and comparative morphology of nomurai and taiwanensis species groups. Zootaxa, 2145, 1 - 35." type="journal article" year="2009" yearSuffix="a">Jałoszyński 2009a</bibRefCitation>
). However, in contrast to other small-bodied
<taxonomicName id="4C084D3C2921FFE9FEEDC4D29F51FDC9" ID-CoL="8VR7K" box="[377,517,511,536]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="tribe" tribe="Cephenniini">Cephenniini</taxonomicName>
,
<taxonomicName id="4C084D3C2921FFE9FD80C4D29F86FDC9" ID-CoL="8XKBK" box="[532,722,511,536]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2921FFE9FD80C4D29F86FDC9" box="[532,722,511,536]" italics="true" pageId="24" pageNumber="25">Cephennomicrus</emphasis>
</taxonomicName>
shows a relatively large diversity of body shapes, sculpture, vestiture of setae and male secondary sexual characters. This allows for defining species on the basis of external characters, especially when they show conspicuous features. As most
<taxonomicName id="4C084D3C2921FFE9FC36C76A9934FDB1" ID-CoL="8XKBK" box="[930,1120,583,608]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2921FFE9FC36C76A9934FDB1" box="[930,1120,583,608]" italics="true" pageId="24" pageNumber="25">Cephennomicrus</emphasis>
</taxonomicName>
adults do not reach
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in length, a large body over
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is remarkable enough to facilitate species identification.
<taxonomicName id="4C084D3C2921FFE9FB4AC7419C50FD79" ID-CoL="SFN7" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B97CEAAD2921FFE9FB4AC7419C50FD79" italics="true" pageId="24" pageNumber="25">Cephennomicrus giganteus</emphasis>
</taxonomicName>
is the largest of all hitherto described species of this genus. The 74 extant species have the body length ranging from
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; 30 of them do not reach
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, and only four reach or exceed
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. None of them is as large as
<taxonomicName id="4C084D3C2921FFE9FED4C7F59C87FD21" ID-CoL="SFN7" box="[320,467,728,752]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B97CEAAD2921FFE9FED4C7F59C87FD21" box="[320,467,728,752]" italics="true" pageId="24" pageNumber="25">C. giganteus</emphasis>
</taxonomicName>
, which measures
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, and this feature alone can be used to distinguish this remarkable extinct species from its extant congeners.
</paragraph>
</subSubSection>
</treatment>
</document>