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<document id="5E0B9AB9680A0EB49019B41E6EC2C3CF" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92929FFE0FF03C7AF984AF971" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Vertheia Jałoszyński &amp; Perkovsky, 2016, gen. n." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="17" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="510478BECAE48D68CE4A47C81A5E2E94">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="83A85A8F0E54F61BF62F759ED422D972">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="B3C0CF74C7B7AF8B5716B97AE1D21680">Perkovsky, Evgeny</mods:namePart>
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<mods:title id="2B2B33740F3EB6EEC244DCC933C788F4">Zootaxa</mods:title>
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<mods:date id="9AA7C2C55B861007D5FED1E4C4708C8B">2016</mods:date>
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<mods:number id="3E98657BF09E28F162CBEEEA21DC4466">4157</mods:number>
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<mods:identifier id="EE716C86F410B854F365D0F7D4CB838F" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<treatment id="03A187A92929FFE0FF03C7AF984AF971" ID-DOI="http://doi.org/10.5281/zenodo.5681735" ID-GBIF-Taxon="124407822" ID-Zenodo-Dep="5681735" LSID="urn:lsid:plazi:treatment:03A187A92929FFE0FF03C7AF984AF971" httpUri="http://treatment.plazi.org/id/03A187A92929FFE0FF03C7AF984AF971" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C31265342929FFE1FF03C7AF9C30FD4D" box="[151,356,642,668]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BB736BF2929FFE1FF03C7AF9C30FD4D" blockId="16.[151,356,642,668]" box="[151,356,642,668]" pageId="16" pageNumber="17">
<heading id="D0FF81D32929FFE1FF03C7AF9C30FD4D" bold="true" box="[151,356,642,668]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B97CEAAD2929FFE1FF03C7AF9C30FD4D" bold="true" box="[151,356,642,668]" pageId="16" pageNumber="17">
†
<taxonomicName id="4C084D3C2929FFE1FF31C7AF9C53FD4D" box="[165,263,642,668]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2929FFE1FF31C7AF9C53FD4D" bold="true" box="[165,263,642,668]" italics="true" pageId="16" pageNumber="17">Vertheia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62929FFE1FE86C7AF9C30FD4D" box="[274,356,642,668]" pageId="16" pageNumber="17" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31265342929FFE1FF03C7E79F94FD0E" box="[151,704,713,735]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BB736BF2929FFE1FF03C7E79F94FD0E" blockId="16.[151,704,713,735]" box="[151,704,713,735]" pageId="16" pageNumber="17">
Type species:
<taxonomicName id="4C084D3C2929FFE1FEB2C7E49F50FD0E" box="[294,516,713,735]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quadrisetosa">
<emphasis id="B97CEAAD2929FFE1FEB2C7E49F50FD0E" box="[294,516,713,735]" italics="true" pageId="16" pageNumber="17">Vertheia quadrisetosa</emphasis>
</taxonomicName>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="C31265342929FFE1FF03C6219C72FC41" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BB736BF2929FFE1FF03C6219C72FC41" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
<emphasis id="B97CEAAD2929FFE1FF03C6219C45FCF4" bold="true" box="[151,273,780,805]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
A genus of
<taxonomicName id="4C084D3C2929FFE1FE31C6219F58FCF4" box="[421,524,780,805]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="tribe" tribe="Eutheiini">Eutheiini</taxonomicName>
with a pair of pits on posterior portion of frons, each pit connected to oblique groove extending posterolaterally; submentum long and subhexagonal; procoxae contiguous; mesocoxae broadly separated by flat and subtrapezoidal mesoventral process; elytra truncated at apices; basal elytral foveae absent or rudimentary.
</paragraph>
</subSubSection>
<subSubSection id="C31265342929FFE0FF53C6B19E61FE59" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BB736BF2929FFE1FF53C6B19E95FC06" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
<emphasis id="B97CEAAD2929FFE1FF53C6B19C0DFC64" bold="true" box="[199,345,924,949]" pageId="16" pageNumber="17">Description.</emphasis>
Body (
<figureCitation id="13332A3A2929FFE1FE3FC6B19F78FC64" box="[427,556,924,949]" captionStart-0="FIGURES 49 – 55" captionStart-1="FIGURES 56 – 60" captionStart-2="FIGURES 61 – 64" captionStartId-0="18.[151,264,1720,1743]" captionStartId-1="19.[151,264,1962,1985]" captionStartId-2="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,443,1689]" captionTargetBox-1="[151,1436,628,1941]" captionTargetBox-2="[160,1393,591,1818]" captionTargetId-0="figure@18.[151,1436,443,1699]" captionTargetId-1="figure@19.[151,1436,628,1941]" captionTargetId-2="figure@21.[151,1436,591,1824]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="21" captionText-0="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." captionText-1="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-2="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri-0="https://zenodo.org/record/262420/files/figure.png" httpUri-1="https://zenodo.org/record/262421/files/figure.png" httpUri-2="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Figs 49–64</figureCitation>
) elongate and flattened, with distinctly marked constriction between head and pronotum but weakly so between pronotum and elytra, BL below
<quantity id="4CF09B5A2929FFE1FCEDC6ED9EE9FC06" box="[889,957,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C6C99EA9FB29" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Head capsule (
<figureCitation id="13332A3A2929FFE1FEE7C6C99CADFC2C" box="[371,505,996,1021]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD93C6C99F1AFC2C" box="[519,590,996,1021]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–63</figureCitation>
) with anterior part (in front of occipital constriction) subtrapezoidal and transverse; eyes located posteriorly, large, strongly convex and moderately coarsely faceted; tempora (
<figureCitation id="13332A3A2929FFE1FA81C12A9838FBF1" box="[1301,1388,1031,1056]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FAECC12A98C3FBF1" box="[1400,1431,1031,1056]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FF03C1009DE7FB95" box="[151,179,1069,1092]" italics="true" pageId="16" pageNumber="17">tm</emphasis>
) barely discernible, very short; occipital constriction (
<figureCitation id="13332A3A2929FFE1FCB5C1019E2EFB94" box="[801,890,1068,1093]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC1CC1019EF3FB94" box="[904,935,1068,1093]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC20C1009E8FFB95" box="[948,987,1069,1092]" italics="true" pageId="16" pageNumber="17">occ</emphasis>
) as broad as frons between eyes; frons (
<figureCitation id="13332A3A2929FFE1FF0BC1629DA2FBB9" box="[159,246,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE97C1629C76FBB9" box="[259,290,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FEBAC17D9C15FBB9" box="[302,321,1104,1128]" italics="true" pageId="16" pageNumber="17">fr</emphasis>
) subtrapezoidal, posteriorly confluent with strongly transverse, subrectangular vertex (
<figureCitation id="13332A3A2929FFE1FA80C1629838FBB9" box="[1300,1388,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FAECC16298C3FBB9" box="[1400,1431,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FF03C1589DFFFB5D" box="[151,171,1141,1164]" italics="true" pageId="16" pageNumber="17">vt</emphasis>
); frons posteriorly with pair of small but distinct pits (
<figureCitation id="13332A3A2929FFE1FCB1C1599E2AFB5C" box="[805,894,1140,1165]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC19C1599EF8FB5C" box="[909,940,1140,1165]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC2EC1599E8FFB5D" box="[954,987,1140,1164]" italics="true" pageId="16" pageNumber="17">frp</emphasis>
), each posteriorly connected to groove extending obliquely to posterolateral margin of vertex; 'neck' region (
<figureCitation id="13332A3A2929FFE1FC27C1BA995FFB61" box="[947,1035,1175,1200]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FB8DC1BA996CFB61" box="[1049,1080,1175,1200]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FBD2C1B4990BFB61" box="[1094,1119,1177,1200]" italics="true" pageId="16" pageNumber="17">nr</emphasis>
) as broad as frons between eyes. Submentum (
<figureCitation id="13332A3A2929FFE1FEE3C1919C9BFB04" box="[375,463,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 63</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FE74C1909F59FB05" box="[480,525,1213,1236]" italics="true" pageId="16" pageNumber="17">smn</emphasis>
) subhexagonal, about as broad as long; posterior tentorial pits (
<figureCitation id="13332A3A2929FFE1FA92C191980AFB04" box="[1286,1374,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FAFBC19098C7FB05" box="[1391,1427,1213,1236]" italics="true" pageId="16" pageNumber="17">ptp</emphasis>
) distinct, located in transverse groove demarcating anteriorly the 'neck' region.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0299F03FA91" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Antennae (
<figureCitation id="13332A3A2929FFE1FED2C0299C99FACC" box="[326,461,1284,1309]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE48C0299F71FACC" box="[476,549,1284,1309]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDA1C0299F28FACC" box="[565,636,1284,1309]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
) moderately long and slender, with enlarged scape and pedicel and gradually but weakly thickened distally.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0609942FA25" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Pronotum (
<figureCitation id="13332A3A2929FFE1FEDCC0619F72FAB4" box="[328,550,1356,1381]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 52–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDA1C0619F29FAB4" box="[565,637,1356,1381]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–57</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD19C0619E56FAB4" box="[653,770,1356,1381]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
) in dorsal view subrectangular with distinctly or slightly concave anterior margin, feebly marked and obtuse-angled anterior pronotal corners, rounded sides, distinct obtuse-angled posterior pronotal corners and arcuate posterior margin, slightly flattened in front of scutellum. Sides with distinct lateral pronotal carinae or edges (
<figureCitation id="13332A3A2929FFE1FD7EC09A9E6AFA01" box="[746,830,1463,1488]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 64</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCDFC09A9E39FA01" box="[843,877,1463,1488]" italics="true" pageId="16" pageNumber="17">lpc</emphasis>
); base with shallow lateral antebasal impressions (
<figureCitation id="13332A3A2929FFE1FF0BC0F19DA1FA24" box="[159,245,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE95C0F19C74FA24" box="[257,288,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FEB8C0F19C04FA24" box="[300,336,1500,1525]" italics="true" pageId="16" pageNumber="17">abi</emphasis>
) and long, faint transverse antebasal groove (
<figureCitation id="13332A3A2929FFE1FCDAC0F19EF1FA24" box="[846,933,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC25C0F19E84FA24" box="[945,976,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC48C0F19952FA24" box="[988,1030,1500,1525]" italics="true" pageId="16" pageNumber="17">abg</emphasis>
).
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0D29FEDF976" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Basisternal part of prosternum (
<figureCitation id="13332A3A2929FFE1FDBDC0D29FD4F9C9" box="[553,640,1535,1560]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD18C0D29FFFF9C9" box="[652,683,1535,1560]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">63</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FD23C0D29F83F9C9" box="[695,727,1535,1560]" italics="true" pageId="16" pageNumber="17">bst</emphasis>
) nearly as long as coxal part, laterally demarcated by complete notosternal sutures (
<figureCitation id="13332A3A2929FFE1FEE8C3099C85F9EC" box="[380,465,1572,1597]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FE4AC3089F56F9ED" box="[478,514,1573,1596]" italics="true" pageId="16" pageNumber="17">nss</emphasis>
); procoxae contiguous, indicating that prosternal process is either absent or very narrow and not elevated. Prothoracic hypomeron (
<figureCitation id="13332A3A2929FFE1FD45C36A9E71F9B1" box="[721,805,1607,1632]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCA7C36A9E19F9B1" box="[819,845,1607,1632]" italics="true" pageId="16" pageNumber="17">hy</emphasis>
) flat and with distinctly demarcated inner (adcoxal) part by complete and recurved hypomeral ridge (
<figureCitation id="13332A3A2929FFE1FD72C3419E6BF954" box="[742,831,1644,1669]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCC5C3419E22F954" box="[849,886,1644,1669]" italics="true" pageId="16" pageNumber="17">hyr</emphasis>
). Shape of inner (adcoxal) hypomeral margin indicates that procoxal sockets are broadly open.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C3999CE0F921" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Mesoscutellum (
<figureCitation id="13332A3A2929FFE1FE16C3999F51F91C" box="[386,517,1716,1741]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD86C3999F0EF91C" box="[530,602,1716,1741]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
) well-visible between elytral bases, broadly subtriangular, with indistinct median basal impression.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C3D1982DF8C5" blockId="16.[151,1437,780,2029]" box="[199,1401,1788,1813]" pageId="16" pageNumber="17">
Mesoventrite (
<figureCitation id="13332A3A2929FFE1FEFEC3D19CEFF8C4" box="[362,443,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
) with broadly subtrapezoidal and flat mesoventral intercoxal process (
<figureCitation id="13332A3A2929FFE1FB44C3D19877F8C4" box="[1232,1315,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FABBC3D0983DF8C5" box="[1327,1385,1789,1812]" italics="true" pageId="16" pageNumber="17">msvp</emphasis>
).
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C23299B7F851" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Metaventrite (
<figureCitation id="13332A3A2929FFE1FEF2C2329CEBF8E9" box="[358,447,1823,1848]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE58C2329CBFF8E9" box="[460,491,1823,1848]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<date id="FFB6107F2929FFE1FE6CC20D9F46F8E9" box="[504,530,1824,1848]" pageId="16" pageNumber="17">
<emphasis id="B97CEAAD2929FFE1FE6CC20D9F46F8E9" box="[504,530,1824,1848]" italics="true" pageId="16" pageNumber="17">v3</emphasis>
</date>
) subquadrate with weakly rounded sides, anteriorly at middle demarcated from mesoventrite; posteriorly strongly concave laterally in admetacoxal regions and medially forming very short and broad metaventral intercoxal process (
<figureCitation id="13332A3A2929FFE1FDD7C24A9FCDF851" box="[579,665,1895,1920]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD31C24A9F90F851" box="[677,708,1895,1920]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FD44C2449E52F851" box="[720,774,1897,1920]" italics="true" pageId="16" pageNumber="17">mtvp</emphasis>
) with distinctly concave posterior margin.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C2A19C50F83C" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Elytra (
<figureCitation id="13332A3A2929FFE1FE89C2A19CA3F874" box="[285,503,1932,1957]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 53–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD92C2A19F1AF874" box="[518,590,1932,1957]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDC9C2A19F84F874" box="[605,720,1932,1957]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
) elongate and nearly oval, except for truncated apices; elytral base with foveae absent or rudimentary (indiscernible in studied specimens, in which the elytral base is wellexposed).
</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C5BA9FE9FF61" blockId="17.[151,1437,151,1544]" box="[199,701,151,176]" pageId="17" pageNumber="18">Hind wings not visible in studied specimens.</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C5909FADFECC" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
Legs (
<figureCitation id="13332A3A2928FFE0FE99C5919C30FF04" box="[269,356,188,213]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FEE5C5919CEDFF04" box="[369,441,188,213]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
) moderately long, slender; pro- and mesocoxae suboval, metacoxae strongly transverse; all trochanters small and subtriangular; femora weakly clavate; tibiae slender, straight or nearly straight; tarsi long and slender, all tarsomeres elongate, slightly flattened.
</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C40A9E61FE59" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
Abdomen (
<figureCitation id="13332A3A2928FFE0FED7C40A9CCEFE91" box="[323,410,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FE31C40A9CB8FE91" box="[421,492,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">62–64</figureCitation>
) protruding from under truncated elytra, six sternites visible (
<figureCitation id="13332A3A2928FFE0FB0EC40A99A5FE91" box="[1178,1265,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FB6FC40A9810FE91" box="[1275,1348,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
;
<emphasis id="B97CEAAD2928FFE0FADBC40598DFFE91" box="[1359,1419,296,320]" italics="true" pageId="17" pageNumber="18">st3–8</emphasis>
), sternite III only slightly longer than IV, sternites VII and VIII longest, suture between them partly indistinct. Pygidium (
<figureCitation id="13332A3A2928FFE0FE87C4429C32FE59" box="[275,358,367,392]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">Fig. 62</figureCitation>
;
<emphasis id="B97CEAAD2928FFE0FEE6C45C9CDAFE59" box="[370,398,369,392]" italics="true" pageId="17" pageNumber="18">pg</emphasis>
) visible in dorsal view, subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C31265342928FFE0FF53C4B99E9FFE01" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BB736BF2928FFE0FF53C4B99E9FFE01" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FF53C4B99F79FE7C" bold="true" box="[199,557,404,429]" pageId="17" pageNumber="18">Distribution and composition.</emphasis>
<taxonomicName id="4C084D3C2928FFE0FDA1C4B99FC4FE7C" box="[565,656,404,429]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FDA1C4B99FC4FE7C" box="[565,656,404,429]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
is represented by one nominal and one undescribed species from the Eocene of Europe (
<figureCitation id="13332A3A2928FFE0FEFBC49A9C96FE01" box="[367,450,439,464]" captionStart="FIGURES 35 – 38" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[181,1396,193,1909]" captionTargetId="figure@10.[181,1396,193,1910]" captionTargetPageId="10" captionText="FIGURES 35 – 38. Location of the Rovno Region (35 − 36) and the Klesov, Dubrovitsa and Vyrka amber mines (37) in the present-day Europe; location of the Rovno Region during Eocene (38)." httpUri="https://zenodo.org/record/262416/files/figure.png" pageId="17" pageNumber="18">Fig. 38</figureCitation>
), area currently within north-western
<collectingCountry id="F31F762F2928FFE0FCFDC49A9E93FE01" box="[873,967,439,464]" name="Ukraine" pageId="17" pageNumber="18">Ukraine</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31265342928FFE0FF53C4F19F87FDC9" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BB736BF2928FFE0FF53C4F19F87FDC9" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FF53C4F19C1AFE24" bold="true" box="[199,334,476,501]" pageId="17" pageNumber="18">Etymology.</emphasis>
The name
<emphasis id="B97CEAAD2928FFE0FE5DC4F09F77FE24" box="[457,547,476,501]" italics="true" pageId="17" pageNumber="18">Vert heia</emphasis>
is a result of combining
<taxonomicName id="4C084D3C2928FFE0FCADC4F19ECFFE24" box="[825,923,476,501]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FCADC4F19ECFFE24" box="[825,923,476,501]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D3C2928FFE0FC44C4F19973FE24" box="[976,1063,476,501]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC44C4F19973FE24" box="[976,1063,476,501]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
, to underline the similarity of the fossil taxon to both these genera. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C31265342928FFE0FF53C709984AF971" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BB736BF2928FFE0FF53C7099E64FAA9" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FF53C7099C6FFDEC" bold="true" box="[199,315,548,573]" pageId="17" pageNumber="18">Remarks.</emphasis>
The new genus is placed in the tribe
<taxonomicName id="4C084D3C2928FFE0FD65C7099E03FDEC" box="[753,855,548,573]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Eutheiini">Eutheiini</taxonomicName>
on the basis of the following characters: the body weakly constricted between the pronotum and elytra; the head with very short tempora and the occipital constriction as broad as vertex; the maxillary palpomere IV much smaller than III and not longer than broad; the pronotum with sharp lateral edges; the notosternal sutures present and complete; and the metacoxae broadly separated by short metaventral intercoxal process. The specimens found in
<collectingRegion id="49CCF85D2928FFE0FB83C7989936FD1D" box="[1047,1122,693,716]" country="Ukraine" name="Rivne" pageId="17" pageNumber="18">Rovno</collectingRegion>
amber have the maxillary palpomere IV clearly visible, which excludes a placement in the extant genus
<taxonomicName id="4C084D3C2928FFE0FBB1C7FA983CFD21" authority="Franz, 1986a" authorityName="Franz" authorityYear="1986" box="[1061,1384,727,752]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FBB1C7FA999DFD21" box="[1061,1225,727,752]" italics="true" pageId="17" pageNumber="18">Paraneseuthia</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FB47C7F5983CFD21" author="Franz" box="[1235,1384,727,752]" pageId="17" pageNumber="18" refString="Franz, H. (1986 a) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna der Fiji-Inseln. Entomologische Blatter, 82, 147 - 178." type="journal article" year="1986" yearSuffix="a">Franz, 1986a</bibRefCitation>
</taxonomicName>
that lacks the palpomere IV (illustrated in
<bibRefCitation id="EF994B4E2928FFE0FDD2C7D19E49FCC5" author="Jaloszynski" box="[582,797,764,789]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2010) Paraneseuthia Franz, 1986 of Australo-Oriental region, with notes on Palearctic species (Coleoptera, Staphylinidae, Scydmaeninae). Annales Zoologici, 60, 181 - 193. http: // dx. doi. org / 10.3161 / 000345410 X 516830" type="journal article" year="2010">Jałoszyński (2010)</bibRefCitation>
. All remaining genera of
<taxonomicName id="4C084D3C2928FFE0FBD5C7D199FCFCC4" box="[1089,1192,764,789]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Eutheiini">Eutheiini</taxonomicName>
have four-segmented maxillary palps, but
<taxonomicName id="4C084D3C2928FFE0FEEFC6329F83FCE9" authority="Jaloszynski, 2003" authorityName="Jaloszynski" authorityYear="2003" box="[379,727,799,824]" class="Insecta" family="Staphylinidae" genus="Paeneutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FEEFC6329F53FCE9" box="[379,519,799,824]" italics="true" pageId="17" pageNumber="18">Paeneutheia</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FD99C6329F83FCE9" author="Jaloszynski" box="[525,727,799,824]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2003) Description of Paeneutheia pallida gen. et sp. nov. from Borneo, with notes on allied genera of the tribe Eutheiini (Coleoptera, Scydmaenidae). Bulletin of the National Science Museum, Tokyo, 29, 213 - 223." type="journal article" year="2003">Jałoszyński, 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C084D3C2928FFE0FD75C6329EBAFCE9" authority="Muller, 1925" authorityName="Muller" authorityYear="1925" box="[737,1006,799,824]" class="Insecta" family="Staphylinidae" genus="Euthiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FD75C6329E03FCE9" box="[737,855,799,824]" italics="true" pageId="17" pageNumber="18">Euthiopsis</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FCCAC6329EBAFCE9" author="Muller" box="[862,1006,799,824]" pageId="17" pageNumber="18" refString="Muller, G. (1925) Nota su due specie del genere Euthia Steph. Studi Entomologici, 1, 17." type="journal article" year="1925">Müller, 1925</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C084D3C2928FFE0FBB7C632986CFCE9" authority="Reitter, 1881" authorityName="Reitter" authorityYear="1881" box="[1059,1336,799,824]" class="Insecta" family="Staphylinidae" genus="Euthiconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FBB7C63299F5FCE9" box="[1059,1185,799,824]" italics="true" pageId="17" pageNumber="18">Euthiconus</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FB3CC632986CFCE9" author="Reitter" box="[1192,1336,799,824]" pageId="17" pageNumber="18" refString="Reitter, E. (1881) Bestimmungs-Tabellen der europaischen Coleopteren. V. Enthaltend die Familien: Paussidae, Clavigeridae, Pselaphidae und Scydmaenidae. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 31, 443 - 593." type="journal article" year="1881">Reitter, 1881</bibRefCitation>
</taxonomicName>
have the palpomere IV very short, button-like (illustrated in
<bibRefCitation id="EF994B4E2928FFE0FD41C6699E96FC8C" author="Jaloszynski" box="[725,962,836,861]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2014 b) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. http: // dx. doi. org / 10.1111 / syen. 12044" type="journal article" year="2014" yearSuffix="b">Jałoszyński (2014b))</bibRefCitation>
, whereas that found in the studied fossil is distinctly subconical. Only the extant genera
<taxonomicName id="4C084D3C2928FFE0FD31C64A9EFEFC51" authority="Stephens,1830" authorityName="Stephens" authorityYear="1830" box="[677,938,871,896]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FD31C64A9FA8FC51" box="[677,764,871,896]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FC92C64A9EFEFC51" author="Stephens" box="[774,938,871,896]" pageId="17" pageNumber="18" refString="Stephens, J. F. (1830) Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. 3. Baldwin &amp; Cradock, London, 374 pp." type="book" year="1830">Stephens,1830</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C084D3C2928FFE0FC73C64A99B6FC51" authority="Casey, 1897" authorityName="Casey" authorityYear="1897" box="[999,1250,871,896]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC73C64A991DFC51" box="[999,1097,871,896]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FBC7C64599B6FC51" author="Casey" box="[1107,1250,872,896]" pageId="17" pageNumber="18" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Science, 9, 285 - 684. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" type="journal article" year="1897">Casey, 1897</bibRefCitation>
</taxonomicName>
, and the extinct Early Cretaceous
<taxonomicName id="4C084D3C2928FFE0FEF7C6A19E4BFC74" authority="Jaloszynski &amp; Peris, 2016" authorityName="Jaloszynski &amp; Peris" authorityYear="2016" box="[355,799,908,933]" class="Insecta" family="Staphylinidae" genus="Archeutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FEF7C6A19CBFFC74" box="[355,491,908,933]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
<bibRefCitation id="EF994B4E2928FFE0FE60C6A19E4BFC74" author="Jaloszynski" box="[500,799,908,933]" pageId="17" pageNumber="18" refString="Jaloszynski, P. &amp; Peris, D. (2016) Cretaceous amber inclusions of Spain and Myanmar demonstrate early diversification and wide dispersal of Cephenniitae (Coleoptera: Staphylinidae: Scydmaeninae). Cretaceous Research, 57, 190 - 198. http: // dx. doi. org / 10.1016 / j. cretres. 2015.09.002" type="journal article" year="2016">Jałoszyński &amp; Peris, 2016</bibRefCitation>
</taxonomicName>
have the maxillary palpomere IV subconical. The new genus is very similar to all these three taxa in the general body form, the truncate elytral apices and the procoxae not separated by prosternal process.
<taxonomicName id="4C084D3C2928FFE0FDB8C6F99FD3FC3C" box="[556,647,980,1005]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FDB8C6F99FD3FC3C" box="[556,647,980,1005]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
has the pair of distinct pits on the posterior portion of the frons, with a groove extending from each pit posterolaterally; previously this character was thought to be a hallmark of
<taxonomicName id="4C084D3C2928FFE0FAAEC6DA98C8FBC1" box="[1338,1436,1015,1040]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FAAEC6DA98C8FBC1" box="[1338,1436,1015,1040]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
(illustrated in
<bibRefCitation id="EF994B4E2928FFE0FEA8C1319FFCFBE4" author="Jaloszynski" box="[316,680,1052,1077]" pageId="17" pageNumber="18" refString="Jaloszynski, P. &amp; Hoshina, H. (2005) Revision of Palearctic species of Veraphis Casey (Coleoptera, Scydmaenidae). Entomological Review of Japan, 60, 101 - 136." type="journal article" year="2005">Jałoszyński &amp; Hoshina (2005))</bibRefCitation>
, although some species of
<taxonomicName id="4C084D3C2928FFE0FC7CC131996BFBE4" box="[1000,1087,1052,1077]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC7CC131996BFBE4" box="[1000,1087,1052,1077]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
have traces of similar pits or grooves, which are rather faint compared to those known in
<taxonomicName id="4C084D3C2928FFE0FCD3C1129EFDFB89" box="[839,937,1087,1112]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FCD3C1129EFDFB89" box="[839,937,1087,1112]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="EF994B4E2928FFE0FC67C1129999FB89" author="Jaloszynski" box="[1011,1229,1087,1112]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2008 a) Eutheia puetzi n. sp. from China (Coleoptera, Scydmaenidae). Genus, 19 (1), 33 - 36." type="journal article" year="2008" yearSuffix="a">Jałoszyński (2008a</bibRefCitation>
,
<bibRefCitation id="EF994B4E2928FFE0FB4DC11299BDFB89" author="Jaloszynski" box="[1241,1257,1087,1112]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2008 b) Revision of Eutheia Stephens of Taiwan (Coleoptera, Scydmaenidae). Genus, 19 (2), 177 - 190." type="journal article" year="2008" yearSuffix="b">b</bibRefCitation>
)). Interestingly, although this character makes
<taxonomicName id="4C084D3C2928FFE0FE65C1499F18FBAC" box="[497,588,1124,1149]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FE65C1499F18FBAC" box="[497,588,1124,1149]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
very similar to
<taxonomicName id="4C084D3C2928FFE0FC92C1499E3CFBAC" box="[774,872,1124,1149]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC92C1499E3CFBAC" box="[774,872,1124,1149]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
in dorsal view, the broad, flat and subtrapezoidal mesoventral intercoxal process in the Eocene taxon is not even remotely similar to the structure known in
<taxonomicName id="4C084D3C2928FFE0FAA7C1AA98C1FB71" box="[1331,1429,1159,1184]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FAA7C1AA98C1FB71" box="[1331,1429,1159,1184]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
. The latter genus has the mesoventral process narrowly carinate and elevated (illustrated in
<bibRefCitation id="EF994B4E2928FFE0FB3DC18198CCFB14" author="Jaloszynski" box="[1193,1432,1196,1221]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2014 b) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. http: // dx. doi. org / 10.1111 / syen. 12044" type="journal article" year="2014" yearSuffix="b">Jałoszyński (2014b))</bibRefCitation>
. The intermesocoxal region in
<taxonomicName id="4C084D3C2928FFE0FE7DC1E29F10FB39" box="[489,580,1231,1256]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FE7DC1E29F10FB39" box="[489,580,1231,1256]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
is nearly identical to that of
<taxonomicName id="4C084D3C2928FFE0FC1FC1E29EB6FB39" box="[907,994,1231,1256]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC1FC1E29EB6FB39" box="[907,994,1231,1256]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D3C2928FFE0FB8EC1E299F6FB39" box="[1050,1186,1231,1256]" class="Insecta" family="Staphylinidae" genus="Archeutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FB8EC1E299F6FB39" box="[1050,1186,1231,1256]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C084D3C2928FFE0FB24C1E2985FFB39" box="[1200,1291,1231,1256]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FB24C1E2985FFB39" box="[1200,1291,1231,1256]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
differs from these two genera in absent or rudimentary basal elytral foveae (the elytral base is well exposed in two studied inclusions, but the foveae are not discernible), and from
<taxonomicName id="4C084D3C2928FFE0FCB2C03A9EFAFAE1" box="[806,942,1303,1328]" class="Insecta" family="Staphylinidae" genus="Archeutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FCB2C03A9EFAFAE1" box="[806,942,1303,1328]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
</taxonomicName>
additionally in complete hypomeral ridges (absent in the Cretaceous taxon). Moreover, the submentum in
<taxonomicName id="4C084D3C2928FFE0FCCDC0119EE4FA84" box="[857,944,1340,1365]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FCCDC0119EE4FA84" box="[857,944,1340,1365]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
is short and transverse, whereas in
<taxonomicName id="4C084D3C2928FFE0FAAAC01198CCFA84" box="[1342,1432,1340,1365]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FAAAC01198CCFA84" box="[1342,1432,1340,1365]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
it is similar to that in
<emphasis id="B97CEAAD2928FFE0FE12C0729CBBFAA9" box="[390,495,1375,1400]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C084D3C2928FFE0FE12C0729CBFFAA9" box="[390,491,1375,1400]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Veraphis</taxonomicName>
,
</emphasis>
i.e., long and subhexagonal.
</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C0A99F13F9D9" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
The morphological structures of the new genus seem to represent a link between
<taxonomicName id="4C084D3C2928FFE0FBE9C0A99980FA4C" box="[1149,1236,1412,1437]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FBE9C0A99980FA4C" box="[1149,1236,1412,1437]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D3C2928FFE0FA85C0A99827FA4C" box="[1297,1395,1412,1437]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FA85C0A99827FA4C" box="[1297,1395,1412,1437]" italics="true" pageId="17" pageNumber="18">Veraphis</emphasis>
</taxonomicName>
, or possibly even an ancestral combination of characters, some of which have been preserved in extant species of these two genera. Key morphological characters of
<taxonomicName id="4C084D3C2928FFE0FD45C0E19E6CFA34" box="[721,824,1484,1509]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Eutheiini">Eutheiini</taxonomicName>
with four-segmented maxillary palps and conical palpomere IV are compiled in
<tableCitation id="C68A03042928FFE0FE7AC0C29F10F9D9" box="[494,580,1519,1544]" captionStart="TABLE 2" captionStartId="17.[151,239,1592,1615]" captionTargetBox="[159,1409,1716,1995]" captionTargetPageId="17" captionText="TABLE 2. Comparison of key morphological characters of the genera of Eutheiini with four-segmented maxillary palps and subconical maxillary palpomere IV." httpUri="http://table.plazi.org/id/DF7766372928FFE0FF03C3159F15F9BE" pageId="17" pageNumber="18" tableUuid="DF7766372928FFE0FF03C3159F15F9BE">Table 2</tableCitation>
.
</paragraph>
<caption id="DF7766372928FFE0FF03C3159F15F9BE" ID-Table-UUID="DF7766372928FFE0FF03C3159F15F9BE" httpUri="http://table.plazi.org/id/DF7766372928FFE0FF03C3159F15F9BE" pageId="17" pageNumber="18" targetBox="[159,1409,1716,1995]" targetIsTable="true" targetPageId="17">
<paragraph id="8BB736BF2928FFE0FF03C3159F15F9BE" blockId="17.[151,1436,1592,1647]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FF03C3159C5EF99E" bold="true" box="[151,266,1592,1615]" pageId="17" pageNumber="18">TABLE 2.</emphasis>
Comparison of key morphological characters of the genera of
<taxonomicName id="4C084D3C2928FFE0FC3DC315995FF99E" box="[937,1035,1592,1615]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Eutheiini">Eutheiini</taxonomicName>
with four-segmented maxillary palps and subconical maxillary palpomere IV.
</paragraph>
</caption>
<paragraph id="8BB736BF2928FFE0FE5BC3A7984AF971" blockId="17.[463,1310,1674,1696]" box="[463,1310,1674,1696]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FE5BC3A79E76F971" box="[463,802,1674,1696]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C084D3C2928FFE0FE5BC3A79F4AF971" box="[463,542,1674,1696]" class="Insecta" family="Staphylinidae" genus="Eutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Eutheia</taxonomicName>
<taxonomicName id="4C084D3C2928FFE0FD5DC3A79E76F971" box="[713,802,1674,1696]" class="Insecta" family="Staphylinidae" genus="Veraphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Veraphis</taxonomicName>
</emphasis>
†
<taxonomicName id="4C084D3C2928FFE0FC77C3A7990AF971" authority="Vertheia" authorityName="Vertheia" box="[995,1118,1674,1696]" class="Insecta" family="Staphylinidae" genus="Archeutheia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD2928FFE0FC77C3A7990AF971" box="[995,1118,1674,1696]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
</taxonomicName>
†
<taxonomicName id="4C084D3C2928FFE0FB5EC3A7984AF971" box="[1226,1310,1674,1696]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FB5EC3A7984AF971" box="[1226,1310,1674,1696]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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