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<document id="395E796B7825B8FD59DB0980545DF6FD" ID-DOI="10.12782/specdiv.28.165" ID-ISSN="2189-7301" ID-Zenodo-Dep="10833160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704916610969" checkinUser="felipe" docAuthor="Kobayashi, Hirozumi, Nishigaki, Koji, Saeki, Toshifumi &amp; Maeda, Ken" docDate="2023" docId="03A187A91264820EFE8EF834FDA3FAD0" docLanguage="en" docName="SpeciesDiversity.28.2.165-175.pdf" docOrigin="Species Diversity 28 (2)" docSource="http://dx.doi.org/10.12782/specdiv.28.165" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Odonteleotris macrodon" docType="treatment" docVersion="2" lastPageNumber="173" masterDocId="FF98FFD112658206FF8FFF97FFF8FFA1" masterDocTitle="First Records of Paloa villadolidi from Japan with a Redescription of Odonteleotris macrodon (Teleostei: Butidae)" masterLastPageNumber="175" masterPageNumber="165" pageNumber="166" updateTime="1710777812586" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B3419AC2CE01E932F16CD0F5DC23D0B1">First Records of Paloa villadolidi from Japan with a Redescription of Odonteleotris macrodon (Teleostei: Butidae)</mods:title>
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<mods:affiliation id="45E790BE890C6CECF1C0C066EC58568F">Okinawa Institute for the Conservation of the Environment Co. Ltd., 7 - 11 Suzaki, Uruma, Okinawa 904 - 2234, Japan</mods:affiliation>
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<mods:namePart id="C192DA506618B838E3F84A50420E23A6">Maeda, Ken</mods:namePart>
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<taxonomicName id="4C084D3C12648207FE8EF834FD62F81A" ID-CoL="48NPM" authority="(Bleeker, 1853)" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[257,666,1955,1979]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12648207FE8EF834FE05F81A" bold="true" box="[257,509,1955,1979]" italics="true" pageId="1" pageNumber="166">Odonteleotris macrodon</emphasis>
(
<bibRefCitation id="EF994B4E12648207FD86F834FD6BF81A" author="Bleeker, P." box="[521,659,1955,1979]" pageId="1" pageNumber="166" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="13332A3A12648207FEC3F853FE72F87D" box="[332,394,1988,2012]" captionStart="Fig" captionStartId="2.[156,187,528,550]" captionTargetBox="[208,1378,208,487]" captionTargetId="figure-411@2.[206,1381,205,489]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Eleotris macrodon Bleeker, 1853 (RMNH.PISC. 4472, 132.5mm SL)." figureDoi="http://doi.org/10.5281/zenodo.10833162" httpUri="https://zenodo.org/record/10833162/files/figure.png" pageId="1" pageNumber="166">Figs 1</figureCitation>
,
<figureCitation id="13332A3A12648207FE1AF853FE4DF87D" box="[405,437,1988,2012]" captionStart="Fig" captionStartId="2.[156,187,1350,1372]" captionTargetBox="[208,1379,610,1309]" captionTargetId="figure-426@2.[206,1381,609,1311]" captionTargetPageId="2" captionText="Fig. 2. Dentition in upper and lower jaws of Odonteleotris macrodon (A: RMNH.PISC. 4472, holotype of Eleotris macrodon, 132.5mm SL) and Paloa villadolidi (B: URM-P 49659, 55.1mm SL). UCT and LCT, magnified illustrations of enlarged canine teeth in the upper and lower jaws, respectively." figureDoi="http://doi.org/10.5281/zenodo.10833164" httpUri="https://zenodo.org/record/10833164/files/figure.png" pageId="1" pageNumber="166">2A</figureCitation>
,
<figureCitation id="13332A3A12648207FE31F853FE03F87D" box="[446,507,1988,2012]" captionStart="Fig" captionStartId="3.[156,187,1191,1213]" captionTargetBox="[177,745,206,1151]" captionTargetId="figure-696@3.[176,746,205,1153]" captionTargetPageId="3" captionText="Fig. 3. Illustrations of specimens of Odonteleotris and Paloa. A, Holotype of Eleotris macrodon from Bleeker (1853: pl. 2); B, E. macrodon from Day (1878: pl. 65, fig. 5); C, O. macrodon from Koumans (1953: fig. 81); D, holotype of P. polylepis from Herre (1927: pl. 4, fig. 3); E: holotype of P. villadolidi from Roxas and Ablan (1940: pl. 3)." figureDoi="http://doi.org/10.5281/zenodo.10833166" httpUri="https://zenodo.org/record/10833166/files/figure.png" pageId="1" pageNumber="166">3AC</figureCitation>
;
<tableCitation id="C68A030412648207FD8BF853FDB3F87D" box="[516,587,1988,2012]" captionStart="Table 1" captionStartId="5.[156,204,190,212]" captionText="Table 1. Meristic and morphometric characters of Odonteleotris macrodon and species of Paloa. Data of P. polylepis and P. villadolidi were compiled from descriptions of Herre (1927) and Roxas and Ablan (1940), respectively. Measurements were expressed as a percentage of standard length (SL). Head lengths were expressed also as a percentage of total length (TL) for comparison. Mode of counts was shown in parentheses." pageId="1" pageNumber="166">Table 1</tableCitation>
)
</heading>
</paragraph>
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<emphasis id="B97CEAAD12648207FF13F792FEA9F7BC" box="[156,337,2053,2077]" italics="true" pageId="1" pageNumber="166">Eleotris macrodon</emphasis>
<bibRefCitation id="EF994B4E12648207FED8F792FDF6F7BC" author="Bleeker, P." box="[343,526,2053,2077]" pageId="1" pageNumber="166" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker, 1853: 104</bibRefCitation>
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(
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locality: Hooghly River in Kolkata,
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: pl. 2, fig. 1 [plates of
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separately printed];
<treatmentCitation id="0AA910AE12648207FB59FF70FA6FFF5E" author="Gunther, A." box="[1238,1431,231,255]" page="129" pageId="1" pageNumber="166" year="1861">
<bibRefCitation id="EF994B4E12648207FB59FF70FA6FFF5E" author="Gunther, A." box="[1238,1431,231,255]" pageId="1" pageNumber="166" refId="ref8321" refString="Gunther, A. 1861. Catalogue of the Acanthopterygian Fishes in the Collection of the British Museum. Vol. 3. British Museum, London, xxv + 586 pp. doi: 10.5962 / bhl. title. 8809 [in part]" type="book" year="1861">Günther 1861: 129</bibRefCitation>
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[Calcutta (currently Kolkata),
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];
<treatmentCitation id="0AA910AE12648207FB57FE9FFA88FE81" author="Day, F." box="[1240,1392,264,288]" page="311" pageId="1" pageNumber="166" year="1878">
<bibRefCitation id="EF994B4E12648207FB57FE9FFA88FE81" author="Day, F." box="[1240,1392,264,288]" pageId="1" pageNumber="166" pagination="139 - 195" refId="ref8162" refString="Day, F. 1878. The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma, and Ceylon. Quaritsch, London, i - xx + 553 - 778 pp., pls 139 - 195. doi: 10.5962 / bhl. title. 62705 [in part]" type="book chapter" year="1878">Day 1878: 311</bibRefCitation>
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, pl. 65, fig. 5 [Bengal,
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;
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(
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<emphasis id="B97CEAAD12648207FCBAFEDFFBCAFEC1" box="[821,1074,328,352]" italics="true" pageId="1" pageNumber="166">Odonteleotris macrodon</emphasis>
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:
<treatmentCitation id="0AA910AE12648207FBC2FEDFFAE7FEC0" author="Bleeker, P." box="[1101,1311,328,353]" page="302" pageId="1" pageNumber="166" year="1874">
<bibRefCitation id="EF994B4E12648207FBC2FEDFFAE7FEC0" author="Bleeker, P." box="[1101,1311,328,353]" pageId="1" pageNumber="166" pagination="289 - 331" refId="ref8136" refString="Bleeker, P. 1874. Esquisse d'un systeme naturel des Gobioides. Archives neerlandaises des sciences exactes et naturelles 9: 289 - 331." type="journal article" year="1874">Bleeker 1874: 302</bibRefCitation>
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;
<treatmentCitation id="0AA910AE12648207FAB9FEDEFC4AFE20" author="Koumans, F. P." page="328" pageId="1" pageNumber="166" year="1941">
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[Calcutta (Kolkata) and Bengal,
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; Akyab (Sittwe),
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; Nias and
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,
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<treatmentCitation id="0AA910AE12648207FAB9FE1DFC40FE63" author="Koumans, F. P." page="330" pageId="1" pageNumber="166" year="1953">
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</treatmentCitation>
, fig. 81 [
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, Nias, and
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; Calcutta (Kolkata),
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; Akyab (Sittwe),
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; Malaya (
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); Port Darwin,
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(with question)];
<treatmentCitation id="0AA910AE12648207FB8CFD9BFABAFD85" author="Talwar, P. K. &amp; Jhingran, A. G." box="[1027,1346,524,548]" page="978" pageId="1" pageNumber="166" year="1991">
<bibRefCitation id="EF994B4E12648207FB8CFD9BFABAFD85" author="Talwar, P. K. &amp; Jhingran, A. G." box="[1027,1346,524,548]" pageId="1" pageNumber="166" refId="ref9081" refString="Talwar, P. K. and Jhingran, A. G. 1991. Inland Fishes of India and Adjacent Countries. Oxford &amp; IBH Publishing Company, New Delhi, 1158 pp., 3 pls [two volumes]." type="book" year="1991">Talwar and Jhingran 1991: 978</bibRefCitation>
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(
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);
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<bibRefCitation id="EF994B4E12648207FCC1FDBBFBBAFDE5" author="Kottelat, M. &amp; Whitten, T. &amp; Kartikasari, S. N. &amp; Wirjoatmodjo, S." box="[846,1090,556,580]" pageId="1" pageNumber="166" refId="ref8513" refString="Kottelat, M., Whitten, T., Kartikasari, S. N., and Wirjoatmodjo, S. 1993. Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp., 84 pls." type="book" year="1993">Kottelat et al. 1993: 186</bibRefCitation>
</treatmentCitation>
, fig. 272 (
<collectingCountry id="F31F762F12648207FB27FDBBFAE9FDE5" box="[1192,1297,556,580]" name="Indonesia" pageId="1" pageNumber="166">Indonesia</collectingCountry>
);
<treatmentCitation id="0AA910AE12648207FAABFDBAFBFFFDC4" author="Larson, H. K. &amp; Murdy, E. O." page="3577" pageId="1" pageNumber="166" year="2001">
<bibRefCitation id="EF994B4E12648207FAABFDBAFBFFFDC4" author="Larson, H. K. &amp; Murdy, E. O." pageId="1" pageNumber="166" pagination="3574 - 3603" refId="ref8631" refString="Larson, H. K. and Murdy, E. O. 2001. Families Eleotridae, Gobiidae. Pp. 3574 - 3603. In: Carpenter, K. E. and Niem, V. H. (Eds) FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Volume 6 Bony Fishes Part 4 (Labridae to Latimeriidae), Estuarine Crocodiles, Sea Turtles, Sea Snakes and Marine Mammals. FAO, Rome." type="book chapter" year="2001">Larson and Murdy 2001: 3577</bibRefCitation>
</treatmentCitation>
(list only);
<treatmentCitation id="0AA910AE12648207FBF2FDDAFAAAFDC4" author="Satapoomin, U." box="[1149,1362,589,613]" page="65" pageId="1" pageNumber="166" year="2011">
<bibRefCitation id="EF994B4E12648207FBF2FDDAFAAAFDC4" author="Satapoomin, U." box="[1149,1362,589,613]" pageId="1" pageNumber="166" pagination="29 - 77" refId="ref9042" refString="Satapoomin, U. 2011. The fishes of southwestern Thailand, the Andaman Sea - a review of research and a provisional checklist of species. Phuket Marine Biological Center Research Bulletin 70: 29 - 77." type="journal article" year="2011">Satapoomin 2011: 65</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F31F762F12648207FAEFFDDAFC86FD24" name="Thailand" pageId="1" pageNumber="166">Thailand</collectingCountry>
);
<treatmentCitation id="0AA910AE12648207FC1FFDFAFBB2FD24" author="Kottelat, M." box="[912,1098,621,645]" page="394" pageId="1" pageNumber="166" year="2013">
<bibRefCitation id="EF994B4E12648207FC1FFDFAFBB2FD24" author="Kottelat, M." box="[912,1098,621,645]" pageId="1" pageNumber="166" pagination="1 - 663" refId="ref8467" refString="Kottelat, M. 2013. The fishes of the inland waters of Southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology, Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat 2013: 394</bibRefCitation>
</treatmentCitation>
(list only);
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<bibRefCitation id="EF994B4E12648207FB4CFDFAFA91FD24" author="Parenti, P." box="[1219,1385,621,645]" pageId="1" pageNumber="166" pagination="1 - 480" refId="ref8866" refString="Parenti, P. 2021. A checklist of the gobioid fishes of the world (Percomorpha: Gobiiformes). Iranian Journal of Ichthyology 8 (Supplement 1): 1 - 480. doi: 10.22034 / iji. v 8 i 0.556" type="journal article" year="2021">Parenti 2021: 23</bibRefCitation>
</treatmentCitation>
(list only);
<bibRefCitation id="EF994B4E12648207FC1EFD19FB6EFD07" author="Larson, H. K. &amp; Nguyen, T. H." box="[913,1174,654,678]" pageId="1" pageNumber="166" pagination="1139300" refId="ref8721" refString="Larson, H. K. and Nguyen, T. H. 2021. Odonteleotris macrodon. The IUCN Red List of Threatened Species 2021: e. T 166620 A 1139300. doi: 10.2305 / IUCN. UK. 2021 - 1. RLTS. T 166620 A 1139300. en" type="journal article" year="2021">Larson and Nguyen 2021</bibRefCitation>
: [1] [
<collectingRegion id="49CCF85D12648207FB42FD19FAA8FD07" box="[1229,1360,654,678]" country="India" name="West Bengal" pageId="1" pageNumber="166">West Bengal</collectingRegion>
,
<collectingCountry id="F31F762F12648207FAD3FD19FA6BFD07" box="[1372,1427,654,678]" name="India" pageId="1" pageNumber="166">India</collectingCountry>
;
<collectingRegion id="49CCF85D12648207FCC1FD39FC5CFD67" box="[846,932,686,710]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="166">Sulawesi</collectingRegion>
and
<collectingRegion id="49CCF85D12648207FC6BFD39FBC3FD67" box="[996,1083,686,710]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="166">Sumatra</collectingRegion>
,
<collectingCountry id="F31F762F12648207FBC4FD39FB49FD67" box="[1099,1201,686,710]" name="Indonesia" pageId="1" pageNumber="166">Indonesia</collectingCountry>
;
<collectingCountry id="F31F762F12648207FB4DFD39FAE6FD67" box="[1218,1310,686,710]" name="Malaysia" pageId="1" pageNumber="166">Malaysia</collectingCountry>
;
<collectingCountry id="F31F762F12648207FAA1FD38FA6CFD66" box="[1326,1428,687,711]" name="Myanmar" pageId="1" pageNumber="166">Myanmar</collectingCountry>
;
<collectingCountry id="F31F762F12648207FCC1FD58FC5FFD46" box="[846,935,719,743]" name="Thailand" pageId="1" pageNumber="166">Thailand</collectingCountry>
(see Remarks)].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C312653412648207FCC1FC87FB44FCC8" pageId="1" pageNumber="166" type="materials_examined">
<paragraph id="8BB736BF12648207FCC1FC87FB44FCC8" blockId="1.[821,1432,784,2077]" pageId="1" pageNumber="166">
<emphasis id="B97CEAAD12648207FCC1FC87FBEFFC89" bold="true" box="[846,1047,784,808]" pageId="1" pageNumber="166">Material examined.</emphasis>
<collectionCode id="ED19AE7A12648207FBBFFC87FB86FC89" box="[1072,1150,784,808]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="166" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED19AE7A12648207FB0FFC87FB4EFC89" box="[1152,1206,784,808]" pageId="1" pageNumber="166">PISC</collectionCode>
.
<materialsCitation id="3B603CE212648207FB30FC87FBB4FCC8" collectionCode="SL" country="India" location="Hooghly River in Kolkata" pageId="1" pageNumber="166" specimenCount="1" typeStatus="holotype">
4472, male,
<quantity id="4CF09B5A12648207FAB9FC87FA6FFC86" box="[1334,1431,784,808]" metricMagnitude="-1" metricUnit="m" metricValue="1.1059999999999999" pageId="1" pageNumber="166" unit="mm" value="110.6">110.6 mm</quantity>
<collectionCode id="ED19AE7A12648207FCBAFCA6FCA9FCE8" box="[821,849,817,841]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="1" pageNumber="166" type="Herbarium">SL</collectionCode>
,
<typeStatus id="54B3881D12648207FCD1FCA7FC4FFCE9" box="[862,951,816,840]" pageId="1" pageNumber="166" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C084D3C12648207FC6FFCA7FAC9FCE9" authority="Bleeker, 1853" authorityName="Bleeker" authorityYear="1853" box="[992,1329,816,840]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12648207FC6FFCA7FB61FCE9" box="[992,1177,816,840]" italics="true" pageId="1" pageNumber="166">Eleotris macrodon</emphasis>
<bibRefCitation id="EF994B4E12648207FB2BFCA7FAC9FCE9" author="Bleeker, P." box="[1188,1329,816,840]" pageId="1" pageNumber="166" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
</taxonomicName>
,
<location id="8ED7606412648207FAB0FCA7FC23FCC8" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412648207FAB0FCA7FC23FCC8" country="India" name="Hooghly River in Kolkata" pageId="1" pageNumber="166">Hooghly River in Kolkata</location>
,
<collectingCountry id="F31F762F12648207FC6BFCC6FBE3FCC8" box="[996,1051,849,873]" name="India" pageId="1" pageNumber="166">India</collectingCountry>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212648207FBC3FCC6FB9BFCC9" box="[1100,1123,849,873]" collectionCode="SL" country="India" isTailEnumeration="true" location="Hooghly River in Kolkata" pageId="1" pageNumber="166" specimenCount="1" typeStatus="holotype">. P</materialsCitation>
. Bleeker.
</paragraph>
</subSubSection>
<subSubSection id="C312653412648207FCC1FCE6FB02FA33" pageId="1" pageNumber="166" type="diagnosis">
<paragraph id="8BB736BF12648207FCC1FCE6FB02FA33" blockId="1.[821,1432,784,2077]" pageId="1" pageNumber="166">
<emphasis id="B97CEAAD12648207FCC1FCE6FC46FC28" bold="true" box="[846,958,881,905]" pageId="1" pageNumber="166">Diagnosis.</emphasis>
<taxonomicName id="4C084D3C12648207FC58FCE6FB2DFC28" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[983,1237,881,905]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12648207FC58FCE6FB2DFC28" box="[983,1237,881,905]" italics="true" pageId="1" pageNumber="166">Odonteleotris macrodon</emphasis>
</taxonomicName>
is distinguished from all other species of
<taxonomicName id="4C084D3C12648207FBCEFC05FB77FC0B" authorityName="Bleeker" authorityYear="1874" box="[1089,1167,914,938]" class="Actinopteri" family="Butidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiiformes" pageId="1" pageNumber="166" phylum="Chordata" rank="family">Butidae</taxonomicName>
by a combination of the following characters. Enlarged canine teeth on the upper and lower jaws (five and twelve, respectively); canines placed in the outermost tooth row in the upper and anterior lower jaw, while canines placed in the inner tooth row in the posterior lower jaw. Upper jaw ending below center of the eye. Interorbital sensory pore D paired with wide interval. Fourth infraocular transverse row of sensory papillae crossed with a longitudinal row on the cheek. Longitudinal scales 103; transverse scales 32; predorsal scales 63; transverse scales at caudal peduncle 21; circumpeduncle scales 39. Dorsal-fin rays VI-I, 10; anal-fin rays I, 8; pectoral-fin rays 17. Predorsal length 38.2% of SL; body depth at anal-fin origin 18.4% of SL; eye diameter 3.3% of SL. Unpaired fin rays variegated by small blackish spots; a red spot on the upper part of the caudal-fin base when fresh.
</paragraph>
</subSubSection>
<subSubSection id="C312653412648204FCC1FA0DFB7CF916" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166" type="description">
<paragraph id="8BB736BF12648204FCC1FA0DFD16F898" blockId="1.[821,1432,784,2077]" lastBlockId="2.[156,768,1467,2077]" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166">
<emphasis id="B97CEAAD12648207FCC1FA0DFC2AFA13" bold="true" box="[846,978,1434,1458]" pageId="1" pageNumber="166">Description.</emphasis>
Body elongated and cylindrical anteriorly, somewhat compressed posteriorly (
<figureCitation id="13332A3A12648207FB6EFA2CFAD8FA72" box="[1249,1312,1467,1491]" captionStart="Fig" captionStartId="2.[156,187,528,550]" captionTargetBox="[208,1378,208,487]" captionTargetId="figure-411@2.[206,1381,205,489]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Eleotris macrodon Bleeker, 1853 (RMNH.PISC. 4472, 132.5mm SL)." figureDoi="http://doi.org/10.5281/zenodo.10833162" httpUri="https://zenodo.org/record/10833162/files/figure.png" pageId="1" pageNumber="166">Fig. 1</figureCitation>
). No bony crests in the interorbital region. Mouth large and superior, and oblique with an approximately 45º upward angle; upper jaw ending below the center of the eye. Snout length greater than eye diameter. Anterior nostrils tubular and posterior nostrils with low rim. Upper and lower jaws with five and twelve enlarged, curved canines, respectively (
<figureCitation id="13332A3A12648207FA84F9E9FAA3F937" box="[1291,1371,1662,1686]" captionStart="Fig" captionStartId="2.[156,187,1350,1372]" captionTargetBox="[208,1379,610,1309]" captionTargetId="figure-426@2.[206,1381,609,1311]" captionTargetPageId="2" captionText="Fig. 2. Dentition in upper and lower jaws of Odonteleotris macrodon (A: RMNH.PISC. 4472, holotype of Eleotris macrodon, 132.5mm SL) and Paloa villadolidi (B: URM-P 49659, 55.1mm SL). UCT and LCT, magnified illustrations of enlarged canine teeth in the upper and lower jaws, respectively." figureDoi="http://doi.org/10.5281/zenodo.10833164" httpUri="https://zenodo.org/record/10833164/files/figure.png" pageId="1" pageNumber="166">Fig. 2A</figureCitation>
). Five canines in the upper jaw aligned to the anterior half of the outermost tooth row. Five canines in the anterior half of the lower jaw aligned to the outermost tooth row, while seven in the posterior half aligned to the innermost row. Two to four and one to four rows of small conical teeth on upper and lower jaws, respectively. No teeth on the vomer. Small cycloid scales covering the head and body, except around the snout; no accessory scales. Longitudinal scales 98 on the right side, 103 on the left side; transverse scales 32; predorsal scales 63. Urogenital papilla small and oval-shaped in male. First dorsal fin with six spines; second dorsal fin with one spine and ten soft rays. Anal fin with one spine and eight soft rays; anal-fin origin posterior to the second-dorsal-fin origin. Caudal fin with 17 segmented rays. Pectoral fin with 17 soft rays. Pelvic-fin insertion slightly anterior to the pectoral-fin base. Left and right pelvic fins separated from each other, both with one spine and five soft rays. Ten sensory pores (A, B, C, D, E, F, G, H, K, and L) in the oculoscapular canal, three (M, N, and O) and four pores (M, N, O, and P) in the left and right preopercular canals, respectively; pore D paired with an interval of approximately half of the interorbital width. Arrangement of the cephalic sensory papillae in transverse pattern: six transverse and two longitudinal rows of the sensory papillae on the cheek.
</paragraph>
<caption id="DF77663712678204FF13FD87FC3FFD84" ID-DOI="http://doi.org/10.5281/zenodo.10833162" ID-Zenodo-Dep="10833162" box="[156,967,528,550]" httpUri="https://zenodo.org/record/10833162/files/figure.png" pageId="2" pageNumber="167" startId="2.[156,187,528,550]" targetBox="[208,1378,208,487]" targetPageId="2" targetType="figure">
<paragraph id="8BB736BF12678204FF13FD87FC3FFD84" blockId="2.[156,967,528,550]" box="[156,967,528,550]" pageId="2" pageNumber="167">
Fig. 1. Holotype of
<taxonomicName id="4C084D3C12678204FEEEFD87FD87FD87" authority="Bleeker, 1853" authorityName="Bleeker" authorityYear="1853" box="[353,639,528,550]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12678204FEEEFD87FDFAFD84" box="[353,514,528,549]" italics="true" pageId="2" pageNumber="167">Eleotris macrodon</emphasis>
<bibRefCitation id="EF994B4E12678204FD88FD87FD87FD87" author="Bleeker, P." box="[519,639,528,550]" pageId="2" pageNumber="167" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
</taxonomicName>
(RMNH.PISC. 4472, 132.5mm SL).
</paragraph>
</caption>
<caption id="DF77663712678204FF13FAD1FEC5FA35" ID-DOI="http://doi.org/10.5281/zenodo.10833164" ID-Zenodo-Dep="10833164" httpUri="https://zenodo.org/record/10833164/files/figure.png" pageId="2" pageNumber="167" startId="2.[156,187,1350,1372]" targetBox="[208,1379,610,1309]" targetPageId="2" targetType="figure">
<paragraph id="8BB736BF12678204FF13FAD1FEC5FA35" blockId="2.[156,1432,1350,1428]" pageId="2" pageNumber="167">
Fig. 2. Dentition in upper and lower jaws of
<taxonomicName id="4C084D3C12678204FDC8FAD1FCE6FAFA" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[583,798,1350,1371]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12678204FDC8FAD1FCE6FAFA" box="[583,798,1350,1371]" italics="true" pageId="2" pageNumber="167">Odonteleotris macrodon</emphasis>
</taxonomicName>
(A: RMNH.PISC. 4472, holotype of
<taxonomicName id="4C084D3C12678204FBE2FAD1FAE8FAFA" authorityName="Bleeker" authorityYear="1853" box="[1133,1296,1350,1371]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12678204FBE2FAD1FAE8FAFA" box="[1133,1296,1350,1371]" italics="true" pageId="2" pageNumber="167">Eleotris macrodon</emphasis>
</taxonomicName>
, 132.5 mm SL) and
<taxonomicName id="4C084D3C12678204FF4BFAF5FEADFAD6" authorityName="Roxas and Ablan" authorityYear="1940" box="[196,341,1378,1399]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12678204FF4BFAF5FEADFAD6" box="[196,341,1378,1399]" italics="true" pageId="2" pageNumber="167">Paloa villadolidi</emphasis>
</taxonomicName>
(B: URM-P 49659, 55.1 mm SL).
<emphasis id="B97CEAAD12678204FD08FAF5FD4BFAD6" box="[647,691,1378,1399]" italics="true" pageId="2" pageNumber="167">UCT</emphasis>
and
<emphasis id="B97CEAAD12678204FD6EFAF5FCF1FAD6" box="[737,777,1378,1399]" italics="true" pageId="2" pageNumber="167">LCT</emphasis>
, magnified illustrations of enlarged canine teeth in the upper and lower jaws, respectively.
</paragraph>
</caption>
<paragraph id="8BB736BF12678204FF3AF8D6FB7CF916" blockId="2.[156,768,1467,2077]" lastBlockId="2.[820,1432,1467,2077]" pageId="2" pageNumber="167">
According to
<bibRefCitation id="EF994B4E12678204FEC6F8D6FDB2F8F8" author="Bleeker, P." box="[329,586,1857,1881]" pageId="2" pageNumber="167" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker (1853: 104105)</bibRefCitation>
, the original description, body blackish green dorsally and rosy ventrally. Cheek with 24 black bars radiating from the eye. Dorsal and caudal fins with reddish rays variegated by black spots. Dorsal rays of caudal fin bordered with red. Pectoral fin olive; pelvic and anal fins violet. Anterior part of the pelvic fin and lower part of the anal fin bordered with red. A red spot on the upper part of the caudal-fin base. Anal-fin rays also variegated by black spots (
<figureCitation id="13332A3A12678204FB12FA4BFB14FA55" box="[1181,1260,1500,1524]" captionStart="Fig" captionStartId="3.[156,187,1191,1213]" captionTargetBox="[177,745,206,1151]" captionTargetId="figure-696@3.[176,746,205,1153]" captionTargetPageId="3" captionText="Fig. 3. Illustrations of specimens of Odonteleotris and Paloa. A, Holotype of Eleotris macrodon from Bleeker (1853: pl. 2); B, E. macrodon from Day (1878: pl. 65, fig. 5); C, O. macrodon from Koumans (1953: fig. 81); D, holotype of P. polylepis from Herre (1927: pl. 4, fig. 3); E: holotype of P. villadolidi from Roxas and Ablan (1940: pl. 3)." figureDoi="http://doi.org/10.5281/zenodo.10833166" httpUri="https://zenodo.org/record/10833166/files/figure.png" pageId="2" pageNumber="167">Fig. 3A</figureCitation>
).
<bibRefCitation id="EF994B4E12678204FB71FA4BFA88FA55" author="Day, F." box="[1278,1392,1500,1524]" pageId="2" pageNumber="167" pagination="139 - 195" refId="ref8162" refString="Day, F. 1878. The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma, and Ceylon. Quaritsch, London, i - xx + 553 - 778 pp., pls 139 - 195. doi: 10.5962 / bhl. title. 62705 [in part]" type="book chapter" year="1878">Day (1878)</bibRefCitation>
described “a dark ocellus edged with light” at the upper part of the caudal-fin base instead of the red spot (
<figureCitation id="13332A3A12678204FB65F98AFACEF994" box="[1258,1334,1565,1589]" captionStart="Fig" captionStartId="3.[156,187,1191,1213]" captionTargetBox="[177,745,206,1151]" captionTargetId="figure-696@3.[176,746,205,1153]" captionTargetPageId="3" captionText="Fig. 3. Illustrations of specimens of Odonteleotris and Paloa. A, Holotype of Eleotris macrodon from Bleeker (1853: pl. 2); B, E. macrodon from Day (1878: pl. 65, fig. 5); C, O. macrodon from Koumans (1953: fig. 81); D, holotype of P. polylepis from Herre (1927: pl. 4, fig. 3); E: holotype of P. villadolidi from Roxas and Ablan (1940: pl. 3)." figureDoi="http://doi.org/10.5281/zenodo.10833166" httpUri="https://zenodo.org/record/10833166/files/figure.png" pageId="2" pageNumber="167">Fig. 3B</figureCitation>
). The
<typeStatus id="54B3881D12678204FAFBF98AFC8BF9F4" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
observed after more than 167 years of preservation, having uniformly brown head and body without any markings, and pale-brown dorsal, caudal, pectoral, and pelvic fins anal dark-brown anal fin (
<figureCitation id="13332A3A12678204FBB4F908FB8DF916" box="[1083,1141,1695,1719]" captionStart="Fig" captionStartId="2.[156,187,528,550]" captionTargetBox="[208,1378,208,487]" captionTargetId="figure-411@2.[206,1381,205,489]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Eleotris macrodon Bleeker, 1853 (RMNH.PISC. 4472, 132.5mm SL)." figureDoi="http://doi.org/10.5281/zenodo.10833162" httpUri="https://zenodo.org/record/10833162/files/figure.png" pageId="2" pageNumber="167">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C312653412678205FCC1F928FDA7F83A" lastPageId="3" lastPageNumber="168" pageId="2" pageNumber="167" type="discussion">
<paragraph id="8BB736BF12678205FCC1F928FE43FA12" blockId="2.[820,1432,1467,2077]" lastBlockId="3.[156,767,1402,1816]" lastPageId="3" lastPageNumber="168" pageId="2" pageNumber="167">
<emphasis id="B97CEAAD12678204FCC1F928FC49F976" bold="true" box="[846,945,1727,1751]" pageId="2" pageNumber="167">Remarks.</emphasis>
The morphology of RMNH.PISC. 4472 corresponds well to that described by
<bibRefCitation id="EF994B4E12678204FB29F977FAB8F959" author="Bleeker, P." box="[1190,1344,1760,1784]" pageId="2" pageNumber="167" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
.
<bibRefCitation id="EF994B4E12678204FAC3F977FC8EF8B9" author="Bleeker, P." pageId="2" pageNumber="167" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker (1853)</bibRefCitation>
noted that the
<typeStatus id="54B3881D12678204FBAEF897FB82F8B9" box="[1057,1146,1792,1816]" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
had a total length (TL) of
<quantity id="4CF09B5A12678204FCBBF8B6FC7FF899" box="[820,903,1825,1849]" metricMagnitude="-1" metricUnit="m" metricValue="1.34" pageId="2" pageNumber="167" unit="mm" value="134.0">134 mm</quantity>
, and the specimen was measured as
<quantity id="4CF09B5A12678204FA9EF8B6FA8CF899" box="[1297,1396,1825,1849]" metricMagnitude="-1" metricUnit="m" metricValue="1.1059999999999999" pageId="2" pageNumber="167" unit="mm" value="110.6">110.6 mm</quantity>
SL and
<quantity id="4CF09B5A12678204FCE8F8D6FC31F8F8" box="[871,969,1857,1881]" metricMagnitude="-1" metricUnit="m" metricValue="1.325" pageId="2" pageNumber="167" unit="mm" value="132.5">132.5 mm</quantity>
TL in our examinations (probably due to shrinkage over time). Although our examination of the
<typeStatus id="54B3881D12678204FAFBF8F5FC8BF83B" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
shows more canines in the lower jaw (12) than the upper jaw (5),
<bibRefCitation id="EF994B4E12678204FC59F834FB7BF81A" author="Koumans, F. P." box="[982,1155,1955,1979]" pageId="2" pageNumber="167" refId="ref8582" refString="Koumans, F. P. 1953. Gobioidea. In: Weber, M. and de Beaufort, L. F. (Eds) Fishes of the Indo-Australian Archipelago. Volume 10. E. J. Brill, Leiden, xiii + 423 pp." type="book" year="1953">Koumans (1953)</bibRefCitation>
reported the opposite arrangement, with more canines in the upper jaw (46) than in the lower jaw (4). However, this difference may be due to the fact that he only counted the canines on the outer row of the lower jaw and did not include those on the inner side of the posterior part of the jaw.
</paragraph>
<caption id="DF77663712668205FF13FB30FEB1FAE8" ID-DOI="http://doi.org/10.5281/zenodo.10833166" ID-Zenodo-Dep="10833166" httpUri="https://zenodo.org/record/10833166/files/figure.png" pageId="3" pageNumber="168" startId="3.[156,187,1191,1213]" targetBox="[177,745,206,1151]" targetPageId="3" targetType="figure">
<paragraph id="8BB736BF12668205FF13FB30FEB1FAE8" blockId="3.[156,767,1191,1353]" pageId="3" pageNumber="168">
Fig. 3. Illustrations of specimens of
<taxonomicName id="4C084D3C12668205FD9BFB30FD73FB1D" authorityName="Gill" authorityYear="1863" box="[532,651,1191,1212]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="genus">
<emphasis id="B97CEAAD12668205FD9BFB30FD73FB1D" box="[532,651,1191,1212]" italics="true" pageId="3" pageNumber="168">Odonteleotris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D3C12668205FD47FB30FD02FB1D" authorityName="Herre" authorityYear="1927" box="[712,762,1191,1212]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="genus">
<emphasis id="B97CEAAD12668205FD47FB30FD02FB1D" box="[712,762,1191,1212]" italics="true" pageId="3" pageNumber="168">Paloa</emphasis>
</taxonomicName>
. A, Holotype of
<taxonomicName id="4C084D3C12668205FEBAFB54FE23FB79" authorityName="Bleeker" authorityYear="1853" box="[309,475,1219,1240]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12668205FEBAFB54FE23FB79" box="[309,475,1219,1240]" italics="true" pageId="3" pageNumber="168">Eleotris macrodon</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF994B4E12668205FD94FB54FD66FB78" author="Bleeker, P." box="[539,670,1219,1241]" pageId="3" pageNumber="168" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker (1853</bibRefCitation>
: pl. 2); B,
<taxonomicName id="4C084D3C12668205FF13FB77FEF7FB55" authorityName="Bleeker" authorityYear="1853" box="[156,271,1247,1269]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12668205FF13FB77FEF7FB55" box="[156,271,1247,1269]" italics="true" pageId="3" pageNumber="168">E. macrodon</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF994B4E12668205FEC4FB77FE54FB54" author="Day, F." box="[331,428,1248,1269]" pageId="3" pageNumber="168" pagination="139 - 195" refId="ref8162" refString="Day, F. 1878. The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma, and Ceylon. Quaritsch, London, i - xx + 553 - 778 pp., pls 139 - 195. doi: 10.5962 / bhl. title. 62705 [in part]" type="book chapter" year="1878">Day (1878</bibRefCitation>
: pl. 65, fig. 5); C,
<taxonomicName id="4C084D3C12668205FDDAFB77FD33FB55" authorityName="Bleeker" authorityYear="1853" box="[597,715,1247,1269]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12668205FDDAFB77FD33FB55" box="[597,715,1247,1269]" italics="true" pageId="3" pageNumber="168">O. macrodon</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF994B4E12668205FF13FB6BFECAFAB0" author="Koumans, F. P." box="[156,306,1275,1297]" pageId="3" pageNumber="168" refId="ref8582" refString="Koumans, F. P. 1953. Gobioidea. In: Weber, M. and de Beaufort, L. F. (Eds) Fishes of the Indo-Australian Archipelago. Volume 10. E. J. Brill, Leiden, xiii + 423 pp." type="book" year="1953">Koumans (1953</bibRefCitation>
: fig. 81); D, holotype of
<taxonomicName id="4C084D3C12668205FDAAFB6BFD72FAB1" authorityName="Herre" authorityYear="1927" box="[549,650,1275,1297]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="B97CEAAD12668205FDAAFB6BFD72FAB1" box="[549,650,1275,1297]" italics="true" pageId="3" pageNumber="168">P. polylepis</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF994B4E12668205FD45FB6BFF2AFA8C" author="Herre, A. W. C. T." pageId="3" pageNumber="168" pagination="1 - 352" refId="ref8367" refString="Herre, A. W. C. T. 1927. Gobies of the Philippines and the China Sea. Monographs of the Bureau of Science Manila, Philippine Islands 23: 1 - 352, pls 1 - 30." type="journal article" year="1927">Herre (1927</bibRefCitation>
: pl. 4, fig. 3); E: holotype of
<taxonomicName id="4C084D3C12668205FE62FA8FFDA6FA8D" authorityName="Roxas and Ablan" authorityYear="1940" box="[493,606,1303,1325]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12668205FE62FA8FFDA6FA8D" box="[493,606,1303,1325]" italics="true" pageId="3" pageNumber="168">P. villadolidi</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF994B4E12668205FD12FA8FFEF5FAE8" author="Roxas, H. A. &amp; Ablan, G. L." pageId="3" pageNumber="168" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan (1940</bibRefCitation>
: pl. 3).
</paragraph>
</caption>
<paragraph id="8BB736BF12668205FF3AFA2CFE43F8B9" blockId="3.[156,767,1402,1816]" pageId="3" pageNumber="168">
<taxonomicName id="4C084D3C12668205FF3AFA2CFE54FA72" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[181,428,1467,1491]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12668205FF3AFA2CFE54FA72" box="[181,428,1467,1491]" italics="true" pageId="3" pageNumber="168">Odonteleotris macrodon</emphasis>
</taxonomicName>
has been reported in Kolkata,
<collectingCountry id="F31F762F12668205FF13FA4BFF29FA55" box="[156,209,1500,1524]" name="India" pageId="3" pageNumber="168">India</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FF6FFA4BFE91FA55" author="Bleeker, P." box="[224,361,1500,1524]" pageId="3" pageNumber="168" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker 1853</bibRefCitation>
);
<collectingCountry id="F31F762F12668205FEF3FA4BFE27FA55" box="[380,479,1500,1524]" name="Myanmar" pageId="3" pageNumber="168">Myanmar</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FE61FA4BFDACFA55" author="Day, F." box="[494,596,1500,1524]" pageId="3" pageNumber="168" pagination="139 - 195" refId="ref8162" refString="Day, F. 1878. The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Fresh Waters of India, Burma, and Ceylon. Quaritsch, London, i - xx + 553 - 778 pp., pls 139 - 195. doi: 10.5962 / bhl. title. 62705 [in part]" type="book chapter" year="1878">Day 1878</bibRefCitation>
;
<bibRefCitation id="EF994B4E12668205FDD0FA4BFD03FA55" author="Koumans, F. P." box="[607,763,1500,1524]" pageId="3" pageNumber="168" refId="ref8582" refString="Koumans, F. P. 1953. Gobioidea. In: Weber, M. and de Beaufort, L. F. (Eds) Fishes of the Indo-Australian Archipelago. Volume 10. E. J. Brill, Leiden, xiii + 423 pp." type="book" year="1953">Koumans 1953</bibRefCitation>
;
<bibRefCitation id="EF994B4E12668205FF13FA6BFE4EF9B5" author="Talwar, P. K. &amp; Jhingran, A. G." box="[156,438,1532,1556]" pageId="3" pageNumber="168" refId="ref9081" refString="Talwar, P. K. and Jhingran, A. G. 1991. Inland Fishes of India and Adjacent Countries. Oxford &amp; IBH Publishing Company, New Delhi, 1158 pp., 3 pls [two volumes]." type="book" year="1991">Talwar and Jhingran 1991</bibRefCitation>
);
<collectingCountry id="F31F762F12668205FE42FA6BFDDEF9B5" box="[461,550,1532,1556]" name="Thailand" pageId="3" pageNumber="168">Thailand</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FDB7FA6BFD0CF9B5" author="Satapoomin, U." box="[568,756,1532,1556]" pageId="3" pageNumber="168" pagination="29 - 77" refId="ref9042" refString="Satapoomin, U. 2011. The fishes of southwestern Thailand, the Andaman Sea - a review of research and a provisional checklist of species. Phuket Marine Biological Center Research Bulletin 70: 29 - 77." type="journal article" year="2011">Satapoomin 2011</bibRefCitation>
); Eastern
<collectingRegion id="49CCF85D12668205FF7AF98AFED8F995" box="[245,288,1565,1588]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="168">Java</collectingRegion>
, Nias, and
<collectingRegion id="49CCF85D12668205FE2DF98AFE00F994" box="[418,504,1565,1589]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="168">Sulawesi</collectingRegion>
in
<collectingCountry id="F31F762F12668205FDAAF98AFD71F994" box="[549,649,1565,1589]" name="Indonesia" pageId="3" pageNumber="168">Indonesia</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FD13F98AFF28F9F4" author="Koumans, F. P." pageId="3" pageNumber="168" refId="ref8582" refString="Koumans, F. P. 1953. Gobioidea. In: Weber, M. and de Beaufort, L. F. (Eds) Fishes of the Indo-Australian Archipelago. Volume 10. E. J. Brill, Leiden, xiii + 423 pp." type="book" year="1953">Koumans 1953</bibRefCitation>
);
<collectingCountry id="F31F762F12668205FF65F9AAFEBBF9F4" box="[234,323,1597,1621]" name="Malaysia" pageId="3" pageNumber="168">Malaysia</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FED6F9AAFE06F9F4" author="Koumans, F. P." box="[345,510,1597,1621]" pageId="3" pageNumber="168" refId="ref8582" refString="Koumans, F. P. 1953. Gobioidea. In: Weber, M. and de Beaufort, L. F. (Eds) Fishes of the Indo-Australian Archipelago. Volume 10. E. J. Brill, Leiden, xiii + 423 pp." type="book" year="1953">Koumans 1953</bibRefCitation>
); and
<collectingCountry id="F31F762F12668205FDC2F9AAFD51F9F4" box="[589,681,1597,1621]" name="Australia" pageId="3" pageNumber="168">Australia</collectingCountry>
(
<bibRefCitation id="EF994B4E12668205FD31F9AAFF28F9D7" author="Hoese, D. F." pageId="3" pageNumber="168" pagination="1596 - 1611" refId="ref8409" refString="Hoese, D. F. 2006. Eleotridae. Gudgeons, sleepers. Pp. 1596 - 1611. In: Beasley, O. L. and Wells, A. (Eds) Zoological Catalogue of Australia. Volume 35, Part 3 (3). ABRS, Canberra and CSIRO, Collingwood." type="book chapter" year="2006">Hoese 2006</bibRefCitation>
), but the distribution should be revised to avoid misidentification.
<bibRefCitation id="EF994B4E12668205FEBAF9E8FDA0F937" author="Larson, H. K. &amp; Nguyen, T. H." box="[309,600,1662,1687]" pageId="3" pageNumber="168" pagination="1139300" refId="ref8721" refString="Larson, H. K. and Nguyen, T. H. 2021. Odonteleotris macrodon. The IUCN Red List of Threatened Species 2021: e. T 166620 A 1139300. doi: 10.2305 / IUCN. UK. 2021 - 1. RLTS. T 166620 A 1139300. en" type="journal article" year="2021">Larson and Nguyen (2021)</bibRefCitation>
pointed that a disjunct subpopulation from southern New
<collectingCountry id="F31F762F12668205FD0EF908FD33F916" box="[641,715,1695,1719]" name="Guinea-Bissau" pageId="3" pageNumber="168">Guinea</collectingCountry>
and northern
<collectingCountry id="F31F762F12668205FE8EF928FEA5F976" box="[257,349,1727,1751]" name="Australia" pageId="3" pageNumber="168">Australia</collectingCountry>
may be an undescribed species. Therefore, further examination is required to determine the true distribution of
<taxonomicName id="4C084D3C12668205FEBBF896FE4DF8B9" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[308,437,1792,1816]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12668205FEBBF896FE4DF8B9" box="[308,437,1792,1816]" italics="true" pageId="3" pageNumber="168">O. macrodon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736BF12668205FE8FF8D6FDA7F83A" blockId="3.[245,678,1857,1947]" pageId="3" pageNumber="168">
<taxonomicName id="4C084D3C12668205FE8FF8D6FD63F8FB" authority="Roxas and Ablan, 1940" authorityName="Roxas and Ablan" authorityYear="1940" box="[256,667,1857,1882]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12668205FE8FF8D6FE56F8F8" bold="true" box="[256,430,1857,1881]" italics="true" pageId="3" pageNumber="168">Paloa villadolidi</emphasis>
<bibRefCitation id="EF994B4E12668205FE3BF8D5FD63F8FB" author="Roxas, H. A. &amp; Ablan, G. L." box="[436,667,1857,1882]" pageId="3" pageNumber="168" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan, 1940</bibRefCitation>
</taxonomicName>
[New standard Japanese name: Shīsā-haze] (
<figureCitation id="13332A3A12668205FECCF814FE68F83A" box="[323,400,1923,1947]" captionStart="Fig" captionStartId="2.[156,187,1350,1372]" captionTargetBox="[208,1379,610,1309]" captionTargetId="figure-426@2.[206,1381,609,1311]" captionTargetPageId="2" captionText="Fig. 2. Dentition in upper and lower jaws of Odonteleotris macrodon (A: RMNH.PISC. 4472, holotype of Eleotris macrodon, 132.5mm SL) and Paloa villadolidi (B: URM-P 49659, 55.1mm SL). UCT and LCT, magnified illustrations of enlarged canine teeth in the upper and lower jaws, respectively." figureDoi="http://doi.org/10.5281/zenodo.10833164" httpUri="https://zenodo.org/record/10833164/files/figure.png" pageId="3" pageNumber="168">Figs 2B</figureCitation>
,
<figureCitation id="13332A3A12668205FE15F814FE2BF83B" box="[410,467,1923,1947]" captionStart="Fig" captionStartId="3.[156,187,1191,1213]" captionTargetBox="[177,745,206,1151]" captionTargetId="figure-696@3.[176,746,205,1153]" captionTargetPageId="3" captionText="Fig. 3. Illustrations of specimens of Odonteleotris and Paloa. A, Holotype of Eleotris macrodon from Bleeker (1853: pl. 2); B, E. macrodon from Day (1878: pl. 65, fig. 5); C, O. macrodon from Koumans (1953: fig. 81); D, holotype of P. polylepis from Herre (1927: pl. 4, fig. 3); E: holotype of P. villadolidi from Roxas and Ablan (1940: pl. 3)." figureDoi="http://doi.org/10.5281/zenodo.10833166" httpUri="https://zenodo.org/record/10833166/files/figure.png" pageId="3" pageNumber="168">3D, E</figureCitation>
,
<figureCitation id="13332A3A12668205FE52F814FDFCF83A" box="[477,516,1923,1947]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="4.[156,187,1887,1909]" captionStartId-1="6.[156,187,1505,1527]" captionStartId-2="6.[156,187,2056,2078]" captionStartId-3="7.[156,187,764,786]" captionStartId-4="7.[822,853,1391,1413]" captionStartId-5="8.[156,187,906,928]" captionTargetBox-0="[207,1380,224,1751]" captionTargetBox-1="[159,765,479,1467]" captionTargetBox-2="[207,1380,1657,2016]" captionTargetBox-3="[207,1380,206,725]" captionTargetBox-4="[823,1429,901,1351]" captionTargetBox-5="[208,1379,207,866]" captionTargetId-0="figure-75@4.[206,1381,205,1848]" captionTargetId-1="figure-544@6.[158,766,478,1468]" captionTargetId-2="figure-589@6.[206,1381,1656,2017]" captionTargetId-3="figure-554@7.[206,1381,205,726]" captionTargetId-4="figure-508@7.[822,1430,900,1352]" captionTargetId-5="figure-616@8.[206,1381,205,868]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="Fig. 4. Specimens of Paloa villadolidi from Japan (A: URM-P 49659, 55.1mm SL; BE: NSMT-P 143769, 55.4mm SL). AD, Specimens immediately after fixation; E, preserved specimen." captionText-1="Fig. 5. Sensory pores (open circles with uppercase notations) and neuromast rows (red dots with lowercase notations) in the head of Paloa villadolidi (NSMT-P 143769, 55.4mm SL). A, Dorsal, B, lateral, and C, ventral views. AN, anterior nostril; PN, posterior nostril." captionText-2="Fig. 6. Neuromast rows (red dots with lowercase notations) of trunk and caudal fin in Paloa villadolidi (NSMT-P 143769, 55.4mm SL)." captionText-3="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." captionText-4="Fig. 8. Urogenital papilla of Paloa villadolidi. A, Male (NSMT- P 143769, 55.4mm SL); B, female (NSMT-P 145829, 75.0mm SL). Broken lines indicate the outer edge of urogenital papilla; white arrows indicate the anus; red circles indicate the anal-fin origin." captionText-5="Fig. 9. A tide pool in mangrove of Okinawa Island in Japan, where four specimens of Paloa villadolidi were collected." figureDoi-0="http://doi.org/10.5281/zenodo.10833168" figureDoi-1="http://doi.org/10.5281/zenodo.10833170" figureDoi-2="http://doi.org/10.5281/zenodo.10833174" figureDoi-3="http://doi.org/10.5281/zenodo.10833176" figureDoi-4="http://doi.org/10.5281/zenodo.10833178" figureDoi-5="http://doi.org/10.5281/zenodo.10833180" httpUri-0="https://zenodo.org/record/10833168/files/figure.png" httpUri-1="https://zenodo.org/record/10833170/files/figure.png" httpUri-2="https://zenodo.org/record/10833174/files/figure.png" httpUri-3="https://zenodo.org/record/10833176/files/figure.png" httpUri-4="https://zenodo.org/record/10833178/files/figure.png" httpUri-5="https://zenodo.org/record/10833180/files/figure.png" pageId="3" pageNumber="168">49</figureCitation>
;
<tableCitation id="C68A030412668205FD81F814FDADF83A" box="[526,597,1923,1947]" captionStart="Table 1" captionStartId="5.[156,204,190,212]" captionText="Table 1. Meristic and morphometric characters of Odonteleotris macrodon and species of Paloa. Data of P. polylepis and P. villadolidi were compiled from descriptions of Herre (1927) and Roxas and Ablan (1940), respectively. Measurements were expressed as a percentage of standard length (SL). Head lengths were expressed also as a percentage of total length (TL) for comparison. Mode of counts was shown in parentheses." pageId="3" pageNumber="168">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C312653412668205FF13F853FF02F7BC" pageId="3" pageNumber="168" type="reference_group">
<paragraph id="8BB736BF12668205FF13F853FF02F7BC" blockId="3.[156,767,1988,2077]" pageId="3" pageNumber="168">
<treatmentCitationGroup id="AB18119112668205FF13F853FF02F7BC" pageId="3" pageNumber="168">
<treatmentCitation id="0AA910AE12668205FF13F853FD87F87D" author="Roxas, H. A. &amp; Ablan, G. L." box="[156,639,1988,2012]" page="304" pageId="3" pageNumber="168" year="1940">
<taxonomicName id="4C084D3C12668205FF13F853FD87F87D" ID-CoL="6TJQS" authority="Roxas and Ablan, 1940: 304" authorityName="Roxas and Ablan" authorityPageNumber="304" authorityYear="1940" box="[156,639,1988,2012]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12668205FF13F853FEBCF87D" box="[156,324,1988,2012]" italics="true" pageId="3" pageNumber="168">Paloa villadolidi</emphasis>
<bibRefCitation id="EF994B4E12668205FEC0F853FD87F87D" author="Roxas, H. A. &amp; Ablan, G. L." box="[335,639,1988,2012]" pageId="3" pageNumber="168" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan, 1940: 304</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl.
<materialsCitation id="3B603CE212668205FD3CF853FF0EF7BC" country="Philippines" location="Luzon" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Dagupan">
3 (
<typeStatus id="54B3881D12668205FD5EF853FD06F87D" box="[721,766,1988,2012]" pageId="3" pageNumber="168">type</typeStatus>
locality:
<collectingRegion id="49CCF85D12668205FE9EF872FE88F85C" box="[273,368,2021,2045]" country="Philippines" name="Dagupan" pageId="3" pageNumber="168">Dagupan</collectingRegion>
,
<collectingRegion id="49CCF85D12668205FEF0F873FDA1F85D" box="[383,601,2020,2044]" country="Philippines" name="Pangasinan" pageId="3" pageNumber="168">Pangasinan Province</collectingRegion>
,
<location id="8ED7606412668205FDE7F872FD52F85D" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FDE7F872FD52F85D" box="[616,682,2021,2044]" country="Philippines" name="Luzon" pageId="3" pageNumber="168" stateProvince="Dagupan">Luzon</location>
,
<collectingCountry id="F31F762F12668205FD36F873FF17F7BC" name="Philippines" pageId="3" pageNumber="168">Philippines</collectingCountry>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C312653412668205FCC1FF51FAFFFDC4" pageId="3" pageNumber="168" type="materials_examined">
<paragraph id="8BB736BF12668205FCC1FF51FAFFFDC4" blockId="3.[821,1432,198,2076]" pageId="3" pageNumber="168">
<emphasis id="B97CEAAD12668205FCC1FF51FBDBFF7F" bold="true" box="[846,1059,198,222]" pageId="3" pageNumber="168">Material examined.</emphasis>
<materialsCitation id="3B603CE212668205FBB2FF51FAA3FF5F" collectionCode="SL" collectorName="Ryukyu Archipelago" country="Japan" location="Japan" pageId="3" pageNumber="168" specimenCount="7">
<specimenCount id="9D0EFD3612668205FBB2FF51FB09FF7F" box="[1085,1265,198,222]" count="7" pageId="3" pageNumber="168" type="generic">Seven specimens</specimenCount>
(19.875.0 mm
<collectionCode id="ED19AE7A12668205FCBAFF70FCA9FF5E" box="[821,849,231,255]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
) from the
<collectorName id="26FD536912668205FC4BFF70FB6DFF5E" box="[964,1173,231,255]" pageId="3" pageNumber="168">Ryukyu Archipelago</collectorName>
in southern
<collectingCountry id="F31F762F12668205FAAFFF70FAA3FF5F" box="[1312,1371,231,254]" name="Japan" pageId="3" pageNumber="168">Japan</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B603CE212668205FAE8FF70FC27FEE1" collectingDate="2021-10-19" collectionCode="SL" country="Japan" location="Island" pageId="3" pageNumber="168" specimenCode="NSMT-P 143769" specimenCount="1" stateProvince="Okinawa">
<emphasis id="B97CEAAD12668205FAE8FF70FC44FEBE" bold="true" pageId="3" pageNumber="168">
<collectingRegion id="49CCF85D12668205FAE8FF70FC94FEBE" country="Japan" name="Okinawa" pageId="3" pageNumber="168">Okinawa</collectingRegion>
<location id="8ED7606412668205FCFBFE90FC40FEBE" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FCFBFE90FC40FEBE" box="[884,952,263,287]" country="Japan" name="Island" pageId="3" pageNumber="168" stateProvince="Okinawa">Island</location>
:
</emphasis>
<specimenCode id="DBAE9EC412668205FC4AFE90FB8CFEBE" box="[965,1140,263,287]" collectionCode="NSMT-P" pageId="3" pageNumber="168">NSMT-P 143769</specimenCode>
, male,
<quantity id="4CF09B5A12668205FB4EFE90FAE5FEBE" box="[1217,1309,263,287]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="3" pageNumber="168" unit="mm" value="55.6">55.6 mm</quantity>
<collectionCode id="ED19AE7A12668205FAA8FE90FABBFEBE" box="[1319,1347,263,287]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FAC0FE90FC51FEE1" pageId="3" pageNumber="168" value="2021-10-19">
<collectingDate id="EFF2E99712668205FAC0FE90FC51FEE1" pageId="3" pageNumber="168" value="2021-10-19">19 October 2021</collectingDate>
</date>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212668205FC50FEBFFBFCFE9E" box="[991,1028,296,320]" collectingDate="2021-10-19" collectionCode="SL" country="Japan" isTailEnumeration="true" location="Island" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">. H</materialsCitation>
<materialsCitation id="3B603CE212668205FB8BFEBFFB98FEC1" collectingDate="2022-07-09" collectionCode="SL" country="Japan" location="Kobayashi" pageId="3" pageNumber="168" specimenCode="NSMT-P 145828" specimenCount="1" stateProvince="Okinawa">
.
<location id="8ED7606412668205FB9CFEBFFB86FEE1" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FB9CFEBFFB86FEE1" box="[1043,1150,296,320]" country="Japan" name="Kobayashi" pageId="3" pageNumber="168" stateProvince="Okinawa">Kobayashi</location>
;
<specimenCode id="DBAE9EC412668205FB01FEBFFAC7FEE1" box="[1166,1343,296,320]" collectionCode="NSMT-P" pageId="3" pageNumber="168">NSMT-P 145828</specimenCode>
, female,
<quantity id="4CF09B5A12668205FCBAFEDFFC77FEC1" box="[821,911,328,352]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" pageId="3" pageNumber="168" unit="mm" value="53.5">53.5 mm</quantity>
<collectionCode id="ED19AE7A12668205FC19FEDFFC4AFEC1" box="[918,946,328,352]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FC33FEDFFBD7FEC1" box="[956,1071,328,352]" pageId="3" pageNumber="168" value="2022-07-09">
<collectingDate id="EFF2E99712668205FC33FEDFFBD7FEC1" box="[956,1071,328,352]" pageId="3" pageNumber="168" value="2022-07-09">9 July 2022</collectingDate>
</date>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212668205FBEFFEDFFB78FEC1" box="[1120,1152,328,352]" collectingDate="2022-07-09" collectionCode="SL" country="Japan" isTailEnumeration="true" location="Kobayashi" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">. H</materialsCitation>
<materialsCitation id="3B603CE212668205FB0FFEDEFAC5FEC1" box="[1152,1341,328,352]" country="Japan" location="Kobayashi" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">
.
<location id="8ED7606412668205FB05FEDFFB0BFEC1" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FB05FEDFFB0BFEC1" box="[1162,1267,328,352]" country="Japan" name="Kobayashi" pageId="3" pageNumber="168" stateProvince="Okinawa">Kobayashi</location>
and H
</materialsCitation>
<materialsCitation id="3B603CE212668205FAB3FEDEFB4DFE00" collectingDate="2022-07-13" collectionCode="SL" country="Japan" location="Miyake" pageId="3" pageNumber="168" specimenCode="NSMT-P 145829, NSMT-P 145830" specimenCount="1" stateProvince="Okinawa">
.
<location id="8ED7606412668205FAC8FEDFFA6CFEC1" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FAC8FEDFFA6CFEC1" box="[1351,1428,328,352]" country="Japan" name="Miyake" pageId="3" pageNumber="168" stateProvince="Okinawa">Miyake</location>
;
<specimenCode id="DBAE9EC412668205FCBAFEFEFC19FE20" box="[821,993,361,385]" collectionCode="NSMT-P" pageId="3" pageNumber="168">NSMT-P 145829</specimenCode>
, female, 75.0 mm
<collectionCode id="ED19AE7A12668205FB11FEFEFB42FE20" box="[1182,1210,361,385]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<specimenCode id="DBAE9EC412668205FB4AFEFEFA89FE20" box="[1221,1393,361,385]" collectionCode="NSMT-P" pageId="3" pageNumber="168">NSMT-P 145830</specimenCode>
, female, 41.0 mm
<collectionCode id="ED19AE7A12668205FC57FE1DFC0CFE03" box="[984,1012,394,418]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FC70FE1DFB7AFE03" box="[1023,1154,393,418]" pageId="3" pageNumber="168" value="2022-07-13">
<collectingDate id="EFF2E99712668205FC70FE1DFB7AFE03" box="[1023,1154,393,418]" pageId="3" pageNumber="168" value="2022-07-13">13 July 2022</collectingDate>
</date>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212668205FB3AFE1EFB2EFE00" box="[1205,1238,393,417]" collectingDate="2022-07-13" collectionCode="SL" country="Japan" isTailEnumeration="true" location="Miyake" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">. H</materialsCitation>
<materialsCitation id="3B603CE212668205FB59FE1DFA6DFE00" box="[1238,1429,393,417]" collectionCode="R" country="Japan" location="Kobayashi" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">
.
<location id="8ED7606412668205FB6DFE1EFAB3FE00" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FB6DFE1EFAB3FE00" box="[1250,1355,393,417]" country="Japan" name="Kobayashi" pageId="3" pageNumber="168" stateProvince="Okinawa">Kobayashi</location>
and
<collectionCode id="ED19AE7A12668205FA0DFE1DFA6DFE00" box="[1410,1429,394,417]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="168">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B603CE212668205FCBAFE3DFCA4FE43" collectingDate="2019-10-16" collectionCode="SL" country="Japan" location="Ueda" pageId="3" pageNumber="168" specimenCode="URM-P 49659" specimenCount="1" stateProvince="Okinawa">
<location id="8ED7606412668205FCBAFE3DFC94FE63" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FCBAFE3DFC94FE63" box="[821,876,426,450]" country="Japan" name="Ueda" pageId="3" pageNumber="168" stateProvince="Okinawa">Ueda</location>
;
<specimenCode id="DBAE9EC412668205FCF7FE3DFBE8FE63" box="[888,1040,426,450]" collectionCode="URM-P" pageId="3" pageNumber="168">URM-P 49659</specimenCode>
, male,
<quantity id="4CF09B5A12668205FBD3FE3DFB4AFE60" box="[1116,1202,426,450]" metricMagnitude="-2" metricUnit="m" metricValue="5.51" pageId="3" pageNumber="168" unit="mm" value="55.1">55.1 mm</quantity>
<collectionCode id="ED19AE7A12668205FB33FE3DFB20FE63" box="[1212,1240,426,450]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FB6BFE3DFA6CFE63" box="[1252,1428,426,450]" pageId="3" pageNumber="168" value="2019-10-16">
<collectingDate id="EFF2E99712668205FB6BFE3DFA6CFE63" box="[1252,1428,426,450]" pageId="3" pageNumber="168" value="2019-10-16">16 October 2019</collectingDate>
</date>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212668205FCD4FE5DFC85FE43" box="[859,893,458,482]" collectingDate="2019-10-16" collectionCode="SL" country="Japan" isTailEnumeration="true" location="Ueda" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">. K</materialsCitation>
<materialsCitation id="3B603CE212668205FCF3FE5CFC10FDA2" collectingDate="2014-08-28" collectionCode="SL" country="Japan" location="Nishigaki" pageId="3" pageNumber="168" specimenCode="URM-P 49660" specimenCount="1" stateProvince="Okinawa">
.
<location id="8ED7606412668205FC05FE5DFC15FE43" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FC05FE5DFC15FE43" box="[906,1005,458,482]" country="Japan" name="Nishigaki" pageId="3" pageNumber="168" stateProvince="Okinawa">Nishigaki</location>
;
<specimenCode id="DBAE9EC412668205FC74FE5CFB6CFE43" box="[1019,1172,458,482]" collectionCode="URM-P" pageId="3" pageNumber="168">URM-P 49660</specimenCode>
, female,
<quantity id="4CF09B5A12668205FB7AFE5CFAB3FE43" box="[1269,1355,459,483]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="3" pageNumber="168" unit="mm" value="31.1">31.1 mm</quantity>
<collectionCode id="ED19AE7A12668205FAD9FE5CFA8AFE42" box="[1366,1394,459,483]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FAF0FE5CFC4FFDA2" pageId="3" pageNumber="168" value="2014-08-28">
<collectingDate id="EFF2E99712668205FAF0FE5CFC4FFDA2" pageId="3" pageNumber="168" value="2014-08-28">28 August 2014</collectingDate>
</date>
, coll
</materialsCitation>
<materialsCitation id="3B603CE212668205FC67FE7CFBF0FDA2" box="[1000,1032,491,515]" collectingDate="2014-08-28" collectionCode="SL" country="Japan" isTailEnumeration="true" location="Nishigaki" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">. H</materialsCitation>
. Yagi (used only for fin-ray counts and observations of teeth, cephalic sensory systems, and coloration in life).
<materialsCitation id="3B603CE212668205FC6BFDBBFB51FDC4" collectingDate="2010-09-28" collectionCode="SL" country="Japan" location="Ishigaki Island" pageId="3" pageNumber="168" specimenCode="URM-P 49661" specimenCount="1" stateProvince="Okinawa">
<emphasis id="B97CEAAD12668205FC6BFDBBFB74FDE5" bold="true" box="[996,1164,556,580]" pageId="3" pageNumber="168">
<location id="8ED7606412668205FC6BFDBBFB70FDE5" LSID="urn:lsid:plazi:treatment:03A187A91264820EFE8EF834FDA3FAD0:8ED7606412668205FC6BFDBBFB70FDE5" box="[996,1160,556,580]" country="Japan" name="Ishigaki Island" pageId="3" pageNumber="168" stateProvince="Okinawa">Ishigaki Island</location>
:
</emphasis>
<specimenCode id="DBAE9EC412668205FB17FDBAFACBFDE5" box="[1176,1331,556,580]" collectionCode="URM-P" pageId="3" pageNumber="168">URM-P 49661</specimenCode>
, juvenile,
<quantity id="4CF09B5A12668205FCBAFDDAFC73FDC5" box="[821,907,589,613]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="3" pageNumber="168" unit="mm" value="19.8">19.8 mm</quantity>
<collectionCode id="ED19AE7A12668205FC1EFDDAFC55FDC4" box="[913,941,589,613]" country="Sierra Leone" lsid="urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="168" type="Herbarium">SL</collectionCode>
,
<date id="FFB6107F12668205FC39FDDAFB81FDC4" box="[950,1145,589,613]" pageId="3" pageNumber="168" value="2010-09-28">
<collectingDate id="EFF2E99712668205FC39FDDAFB81FDC4" box="[950,1145,589,613]" pageId="3" pageNumber="168" value="2010-09-28">28 September 2010</collectingDate>
</date>
, coll
</materialsCitation>
.
<materialsCitation id="3B603CE212668205FB3DFDDAFAFBFDC4" box="[1202,1283,589,613]" collectionCode="T" collectorName="Saeki" country="Japan" pageId="3" pageNumber="168" specimenCount="1" stateProvince="Okinawa">
<collectionCode id="ED19AE7A12668205FB3DFDDAFB3AFDC5" box="[1202,1218,589,612]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="168">T</collectionCode>
.
<collectorName id="26FD536912668205FB44FDDAFAFBFDC4" box="[1227,1283,589,613]" pageId="3" pageNumber="168">Saeki</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312653412668205FCC1FDFAFC27FBCC" pageId="3" pageNumber="168" type="diagnosis">
<paragraph id="8BB736BF12668205FCC1FDFAFC27FBCC" blockId="3.[821,1432,198,2076]" pageId="3" pageNumber="168">
<emphasis id="B97CEAAD12668205FCC1FDFAFC46FD24" bold="true" box="[846,958,621,645]" pageId="3" pageNumber="168">Diagnosis.</emphasis>
<taxonomicName id="4C084D3C12668205FC58FDFAFB79FD24" authorityName="Roxas and Ablan" authorityYear="1940" box="[983,1153,621,645]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12668205FC58FDFAFB79FD24" box="[983,1153,621,645]" italics="true" pageId="3" pageNumber="168">Paloa villadolidi</emphasis>
</taxonomicName>
is distinguished from all other species of
<taxonomicName id="4C084D3C12668205FC50FD19FBD5FD07" authorityName="Bleeker" authorityYear="1874" box="[991,1069,654,678]" class="Actinopteri" family="Butidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiiformes" pageId="3" pageNumber="168" phylum="Chordata" rank="family">Butidae</taxonomicName>
by a combination of the following characters. Upper and lower jaws with enlarged canine teeth on their outermost rows; more canines on upper jaw (711) than on lower jaw (47); no inner row of canines in the posterior half of lower jaw. Upper jaw not extending to anterior margin of eye. Interorbital sensory pore D paired with wide interval. Fourth transverse row of sensory papillae (
<emphasis id="B97CEAAD12668205FA0CFCC6FA77FCC8" box="[1411,1423,849,873]" italics="true" pageId="3" pageNumber="168">4</emphasis>
) crossed with longitudinal papilla row (
<emphasis id="B97CEAAD12668205FB32FCE6FB31FC28" box="[1213,1225,881,905]" italics="true" pageId="3" pageNumber="168">b</emphasis>
) on the cheek. Longitudinal scales 88100, transverse scales 3236, predorsal scales 5165, transverse scales at caudal peduncle 22 or 23, circumpeduncle scales 4043. Dorsal fin VI-I, 910; anal fin I, 8; pectoral fin 17. Predorsal length 38.440.7% of SL; body depth at anal-fin origin 14.617.1% of SL; eye diameter 3.3 5.1% of SL. No distinct markings on the unpaired fins and caudal peduncle.
</paragraph>
</subSubSection>
<subSubSection id="C312653412668201FCC1FBE2FE92F87D" lastPageId="7" lastPageNumber="172" pageId="3" pageNumber="168" type="description">
<paragraph id="8BB736BF12668205FCC1FBE2FC97F81B" blockId="3.[821,1432,198,2076]" pageId="3" pageNumber="168">
<emphasis id="B97CEAAD12668205FCC1FBE2FC2AFB2C" bold="true" box="[846,978,1141,1165]" pageId="3" pageNumber="168">Description.</emphasis>
The morphometric and meristic characters are presented in
<tableCitation id="C68A030412668205FB9FFB01FBA4FB0F" box="[1040,1116,1174,1198]" captionStart="Table 1" captionStartId="5.[156,204,190,212]" captionText="Table 1. Meristic and morphometric characters of Odonteleotris macrodon and species of Paloa. Data of P. polylepis and P. villadolidi were compiled from descriptions of Herre (1927) and Roxas and Ablan (1940), respectively. Measurements were expressed as a percentage of standard length (SL). Head lengths were expressed also as a percentage of total length (TL) for comparison. Mode of counts was shown in parentheses." pageId="3" pageNumber="168">Table 1</tableCitation>
. Body elongated and cylindrical anteriorly, somewhat compressed posteriorly (
<figureCitation id="13332A3A12668205FAC6FB20FA74FB6E" box="[1353,1420,1207,1231]" captionStart="Fig" captionStartId="4.[156,187,1887,1909]" captionTargetBox="[207,1380,224,1751]" captionTargetId="figure-75@4.[206,1381,205,1848]" captionTargetPageId="4" captionText="Fig. 4. Specimens of Paloa villadolidi from Japan (A: URM-P 49659, 55.1mm SL; BE: NSMT-P 143769, 55.4mm SL). AD, Specimens immediately after fixation; E, preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.10833168" httpUri="https://zenodo.org/record/10833168/files/figure.png" pageId="3" pageNumber="168">Fig. 4</figureCitation>
). Head slightly depressed, dorsal profile nearly straight, no bony crests in the interorbital region. Mouth large, superior, and oblique with an approximately 50º upward angle; its end not extending below the anterior margin of the eye. Snout rounded, length longer than eye diameter. Anterior nostrils tubular and extending to anterior margin of upper lip; posterior nostrils oval with low rim and not tubular, located approximately one-third of the distance between the orbit and anterior nostril. The gill opening extends anteriorly to below the posterior margin of the preopercle. Upper and lower jaws with enlarged and curved canines in their outermost tooth row (
<figureCitation id="13332A3A12668205FC51F9AAFBD2F9F4" box="[990,1066,1597,1621]" captionStart="Fig" captionStartId="2.[156,187,1350,1372]" captionTargetBox="[208,1379,610,1309]" captionTargetId="figure-426@2.[206,1381,609,1311]" captionTargetPageId="2" captionText="Fig. 2. Dentition in upper and lower jaws of Odonteleotris macrodon (A: RMNH.PISC. 4472, holotype of Eleotris macrodon, 132.5mm SL) and Paloa villadolidi (B: URM-P 49659, 55.1mm SL). UCT and LCT, magnified illustrations of enlarged canine teeth in the upper and lower jaws, respectively." figureDoi="http://doi.org/10.5281/zenodo.10833164" httpUri="https://zenodo.org/record/10833164/files/figure.png" pageId="3" pageNumber="168">Fig. 2B</figureCitation>
): 711 canines in the upper jaw and
<quantity id="4CF09B5A12668205FCBAF9C9FC8EF9D4" box="[821,886,1629,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.778" metricValueMin="1.016" pageId="3" pageNumber="168" unit="in" value="5.5" valueMax="7.0" valueMin="4.0">47 in</quantity>
the lower jaw; upper jaw always having more canines than lower jaw. Two to four rows of small conical teeth on the upper and lower jaws. No teeth on the vomer. Small and somewhat deciduous cycloid scales covering the head and body, except dorsal surface of head anterior to interorbital pores D, side of snout in front of suborbital papilla row
<emphasis id="B97CEAAD12668205FA09F897FA6AF8B9" box="[1414,1426,1792,1816]" italics="true" pageId="3" pageNumber="168">1</emphasis>
, lower jaw, lips, maxilla, gular region, and gill membrane; embedded scales on the cheek; no accessory scales. Longitudinal scales 88100, transverse scales 3236, predorsal scales 5165. Urogenital papilla small and triangular in both sexes.
</paragraph>
<paragraph id="8BB736BF12668200FCC1F854FC4DFEE1" blockId="3.[821,1432,198,2076]" lastBlockId="6.[820,1432,198,1589]" lastPageId="6" lastPageNumber="171" pageId="3" pageNumber="168">First dorsal fin with six spines; three anterior spines slightly longer than posterior ones. First and second dorsal fins separated by an interval of approximately eight scales along the dorsal midline. Second dorsal fin with one spine and nine or ten soft rays; length of the spine being the same as those of first to third spines of the first dorsal fin; all soft rays branched and longer than spine; soft rays becoming longer from first to fifth soft rays, similar length in fifth to eighth rays, and ninth ray shorter than eighth. Anal fin with one spine and eight soft rays; length of the spine almost the same as that of the second-dorsal-fin spine; all soft rays branched. Anal-fin origin vertically below the base of second or third soft ray of the second dorsal fin. The posterior tip of the second dorsal and anal fins not reaching the caudal-fin base. Caudal fin rounded with 17 segmented rays, including 1417 branched rays. Pectoral fin rounded with 17 soft rays; rays not filamentous, tip of the pectoral fin below the base of the fourth or fifth spine of first dorsal fin. Pelvic-fin insertion slightly anterior to the pectoral-fin base. Left and right pelvic fins separated from each other, both with one spine and five soft rays; pointed distal tip of the pelvic fin not reaching middle of interval between pelvic- and anal-fin bases; third soft ray longest. Number of vertebrae 26, including 11 abdominal and 15 caudal vertebrae. P-V 3/ II I I 0/9: three anal-fin pterygiophores forward to the first hemal spine.</paragraph>
<caption id="DF77663712618202FF13F8C8FD99F830" ID-DOI="http://doi.org/10.5281/zenodo.10833168" ID-Zenodo-Dep="10833168" httpUri="https://zenodo.org/record/10833168/files/figure.png" pageId="4" pageNumber="169" startId="4.[156,187,1887,1909]" targetBox="[207,1380,224,1751]" targetPageId="4" targetType="figure">
<paragraph id="8BB736BF12618202FF13F8C8FD99F830" blockId="4.[156,1432,1887,1937]" pageId="4" pageNumber="169">
Fig. 4. Specimens of
<taxonomicName id="4C084D3C12618202FEFEF8C8FDFCF8D5" authorityName="Roxas and Ablan" authorityYear="1940" box="[369,516,1887,1908]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="169" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12618202FEFEF8C8FDFCF8D5" box="[369,516,1887,1908]" italics="true" pageId="4" pageNumber="169">Paloa villadolidi</emphasis>
</taxonomicName>
from Japan (A: URM-P 49659, 55.1 mm SL; BE: NSMT-P 143769, 55.4 mm SL). AD, Specimens immediately after fixation; E, preserved specimen.
</paragraph>
</caption>
<caption id="DF77663712608203FF13FF29FEF5FE89" pageId="5" pageNumber="170" startId="5.[156,204,190,212]" targetType="table">
<paragraph id="8BB736BF12608203FF13FF29FEF5FE89" blockId="5.[156,1431,190,296]" pageId="5" pageNumber="170">
Table 1. Meristic and morphometric characters of
<taxonomicName id="4C084D3C12608203FDF2FF29FCADFF72" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[637,853,190,211]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12608203FDF2FF29FCADFF72" box="[637,853,190,211]" italics="true" pageId="5" pageNumber="170">Odonteleotris macrodon</emphasis>
</taxonomicName>
and species of
<taxonomicName id="4C084D3C12608203FC6EFF29FBEBFF72" authorityName="Herre" authorityYear="1927" box="[993,1043,190,211]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="genus">
<emphasis id="B97CEAAD12608203FC6EFF29FBEBFF72" box="[993,1043,190,211]" italics="true" pageId="5" pageNumber="170">Paloa</emphasis>
</taxonomicName>
. Data of
<taxonomicName id="4C084D3C12608203FBE7FF28FB31FF72" authorityName="Herre" authorityYear="1927" box="[1128,1225,190,212]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="B97CEAAD12608203FBE7FF28FB31FF72" box="[1128,1225,190,212]" italics="true" pageId="5" pageNumber="170">P. polylepis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D3C12608203FB78FF28FA9DFF72" authorityName="Roxas and Ablan" authorityYear="1940" box="[1271,1381,190,212]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12608203FB78FF28FA9DFF72" box="[1271,1381,190,212]" italics="true" pageId="5" pageNumber="170">P. villadolidi</emphasis>
</taxonomicName>
were compiled from descriptions of
<bibRefCitation id="EF994B4E12608203FE31FF4CFDCEFF4E" author="Herre, A. W. C. T." box="[446,566,218,240]" pageId="5" pageNumber="170" pagination="1 - 352" refId="ref8367" refString="Herre, A. W. C. T. 1927. Gobies of the Philippines and the China Sea. Monographs of the Bureau of Science Manila, Philippine Islands 23: 1 - 352, pls 1 - 30." type="journal article" year="1927">Herre (1927)</bibRefCitation>
and
<bibRefCitation id="EF994B4E12608203FDE6FF4CFCB5FF4E" author="Roxas, H. A. &amp; Ablan, G. L." box="[617,845,218,240]" pageId="5" pageNumber="170" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan (1940)</bibRefCitation>
, respectively. Measurements were expressed as a percentage of standard length (SL). Head lengths were expressed also as a percentage of total length (TL) for comparison. Mode of counts was shown in parentheses.
</paragraph>
</caption>
<paragraph id="8BB736BF12608203FF28FED1FAA8F892" pageId="5" pageNumber="170">
<table id="F908C41F12607DF9FF28FED1FA7EF892" box="[167,1414,326,1843]" gridcols="5" gridrows="47" pageId="5" pageNumber="170">
<tr id="353834FD12607DF9FF28FED1FA7EFEF8" box="[167,1414,326,345]" gridrow="0" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FED1FE70FEF8" box="[167,392,326,345]" gridcol="0" gridrow="0" pageId="5" pageNumber="170">Species</th>
<th id="76E95D8112607DF9FE3BFED1FD54FEF8" box="[436,684,326,345]" gridcol="1" gridrow="0" pageId="5" pageNumber="170">
<taxonomicName id="4C084D3C12608203FE44FED1FD6CFEF8" baseAuthorityName="Bleeker" baseAuthorityYear="1853" box="[459,660,326,345]" family="Eleotridae" genus="Odonteleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="macrodon">
<emphasis id="B97CEAAD12608203FE44FED1FD6CFEF8" box="[459,660,326,345]" italics="true" pageId="5" pageNumber="170">Odonteleotris macrodon</emphasis>
</taxonomicName>
</th>
<th id="76E95D8112607DF9FD35FED1FC38FEF8" box="[698,960,326,345]" gridcol="2" gridrow="0" pageId="5" pageNumber="170">
<taxonomicName id="4C084D3C12608203FC8FFED1FC83FEF8" authorityName="Herre" authorityYear="1927" box="[768,891,326,345]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="B97CEAAD12608203FC8FFED1FC83FEF8" box="[768,891,326,345]" italics="true" pageId="5" pageNumber="170">Paloa polylepis</emphasis>
</taxonomicName>
</th>
<th id="76E95D8112607DF9FC42FED1FA7EFEF8" box="[973,1414,326,345]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="5" pageNumber="170">
<taxonomicName id="4C084D3C12608203FBE5FED1FB0AFEF8" authority="Locality" authorityName="Locality" box="[1130,1266,326,345]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12608203FBE5FED1FB0AFEF8" box="[1130,1266,326,345]" italics="true" pageId="5" pageNumber="170">Paloa villadolidi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353834FD12607DF9FF28FEF8FA7EFE22" box="[167,1414,367,387]" gridrow="1" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FEF8FE70FE22" box="[167,392,367,387]" gridcol="0" gridrow="1" pageId="5" pageNumber="170">Locality</th>
<td id="76E95D8112607DF9FE3BFEF8FD54FE22" box="[436,684,367,387]" gridcol="1" gridrow="1" pageId="5" pageNumber="170">India</td>
<td id="76E95D8112607DF9FD35FEF8FC38FE22" box="[698,960,367,387]" gridcol="2" gridrow="1" pageId="5" pageNumber="170">Philippines</td>
<td id="76E95D8112607DF9FC42FEF8FB4DFE22" box="[973,1205,367,387]" gridcol="3" gridrow="1" pageId="5" pageNumber="170">Philippines</td>
<td id="76E95D8112607DF9FB43FEF8FA7EFE22" box="[1228,1414,367,387]" gridcol="4" gridrow="1" pageId="5" pageNumber="170">Japan</td>
</tr>
<tr id="353834FD12607DF9FF28FE0EFA7EFE0D" box="[167,1414,409,428]" gridrow="2" pageId="5" pageNumber="170" rowspan-0="1">
<td id="76E95D8112607DF9FE3BFE0EFD54FE0D" box="[436,684,409,428]" gridcol="1" gridrow="2" pageId="5" pageNumber="170">Kolkata</td>
<td id="76E95D8112607DF9FD35FE0EFC38FE0D" box="[698,960,409,428]" gridcol="2" gridrow="2" pageId="5" pageNumber="170">Panay</td>
<td id="76E95D8112607DF9FC42FE0EFB4DFE0D" box="[973,1205,409,428]" gridcol="3" gridrow="2" pageId="5" pageNumber="170">Luzon</td>
<td id="76E95D8112607DF9FB43FE0EFA7EFE0D" box="[1228,1414,409,428]" gridcol="4" gridrow="2" pageId="5" pageNumber="170">Okinawa and Ishigaki</td>
</tr>
<tr id="353834FD12607DF9FF28FE54FA7EFE77" box="[167,1414,451,470]" gridrow="3" pageId="5" pageNumber="170" rowspan-0="1">
<td id="76E95D8112607DF9FE3BFE54FD54FE77" box="[436,684,451,470]" gridcol="1" gridrow="3" pageId="5" pageNumber="170">Holotype RMNH.PISC. 4472</td>
<td id="76E95D8112607DF9FD35FE54FC38FE77" box="[698,960,451,470]" gridcol="2" gridrow="3" pageId="5" pageNumber="170">Holotype BSM specimen (lost)</td>
<td id="76E95D8112607DF9FC42FE54FB4DFE77" box="[973,1205,451,470]" gridcol="3" gridrow="3" pageId="5" pageNumber="170">Holotype BSM 31946 (lost)</td>
<td id="76E95D8112607DF9FB43FE54FA7EFE77" box="[1228,1414,451,470]" gridcol="4" gridrow="3" pageId="5" pageNumber="170">
7 specimens
<emphasis id="B97CEAAD12608203FAD7FE54FA9BFE77" bold="true" box="[1368,1379,451,470]" pageId="5" pageNumber="170">*</emphasis>
</td>
</tr>
<tr id="353834FD12607DF9FF28FE7BFA7EFE5E" box="[167,1414,492,511]" gridrow="4" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FE7BFE70FE5E" box="[167,392,492,511]" gridcol="0" gridrow="4" pageId="5" pageNumber="170">Standard length (mm)</th>
<td id="76E95D8112607DF9FE3BFE7BFD54FE5E" box="[436,684,492,511]" gridcol="1" gridrow="4" pageId="5" pageNumber="170">110.6</td>
<td id="76E95D8112607DF9FD35FE7BFC38FE5E" box="[698,960,492,511]" gridcol="2" gridrow="4" pageId="5" pageNumber="170">68</td>
<td id="76E95D8112607DF9FC42FE7BFB4DFE5E" box="[973,1205,492,511]" gridcol="3" gridrow="4" pageId="5" pageNumber="170">56</td>
<td id="76E95D8112607DF9FB43FE7BFA7EFE5E" box="[1228,1414,492,511]" gridcol="4" gridrow="4" pageId="5" pageNumber="170">19.875.0</td>
</tr>
<tr id="353834FD12607DF9FF28FD81FA7EFD88" box="[167,1414,534,553]" gridrow="5" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FD81FA7EFD88" box="[167,1414,534,553]" colspan="5" colspanRight="4" gridcol="0" gridrow="5" pageId="5" pageNumber="170">
<emphasis id="B97CEAAD12608203FF28FD81FF10FD88" bold="true" box="[167,232,534,553]" pageId="5" pageNumber="170">Counts</emphasis>
</th>
</tr>
<tr id="353834FD12607DF9FF28FDA2FA7EFDE9" box="[167,1414,565,584]" gridrow="6" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FDA2FE70FDE9" box="[167,392,565,584]" gridcol="0" gridrow="6" pageId="5" pageNumber="170">D1-D2 elements</th>
<td id="76E95D8112607DF9FE3BFDA2FD54FDE9" box="[436,684,565,584]" gridcol="1" gridrow="6" pageId="5" pageNumber="170">VI-I, 10</td>
<td id="76E95D8112607DF9FD35FDA2FC38FDE9" box="[698,960,565,584]" gridcol="2" gridrow="6" pageId="5" pageNumber="170">V-I, 10</td>
<td id="76E95D8112607DF9FC42FDA2FB4DFDE9" box="[973,1205,565,584]" gridcol="3" gridrow="6" pageId="5" pageNumber="170">VI-I, 9</td>
<td id="76E95D8112607DF9FB43FDA2FA7EFDE9" box="[1228,1414,565,584]" gridcol="4" gridrow="6" pageId="5" pageNumber="170">VI-I, 9 (6) or 10 (1)</td>
</tr>
<tr id="353834FD12607DF9FF28FDC3FA7EFDC6" box="[167,1414,596,615]" gridrow="7" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FDC3FE70FDC6" box="[167,392,596,615]" gridcol="0" gridrow="7" pageId="5" pageNumber="170">A elements</th>
<td id="76E95D8112607DF9FE3BFDC3FD54FDC6" box="[436,684,596,615]" gridcol="1" gridrow="7" pageId="5" pageNumber="170">I, 8</td>
<td id="76E95D8112607DF9FD35FDC3FC38FDC6" box="[698,960,596,615]" gridcol="2" gridrow="7" pageId="5" pageNumber="170">I, 8</td>
<td id="76E95D8112607DF9FC42FDC3FB4DFDC6" box="[973,1205,596,615]" gridcol="3" gridrow="7" pageId="5" pageNumber="170">I, 8</td>
<td id="76E95D8112607DF9FB43FDC3FA7EFDC6" box="[1228,1414,596,615]" gridcol="4" gridrow="7" pageId="5" pageNumber="170">I, 8</td>
</tr>
<tr id="353834FD12607DF9FF28FDE4FA7EFD26" box="[167,1414,627,647]" gridrow="8" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FDE4FE70FD26" box="[167,392,627,647]" gridcol="0" gridrow="8" pageId="5" pageNumber="170">P1 rays</th>
<td id="76E95D8112607DF9FE3BFDE4FD54FD26" box="[436,684,627,647]" gridcol="1" gridrow="8" pageId="5" pageNumber="170">17 (left), 17 (right)</td>
<td id="76E95D8112607DF9FD35FDE4FC38FD26" box="[698,960,627,647]" gridcol="2" gridrow="8" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FDE4FB4DFD26" box="[973,1205,627,647]" gridcol="3" gridrow="8" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FDE4FA7EFD26" box="[1228,1414,627,647]" gridcol="4" gridrow="8" pageId="5" pageNumber="170">17 (5) or 18 (2)</td>
</tr>
<tr id="353834FD12607DF9FF28FD06FA7EFD07" box="[167,1414,657,678]" gridrow="9" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FD06FE70FD07" box="[167,392,657,678]" gridcol="0" gridrow="9" pageId="5" pageNumber="170">P2 rays</th>
<td id="76E95D8112607DF9FE3BFD06FD54FD07" box="[436,684,657,678]" gridcol="1" gridrow="9" pageId="5" pageNumber="170">I, 5</td>
<td id="76E95D8112607DF9FD35FD06FC38FD07" box="[698,960,657,678]" gridcol="2" gridrow="9" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FD06FB4DFD07" box="[973,1205,657,678]" gridcol="3" gridrow="9" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FD06FA7EFD07" box="[1228,1414,657,678]" gridcol="4" gridrow="9" pageId="5" pageNumber="170">I, 5</td>
</tr>
<tr id="353834FD12607DF9FF28FD26FA7EFD65" box="[167,1414,689,708]" gridrow="10" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FD26FE70FD65" box="[167,392,689,708]" gridcol="0" gridrow="10" pageId="5" pageNumber="170">Branched rays in C</th>
<td id="76E95D8112607DF9FE3BFD26FD54FD65" box="[436,684,689,708]" gridcol="1" gridrow="10" pageId="5" pageNumber="170">8 +?</td>
<td id="76E95D8112607DF9FD35FD26FC38FD65" box="[698,960,689,708]" gridcol="2" gridrow="10" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FD26FB4DFD65" box="[973,1205,689,708]" gridcol="3" gridrow="10" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FD26FA7EFD65" box="[1228,1414,689,708]" gridcol="4" gridrow="10" pageId="5" pageNumber="170">8 + 8</td>
</tr>
<tr id="353834FD12607DF9FF28FD47FA7EFD42" box="[167,1414,720,739]" gridrow="11" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FD47FE70FD42" box="[167,392,720,739]" gridcol="0" gridrow="11" pageId="5" pageNumber="170">Segmented rays in C</th>
<td id="76E95D8112607DF9FE3BFD47FD54FD42" box="[436,684,720,739]" gridcol="1" gridrow="11" pageId="5" pageNumber="170">9 + 8</td>
<td id="76E95D8112607DF9FD35FD47FC38FD42" box="[698,960,720,739]" gridcol="2" gridrow="11" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FD47FB4DFD42" box="[973,1205,720,739]" gridcol="3" gridrow="11" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FD47FA7EFD42" box="[1228,1414,720,739]" gridcol="4" gridrow="11" pageId="5" pageNumber="170">9 + 8</td>
</tr>
<tr id="353834FD12607DF9FF28FD78FA7EFCA3" box="[167,1414,751,770]" gridrow="12" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FD78FE70FCA3" box="[167,392,751,770]" gridcol="0" gridrow="12" pageId="5" pageNumber="170">Predorsal scales</th>
<td id="76E95D8112607DF9FE3BFD78FD54FCA3" box="[436,684,751,770]" gridcol="1" gridrow="12" pageId="5" pageNumber="170">63</td>
<td id="76E95D8112607DF9FD35FD78FC38FCA3" box="[698,960,751,770]" gridcol="2" gridrow="12" pageId="5" pageNumber="170">58</td>
<td id="76E95D8112607DF9FC42FD78FB4DFCA3" box="[973,1205,751,770]" gridcol="3" gridrow="12" pageId="5" pageNumber="170">62</td>
<td id="76E95D8112607DF9FB43FD78FA7EFCA3" box="[1228,1414,751,770]" gridcol="4" gridrow="12" pageId="5" pageNumber="170">5165</td>
</tr>
<tr id="353834FD12607DF9FF28FC99FA7EFC80" box="[167,1414,782,801]" gridrow="13" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FC99FE70FC80" box="[167,392,782,801]" gridcol="0" gridrow="13" pageId="5" pageNumber="170">Longitudinal scales</th>
<td id="76E95D8112607DF9FE3BFC99FD54FC80" box="[436,684,782,801]" gridcol="1" gridrow="13" pageId="5" pageNumber="170">103 (left), 98 (right)</td>
<td id="76E95D8112607DF9FD35FC99FC38FC80" box="[698,960,782,801]" gridcol="2" gridrow="13" pageId="5" pageNumber="170">88</td>
<td id="76E95D8112607DF9FC42FC99FB4DFC80" box="[973,1205,782,801]" gridcol="3" gridrow="13" pageId="5" pageNumber="170">100</td>
<td id="76E95D8112607DF9FB43FC99FA7EFC80" box="[1228,1414,782,801]" gridcol="4" gridrow="13" pageId="5" pageNumber="170">8897 (94)</td>
</tr>
<tr id="353834FD12607DF9FF28FCBAFA7EFCE1" box="[167,1414,813,832]" gridrow="14" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FCBAFE70FCE1" box="[167,392,813,832]" gridcol="0" gridrow="14" pageId="5" pageNumber="170">Transverse scales</th>
<td id="76E95D8112607DF9FE3BFCBAFD54FCE1" box="[436,684,813,832]" gridcol="1" gridrow="14" pageId="5" pageNumber="170">32</td>
<td id="76E95D8112607DF9FD35FCBAFC38FCE1" box="[698,960,813,832]" gridcol="2" gridrow="14" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FCBAFB4DFCE1" box="[973,1205,813,832]" gridcol="3" gridrow="14" pageId="5" pageNumber="170">36</td>
<td id="76E95D8112607DF9FB43FCBAFA7EFCE1" box="[1228,1414,813,832]" gridcol="4" gridrow="14" pageId="5" pageNumber="170">3236 (36)</td>
</tr>
<tr id="353834FD12607DF9FF28FCDBFA7EFCFE" box="[167,1414,844,863]" gridrow="15" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FCDBFE70FCFE" box="[167,392,844,863]" gridcol="0" gridrow="15" pageId="5" pageNumber="170">Caudal peduncle scales</th>
<td id="76E95D8112607DF9FE3BFCDBFD54FCFE" box="[436,684,844,863]" gridcol="1" gridrow="15" pageId="5" pageNumber="170">39</td>
<td id="76E95D8112607DF9FD35FCDBFC38FCFE" box="[698,960,844,863]" gridcol="2" gridrow="15" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FCDBFB4DFCFE" box="[973,1205,844,863]" gridcol="3" gridrow="15" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FCDBFA7EFCFE" box="[1228,1414,844,863]" gridcol="4" gridrow="15" pageId="5" pageNumber="170">4043 (42)</td>
</tr>
<tr id="353834FD12607DF9FF28FCFDFA7EFCDF" box="[167,1414,874,894]" gridrow="16" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FCFDFE70FCDF" box="[167,392,874,894]" gridcol="0" gridrow="16" pageId="5" pageNumber="170">Canine teeth on upper jaw</th>
<td id="76E95D8112607DF9FE3BFCFDFD54FCDF" box="[436,684,874,894]" gridcol="1" gridrow="16" pageId="5" pageNumber="170">5</td>
<td id="76E95D8112607DF9FD35FCFDFC38FCDF" box="[698,960,874,894]" gridcol="2" gridrow="16" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FCFDFB4DFCDF" box="[973,1205,874,894]" gridcol="3" gridrow="16" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FCFDFA7EFCDF" box="[1228,1414,874,894]" gridcol="4" gridrow="16" pageId="5" pageNumber="170">711 (8)</td>
</tr>
<tr id="353834FD12607DF9FF28FC1EFA7EFC3C" box="[167,1414,905,925]" gridrow="17" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FC1EFE70FC3C" box="[167,392,905,925]" gridcol="0" gridrow="17" pageId="5" pageNumber="170">Canine teeth on lower jaw</th>
<td id="76E95D8112607DF9FE3BFC1EFD54FC3C" box="[436,684,905,925]" gridcol="1" gridrow="17" pageId="5" pageNumber="170">10</td>
<td id="76E95D8112607DF9FD35FC1EFC38FC3C" box="[698,960,905,925]" gridcol="2" gridrow="17" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FC1EFB4DFC3C" box="[973,1205,905,925]" gridcol="3" gridrow="17" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FC1EFA7EFC3C" box="[1228,1414,905,925]" gridcol="4" gridrow="17" pageId="5" pageNumber="170">47 (5)</td>
</tr>
<tr id="353834FD12607DF9FF28FC24FA7EFC67" box="[167,1414,947,966]" gridrow="18" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FC24FA7EFC67" box="[167,1414,947,966]" colspan="5" colspanRight="4" gridcol="0" gridrow="18" pageId="5" pageNumber="170">
<emphasis id="B97CEAAD12608203FF28FC24FE86FC67" bold="true" box="[167,382,947,966]" pageId="5" pageNumber="170">Measurements (% of SL)</emphasis>
</th>
</tr>
<tr id="353834FD12607DF9FF28FC45FA7EFC44" box="[167,1414,978,997]" gridrow="19" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FC45FE70FC44" box="[167,392,978,997]" gridcol="0" gridrow="19" pageId="5" pageNumber="170">Total length</th>
<td id="76E95D8112607DF9FE3BFC45FD54FC44" box="[436,684,978,997]" gridcol="1" gridrow="19" pageId="5" pageNumber="170">119.8</td>
<td id="76E95D8112607DF9FD35FC45FC38FC44" box="[698,960,978,997]" gridcol="2" gridrow="19" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FC45FB4DFC44" box="[973,1205,978,997]" gridcol="3" gridrow="19" pageId="5" pageNumber="170">124.1</td>
<td id="76E95D8112607DF9FB43FC45FA7EFC44" box="[1228,1414,978,997]" gridcol="4" gridrow="19" pageId="5" pageNumber="170">120.8125.6</td>
</tr>
<tr id="353834FD12607DF9FF28FC66FA7EFBA5" box="[167,1414,1009,1028]" gridrow="20" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FC66FE70FBA5" box="[167,392,1009,1028]" gridcol="0" gridrow="20" pageId="5" pageNumber="170">Head length</th>
<td id="76E95D8112607DF9FE3BFC66FD54FBA5" box="[436,684,1009,1028]" gridcol="1" gridrow="20" pageId="5" pageNumber="170">27.8</td>
<td id="76E95D8112607DF9FD35FC66FC38FBA5" box="[698,960,1009,1028]" gridcol="2" gridrow="20" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FC66FB4DFBA5" box="[973,1205,1009,1028]" gridcol="3" gridrow="20" pageId="5" pageNumber="170">26.8</td>
<td id="76E95D8112607DF9FB43FC66FA7EFBA5" box="[1228,1414,1009,1028]" gridcol="4" gridrow="20" pageId="5" pageNumber="170">28.430.7</td>
</tr>
<tr id="353834FD12607DF9FF28FB87FA7EFB82" box="[167,1414,1040,1059]" gridrow="21" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FB87FE70FB82" box="[167,392,1040,1059]" gridcol="0" gridrow="21" pageId="5" pageNumber="170">Snout length</th>
<td id="76E95D8112607DF9FE3BFB87FD54FB82" box="[436,684,1040,1059]" gridcol="1" gridrow="21" pageId="5" pageNumber="170">7.4</td>
<td id="76E95D8112607DF9FD35FB87FC38FB82" box="[698,960,1040,1059]" gridcol="2" gridrow="21" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FB87FB4DFB82" box="[973,1205,1040,1059]" gridcol="3" gridrow="21" pageId="5" pageNumber="170">7.1</td>
<td id="76E95D8112607DF9FB43FB87FA7EFB82" box="[1228,1414,1040,1059]" gridcol="4" gridrow="21" pageId="5" pageNumber="170">7.68.9</td>
</tr>
<tr id="353834FD12607DF9FF28FBB8FA7EFBE3" box="[167,1414,1071,1090]" gridrow="22" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FBB8FE70FBE3" box="[167,392,1071,1090]" gridcol="0" gridrow="22" pageId="5" pageNumber="170">Eye diameter</th>
<td id="76E95D8112607DF9FE3BFBB8FD54FBE3" box="[436,684,1071,1090]" gridcol="1" gridrow="22" pageId="5" pageNumber="170">3.3</td>
<td id="76E95D8112607DF9FD35FBB8FC38FBE3" box="[698,960,1071,1090]" gridcol="2" gridrow="22" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FBB8FB4DFBE3" box="[973,1205,1071,1090]" gridcol="3" gridrow="22" pageId="5" pageNumber="170">3.6</td>
<td id="76E95D8112607DF9FB43FBB8FA7EFBE3" box="[1228,1414,1071,1090]" gridcol="4" gridrow="22" pageId="5" pageNumber="170">3.35.1</td>
</tr>
<tr id="353834FD12607DF9FF28FBD9FA7EFBC0" box="[167,1414,1102,1121]" gridrow="23" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FBD9FE70FBC0" box="[167,392,1102,1121]" gridcol="0" gridrow="23" pageId="5" pageNumber="170">Interorbital distance</th>
<td id="76E95D8112607DF9FE3BFBD9FD54FBC0" box="[436,684,1102,1121]" gridcol="1" gridrow="23" pageId="5" pageNumber="170">8.5</td>
<td id="76E95D8112607DF9FD35FBD9FC38FBC0" box="[698,960,1102,1121]" gridcol="2" gridrow="23" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FBD9FB4DFBC0" box="[973,1205,1102,1121]" gridcol="3" gridrow="23" pageId="5" pageNumber="170">10.7</td>
<td id="76E95D8112607DF9FB43FBD9FA7EFBC0" box="[1228,1414,1102,1121]" gridcol="4" gridrow="23" pageId="5" pageNumber="170">8.611.2</td>
</tr>
<tr id="353834FD12607DF9FF28FBFAFA7EFB21" box="[167,1414,1133,1152]" gridrow="24" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FBFAFE70FB21" box="[167,392,1133,1152]" gridcol="0" gridrow="24" pageId="5" pageNumber="170">Postorbital length</th>
<td id="76E95D8112607DF9FE3BFBFAFD54FB21" box="[436,684,1133,1152]" gridcol="1" gridrow="24" pageId="5" pageNumber="170">18.5</td>
<td id="76E95D8112607DF9FD35FBFAFC38FB21" box="[698,960,1133,1152]" gridcol="2" gridrow="24" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FBFAFB4DFB21" box="[973,1205,1133,1152]" gridcol="3" gridrow="24" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FBFAFA7EFB21" box="[1228,1414,1133,1152]" gridcol="4" gridrow="24" pageId="5" pageNumber="170">18.219.6</td>
</tr>
<tr id="353834FD12607DF9FF28FB1CFA7EFB3E" box="[167,1414,1163,1183]" gridrow="25" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FB1CFE70FB3E" box="[167,392,1163,1183]" gridcol="0" gridrow="25" pageId="5" pageNumber="170">Upper jaw length</th>
<td id="76E95D8112607DF9FE3BFB1CFD54FB3E" box="[436,684,1163,1183]" gridcol="1" gridrow="25" pageId="5" pageNumber="170">11.6</td>
<td id="76E95D8112607DF9FD35FB1CFC38FB3E" box="[698,960,1163,1183]" gridcol="2" gridrow="25" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FB1CFB4DFB3E" box="[973,1205,1163,1183]" gridcol="3" gridrow="25" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FB1CFA7EFB3E" box="[1228,1414,1163,1183]" gridcol="4" gridrow="25" pageId="5" pageNumber="170">10.512.5</td>
</tr>
<tr id="353834FD12607DF9FF28FB3DFA7EFB1E" box="[167,1414,1194,1215]" gridrow="26" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FB3DFE70FB1E" box="[167,392,1194,1215]" gridcol="0" gridrow="26" pageId="5" pageNumber="170">Body depth at P1 base</th>
<td id="76E95D8112607DF9FE3BFB3DFD54FB1E" box="[436,684,1194,1215]" gridcol="1" gridrow="26" pageId="5" pageNumber="170">18.3</td>
<td id="76E95D8112607DF9FD35FB3DFC38FB1E" box="[698,960,1194,1215]" gridcol="2" gridrow="26" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FB3DFB4DFB1E" box="[973,1205,1194,1215]" gridcol="3" gridrow="26" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FB3DFA7EFB1E" box="[1228,1414,1194,1215]" gridcol="4" gridrow="26" pageId="5" pageNumber="170">14.618.9</td>
</tr>
<tr id="353834FD12607DF9FF28FB5EFA7EFB7C" box="[167,1414,1225,1245]" gridrow="27" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FB5EFE70FB7C" box="[167,392,1225,1245]" gridcol="0" gridrow="27" pageId="5" pageNumber="170">Body depth at D2 base</th>
<td id="76E95D8112607DF9FE3BFB5EFD54FB7C" box="[436,684,1225,1245]" gridcol="1" gridrow="27" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FD35FB5EFC38FB7C" box="[698,960,1225,1245]" gridcol="2" gridrow="27" pageId="5" pageNumber="170">
25.0
<emphasis id="B97CEAAD12608203FCDCFB5EFC90FB7D" bold="true" box="[851,872,1225,1244]" pageId="5" pageNumber="170">**</emphasis>
</td>
<td id="76E95D8112607DF9FC42FB5EFB4DFB7C" box="[973,1205,1225,1245]" gridcol="3" gridrow="27" pageId="5" pageNumber="170">
19.6
<emphasis id="B97CEAAD12608203FBDFFB5EFB9DFB7D" bold="true" box="[1104,1125,1225,1244]" pageId="5" pageNumber="170">**</emphasis>
</td>
<td id="76E95D8112607DF9FB43FB5EFA7EFB7C" box="[1228,1414,1225,1245]" gridcol="4" gridrow="27" pageId="5" pageNumber="170">16.219.4</td>
</tr>
<tr id="353834FD12607DF9FF28FB7FFA7EFB5D" box="[167,1414,1256,1276]" gridrow="28" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FB7FFE70FB5D" box="[167,392,1256,1276]" gridcol="0" gridrow="28" pageId="5" pageNumber="170">Body depth at A base</th>
<td id="76E95D8112607DF9FE3BFB7FFD54FB5D" box="[436,684,1256,1276]" gridcol="1" gridrow="28" pageId="5" pageNumber="170">18.4</td>
<td id="76E95D8112607DF9FD35FB7FFC38FB5D" box="[698,960,1256,1276]" gridcol="2" gridrow="28" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FB7FFB4DFB5D" box="[973,1205,1256,1276]" gridcol="3" gridrow="28" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FB7FFA7EFB5D" box="[1228,1414,1256,1276]" gridcol="4" gridrow="28" pageId="5" pageNumber="170">14.617.1</td>
</tr>
<tr id="353834FD12607DF9FF28FA90FA7EFABA" box="[167,1414,1287,1307]" gridrow="29" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA90FE70FABA" box="[167,392,1287,1307]" gridcol="0" gridrow="29" pageId="5" pageNumber="170">Caudal-peduncle depth</th>
<td id="76E95D8112607DF9FE3BFA90FD54FABA" box="[436,684,1287,1307]" gridcol="1" gridrow="29" pageId="5" pageNumber="170">15.0</td>
<td id="76E95D8112607DF9FD35FA90FC38FABA" box="[698,960,1287,1307]" gridcol="2" gridrow="29" pageId="5" pageNumber="170">20.0</td>
<td id="76E95D8112607DF9FC42FA90FB4DFABA" box="[973,1205,1287,1307]" gridcol="3" gridrow="29" pageId="5" pageNumber="170">15.2</td>
<td id="76E95D8112607DF9FB43FA90FA7EFABA" box="[1228,1414,1287,1307]" gridcol="4" gridrow="29" pageId="5" pageNumber="170">12.613.6</td>
</tr>
<tr id="353834FD12607DF9FF28FAB1FA7EFA9B" box="[167,1414,1318,1338]" gridrow="30" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FAB1FE70FA9B" box="[167,392,1318,1338]" gridcol="0" gridrow="30" pageId="5" pageNumber="170">Caudal-peduncle length</th>
<td id="76E95D8112607DF9FE3BFAB1FD54FA9B" box="[436,684,1318,1338]" gridcol="1" gridrow="30" pageId="5" pageNumber="170">24.4</td>
<td id="76E95D8112607DF9FD35FAB1FC38FA9B" box="[698,960,1318,1338]" gridcol="2" gridrow="30" pageId="5" pageNumber="170">18.0</td>
<td id="76E95D8112607DF9FC42FAB1FB4DFA9B" box="[973,1205,1318,1338]" gridcol="3" gridrow="30" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FAB1FA7EFA9B" box="[1228,1414,1318,1338]" gridcol="4" gridrow="30" pageId="5" pageNumber="170">22.025.3</td>
</tr>
<tr id="353834FD12607DF9FF28FAD2FA7EFAF8" box="[167,1414,1349,1369]" gridrow="31" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FAD2FE70FAF8" box="[167,392,1349,1369]" gridcol="0" gridrow="31" pageId="5" pageNumber="170">Predorsal length</th>
<td id="76E95D8112607DF9FE3BFAD2FD54FAF8" box="[436,684,1349,1369]" gridcol="1" gridrow="31" pageId="5" pageNumber="170">38.2</td>
<td id="76E95D8112607DF9FD35FAD2FC38FAF8" box="[698,960,1349,1369]" gridcol="2" gridrow="31" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FAD2FB4DFAF8" box="[973,1205,1349,1369]" gridcol="3" gridrow="31" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FAD2FA7EFAF8" box="[1228,1414,1349,1369]" gridcol="4" gridrow="31" pageId="5" pageNumber="170">38.740.7</td>
</tr>
<tr id="353834FD12607DF9FF28FAF3FA7EFAD9" box="[167,1414,1380,1400]" gridrow="32" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FAF3FE70FAD9" box="[167,392,1380,1400]" gridcol="0" gridrow="32" pageId="5" pageNumber="170">Preanal-fin length</th>
<td id="76E95D8112607DF9FE3BFAF3FD54FAD9" box="[436,684,1380,1400]" gridcol="1" gridrow="32" pageId="5" pageNumber="170">64.0</td>
<td id="76E95D8112607DF9FD35FAF3FC38FAD9" box="[698,960,1380,1400]" gridcol="2" gridrow="32" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FAF3FB4DFAD9" box="[973,1205,1380,1400]" gridcol="3" gridrow="32" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FAF3FA7EFAD9" box="[1228,1414,1380,1400]" gridcol="4" gridrow="32" pageId="5" pageNumber="170">60.664.8</td>
</tr>
<tr id="353834FD12607DF9FF28FA14FA7EFA36" box="[167,1414,1411,1431]" gridrow="33" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA14FE70FA36" box="[167,392,1411,1431]" gridcol="0" gridrow="33" pageId="5" pageNumber="170">Preanal length</th>
<td id="76E95D8112607DF9FE3BFA14FD54FA36" box="[436,684,1411,1431]" gridcol="1" gridrow="33" pageId="5" pageNumber="170">60.2</td>
<td id="76E95D8112607DF9FD35FA14FC38FA36" box="[698,960,1411,1431]" gridcol="2" gridrow="33" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FA14FB4DFA36" box="[973,1205,1411,1431]" gridcol="3" gridrow="33" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FA14FA7EFA36" box="[1228,1414,1411,1431]" gridcol="4" gridrow="33" pageId="5" pageNumber="170">56.659.2</td>
</tr>
<tr id="353834FD12607DF9FF28FA35FA7EFA17" box="[167,1414,1442,1462]" gridrow="34" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA35FE70FA17" box="[167,392,1442,1462]" gridcol="0" gridrow="34" pageId="5" pageNumber="170">D1 base length</th>
<td id="76E95D8112607DF9FE3BFA35FD54FA17" box="[436,684,1442,1462]" gridcol="1" gridrow="34" pageId="5" pageNumber="170">15.6</td>
<td id="76E95D8112607DF9FD35FA35FC38FA17" box="[698,960,1442,1462]" gridcol="2" gridrow="34" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FA35FB4DFA17" box="[973,1205,1442,1462]" gridcol="3" gridrow="34" pageId="5" pageNumber="170">10.7</td>
<td id="76E95D8112607DF9FB43FA35FA7EFA17" box="[1228,1414,1442,1462]" gridcol="4" gridrow="34" pageId="5" pageNumber="170">13.214.6</td>
</tr>
<tr id="353834FD12607DF9FF28FA56FA7EFA75" box="[167,1414,1473,1492]" gridrow="35" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA56FE70FA75" box="[167,392,1473,1492]" gridcol="0" gridrow="35" pageId="5" pageNumber="170">D1 length</th>
<td id="76E95D8112607DF9FE3BFA56FD54FA75" box="[436,684,1473,1492]" gridcol="1" gridrow="35" pageId="5" pageNumber="170">16.5</td>
<td id="76E95D8112607DF9FD35FA56FC38FA75" box="[698,960,1473,1492]" gridcol="2" gridrow="35" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FA56FB4DFA75" box="[973,1205,1473,1492]" gridcol="3" gridrow="35" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FA56FA7EFA75" box="[1228,1414,1473,1492]" gridcol="4" gridrow="35" pageId="5" pageNumber="170">13.115.2</td>
</tr>
<tr id="353834FD12607DF9FF28FA77FA7EFA52" box="[167,1414,1504,1523]" gridrow="36" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA77FE70FA52" box="[167,392,1504,1523]" gridcol="0" gridrow="36" pageId="5" pageNumber="170">D1-D2 distance</th>
<td id="76E95D8112607DF9FE3BFA77FD54FA52" box="[436,684,1504,1523]" gridcol="1" gridrow="36" pageId="5" pageNumber="170">4.7</td>
<td id="76E95D8112607DF9FD35FA77FC38FA52" box="[698,960,1504,1523]" gridcol="2" gridrow="36" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FA77FB4DFA52" box="[973,1205,1504,1523]" gridcol="3" gridrow="36" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43FA77FA7EFA52" box="[1228,1414,1504,1523]" gridcol="4" gridrow="36" pageId="5" pageNumber="170">4.55.9</td>
</tr>
<tr id="353834FD12607DF9FF28FA68FA7EF9B3" box="[167,1414,1535,1554]" gridrow="37" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28FA68FE70F9B3" box="[167,392,1535,1554]" gridcol="0" gridrow="37" pageId="5" pageNumber="170">D2 base length</th>
<td id="76E95D8112607DF9FE3BFA68FD54F9B3" box="[436,684,1535,1554]" gridcol="1" gridrow="37" pageId="5" pageNumber="170">21.6</td>
<td id="76E95D8112607DF9FD35FA68FC38F9B3" box="[698,960,1535,1554]" gridcol="2" gridrow="37" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42FA68FB4DF9B3" box="[973,1205,1535,1554]" gridcol="3" gridrow="37" pageId="5" pageNumber="170">21.4</td>
<td id="76E95D8112607DF9FB43FA68FA7EF9B3" box="[1228,1414,1535,1554]" gridcol="4" gridrow="37" pageId="5" pageNumber="170">18.120.0</td>
</tr>
<tr id="353834FD12607DF9FF28F989FA7EF990" box="[167,1414,1566,1585]" gridrow="38" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F989FE70F990" box="[167,392,1566,1585]" gridcol="0" gridrow="38" pageId="5" pageNumber="170">D2 length</th>
<td id="76E95D8112607DF9FE3BF989FD54F990" box="[436,684,1566,1585]" gridcol="1" gridrow="38" pageId="5" pageNumber="170">31.2</td>
<td id="76E95D8112607DF9FD35F989FC38F990" box="[698,960,1566,1585]" gridcol="2" gridrow="38" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F989FB4DF990" box="[973,1205,1566,1585]" gridcol="3" gridrow="38" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43F989FA7EF990" box="[1228,1414,1566,1585]" gridcol="4" gridrow="38" pageId="5" pageNumber="170">27.331.0</td>
</tr>
<tr id="353834FD12607DF9FF28F9AAFA7EF9F1" box="[167,1414,1597,1616]" gridrow="39" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F9AAFE70F9F1" box="[167,392,1597,1616]" gridcol="0" gridrow="39" pageId="5" pageNumber="170">A base length</th>
<td id="76E95D8112607DF9FE3BF9AAFD54F9F1" box="[436,684,1597,1616]" gridcol="1" gridrow="39" pageId="5" pageNumber="170">13.1</td>
<td id="76E95D8112607DF9FD35F9AAFC38F9F1" box="[698,960,1597,1616]" gridcol="2" gridrow="39" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F9AAFB4DF9F1" box="[973,1205,1597,1616]" gridcol="3" gridrow="39" pageId="5" pageNumber="170">17.9</td>
<td id="76E95D8112607DF9FB43F9AAFA7EF9F1" box="[1228,1414,1597,1616]" gridcol="4" gridrow="39" pageId="5" pageNumber="170">15.017.2</td>
</tr>
<tr id="353834FD12607DF9FF28F9CBFA7EF9CE" box="[167,1414,1628,1647]" gridrow="40" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F9CBFE70F9CE" box="[167,392,1628,1647]" gridcol="0" gridrow="40" pageId="5" pageNumber="170">A length</th>
<td id="76E95D8112607DF9FE3BF9CBFD54F9CE" box="[436,684,1628,1647]" gridcol="1" gridrow="40" pageId="5" pageNumber="170">23.4</td>
<td id="76E95D8112607DF9FD35F9CBFC38F9CE" box="[698,960,1628,1647]" gridcol="2" gridrow="40" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F9CBFB4DF9CE" box="[973,1205,1628,1647]" gridcol="3" gridrow="40" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43F9CBFA7EF9CE" box="[1228,1414,1628,1647]" gridcol="4" gridrow="40" pageId="5" pageNumber="170">23.226.8</td>
</tr>
<tr id="353834FD12607DF9FF28F9ECFA7EF92E" box="[167,1414,1659,1679]" gridrow="41" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F9ECFE70F92E" box="[167,392,1659,1679]" gridcol="0" gridrow="41" pageId="5" pageNumber="170">P1 base length</th>
<td id="76E95D8112607DF9FE3BF9ECFD54F92E" box="[436,684,1659,1679]" gridcol="1" gridrow="41" pageId="5" pageNumber="170">6.8</td>
<td id="76E95D8112607DF9FD35F9ECFC38F92E" box="[698,960,1659,1679]" gridcol="2" gridrow="41" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F9ECFB4DF92E" box="[973,1205,1659,1679]" gridcol="3" gridrow="41" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FB43F9ECFA7EF92E" box="[1228,1414,1659,1679]" gridcol="4" gridrow="41" pageId="5" pageNumber="170">7.69.1</td>
</tr>
<tr id="353834FD12607DF9FF28F90EFA7EF90F" box="[167,1414,1689,1710]" gridrow="42" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F90EFE70F90F" box="[167,392,1689,1710]" gridcol="0" gridrow="42" pageId="5" pageNumber="170">P1 length</th>
<td id="76E95D8112607DF9FE3BF90EFD54F90F" box="[436,684,1689,1710]" gridcol="1" gridrow="42" pageId="5" pageNumber="170">16.4</td>
<td id="76E95D8112607DF9FD35F90EFC38F90F" box="[698,960,1689,1710]" gridcol="2" gridrow="42" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F90EFB4DF90F" box="[973,1205,1689,1710]" gridcol="3" gridrow="42" pageId="5" pageNumber="170">17.0</td>
<td id="76E95D8112607DF9FB43F90EFA7EF90F" box="[1228,1414,1689,1710]" gridcol="4" gridrow="42" pageId="5" pageNumber="170">15.120.5</td>
</tr>
<tr id="353834FD12607DF9FF28F92FFA7EF96C" box="[167,1414,1720,1741]" gridrow="43" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F92FFE70F96C" box="[167,392,1720,1741]" gridcol="0" gridrow="43" pageId="5" pageNumber="170">P2 length</th>
<td id="76E95D8112607DF9FE3BF92FFD54F96C" box="[436,684,1720,1741]" gridcol="1" gridrow="43" pageId="5" pageNumber="170">14.4</td>
<td id="76E95D8112607DF9FD35F92FFC38F96C" box="[698,960,1720,1741]" gridcol="2" gridrow="43" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F92FFB4DF96C" box="[973,1205,1720,1741]" gridcol="3" gridrow="43" pageId="5" pageNumber="170">17.9</td>
<td id="76E95D8112607DF9FB43F92FFA7EF96C" box="[1228,1414,1720,1741]" gridcol="4" gridrow="43" pageId="5" pageNumber="170">11.620.2</td>
</tr>
<tr id="353834FD12607DF9FF28F940FA7EF94A" box="[167,1414,1751,1771]" gridrow="44" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F940FE70F94A" box="[167,392,1751,1771]" gridcol="0" gridrow="44" pageId="5" pageNumber="170">C length</th>
<td id="76E95D8112607DF9FE3BF940FD54F94A" box="[436,684,1751,1771]" gridcol="1" gridrow="44" pageId="5" pageNumber="170">19.3</td>
<td id="76E95D8112607DF9FD35F940FC38F94A" box="[698,960,1751,1771]" gridcol="2" gridrow="44" pageId="5" pageNumber="170"></td>
<td id="76E95D8112607DF9FC42F940FB4DF94A" box="[973,1205,1751,1771]" gridcol="3" gridrow="44" pageId="5" pageNumber="170">24.1</td>
<td id="76E95D8112607DF9FB43F940FA7EF94A" box="[1228,1414,1751,1771]" gridcol="4" gridrow="44" pageId="5" pageNumber="170">19.924.6</td>
</tr>
<tr id="353834FD12607DF9FF28F896FA7EF8B5" box="[167,1414,1793,1812]" gridrow="45" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F896FA7EF8B5" box="[167,1414,1793,1812]" colspan="5" colspanRight="4" gridcol="0" gridrow="45" pageId="5" pageNumber="170">
<emphasis id="B97CEAAD12608203FF28F896FE7AF8B5" bold="true" box="[167,386,1793,1812]" pageId="5" pageNumber="170">Measurements (% of TL)</emphasis>
</th>
</tr>
<tr id="353834FD12607DF9FF28F8B7FA7EF892" box="[167,1414,1824,1843]" gridrow="46" pageId="5" pageNumber="170">
<th id="76E95D8112607DF9FF28F8B7FE70F892" box="[167,392,1824,1843]" gridcol="0" gridrow="46" pageId="5" pageNumber="170">Head length</th>
<td id="76E95D8112607DF9FE3BF8B7FD54F892" box="[436,684,1824,1843]" gridcol="1" gridrow="46" pageId="5" pageNumber="170">21.6</td>
<td id="76E95D8112607DF9FD35F8B7FC38F892" box="[698,960,1824,1843]" gridcol="2" gridrow="46" pageId="5" pageNumber="170">29.4</td>
<td id="76E95D8112607DF9FC42F8B7FB4DF892" box="[973,1205,1824,1843]" gridcol="3" gridrow="46" pageId="5" pageNumber="170">21.6</td>
<td id="76E95D8112607DF9FB43F8B7FA7EF892" box="[1228,1414,1824,1843]" gridcol="4" gridrow="46" pageId="5" pageNumber="170">23.525.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB736BF12608203FF3EF8DDFAC1F8D6" blockId="5.[177,1401,1866,1912]" pageId="5" pageNumber="170">
<tableNote id="76EE373112608203FF3EF8DDFAC1F8D6" pageId="5" pageNumber="170" targetBox="[167,1414,326,1843]" targetPageId="5">
D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P
<subScript id="178C34FA12608203FD79F8C5FD05F8FF" attach="left" box="[758,765,1874,1886]" fontSize="5" pageId="5" pageNumber="170">1</subScript>
, pectoral fin; P
<subScript id="178C34FA12608203FCF1F8C5FC7DF8FF" attach="left" box="[894,901,1874,1886]" fontSize="5" pageId="5" pageNumber="170">2</subScript>
, pelvic fin.
<emphasis id="B97CEAAD12608203FC69F8DDFC09F8FC" bold="true" box="[998,1009,1866,1885]" pageId="5" pageNumber="170">*</emphasis>
Number of fin rays were counted for all seven specimens, but six specimens excluding URM-P 49660 were used for other characters.
<emphasis id="B97CEAAD12608203FC1EF8F3FC5EF8D6" bold="true" box="[913,934,1892,1911]" pageId="5" pageNumber="170">**</emphasis>
Detailed method of measurement is unknown.
</tableNote>
</paragraph>
<paragraph id="8BB736BF12638200FCC1FEDFFC20FCE9" blockId="6.[820,1432,198,1589]" pageId="6" pageNumber="171">
Cephalic sensory systems illustrated in
<figureCitation id="13332A3A12638200FA83FEDFFAB7FEC1" box="[1292,1359,328,352]" captionStart="Fig" captionStartId="6.[156,187,1505,1527]" captionTargetBox="[159,765,479,1467]" captionTargetId="figure-544@6.[158,766,478,1468]" captionTargetPageId="6" captionText="Fig. 5. Sensory pores (open circles with uppercase notations) and neuromast rows (red dots with lowercase notations) in the head of Paloa villadolidi (NSMT-P 143769, 55.4mm SL). A, Dorsal, B, lateral, and C, ventral views. AN, anterior nostril; PN, posterior nostril." figureDoi="http://doi.org/10.5281/zenodo.10833170" httpUri="https://zenodo.org/record/10833170/files/figure.png" pageId="6" pageNumber="171">Fig. 5</figureCitation>
. Oculoscapular canal with 12 pores (A, B, C, D, E, F, G, H, I, J, K, and L) and preopercular canal with five pores (M, N, O, P, and Q), but
<specimenCount id="9D0EFD3612638200FC58FE3CFB9DFE63" box="[983,1125,426,450]" count="1" pageId="6" pageNumber="171" type="generic">one specimen</specimenCount>
(NSMT-P14769) lacked pore G on both sides; pore D paired with an interval of approximately half of the interorbital width. Arrangement of the sensory papillae transverse pattern: five transverse (
<emphasis id="B97CEAAD12638200FAE5FD9BFA77FD85" box="[1386,1423,524,548]" italics="true" pageId="6" pageNumber="171">15</emphasis>
) and two longitudinal rows (
<emphasis id="B97CEAAD12638200FBD5FDBBFB9EFDE5" box="[1114,1126,556,580]" italics="true" pageId="6" pageNumber="171">b</emphasis>
and
<emphasis id="B97CEAAD12638200FB10FDBBFB54FDE5" box="[1183,1196,556,580]" italics="true" pageId="6" pageNumber="171">d</emphasis>
) of the sensory papillae on the suborbital region and cheek; transverse row
<quantity id="4CF09B5A12638200FA04FDDAFCB1FD24" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="171" unit="in" value="3.0">
<emphasis id="B97CEAAD12638200FA04FDDAFA6FFDC4" box="[1419,1431,589,613]" italics="true" pageId="6" pageNumber="171">3</emphasis>
in
</quantity>
contact with longitudinal row
<emphasis id="B97CEAAD12638200FB01FDFAFB62FD24" box="[1166,1178,621,645]" italics="true" pageId="6" pageNumber="171">b</emphasis>
at the middle (
<figureCitation id="13332A3A12638200FAB1FDFAFA74FD24" box="[1342,1420,621,645]" captionStart="Fig" captionStartId="6.[156,187,1505,1527]" captionTargetBox="[159,765,479,1467]" captionTargetId="figure-544@6.[158,766,478,1468]" captionTargetPageId="6" captionText="Fig. 5. Sensory pores (open circles with uppercase notations) and neuromast rows (red dots with lowercase notations) in the head of Paloa villadolidi (NSMT-P 143769, 55.4mm SL). A, Dorsal, B, lateral, and C, ventral views. AN, anterior nostril; PN, posterior nostril." figureDoi="http://doi.org/10.5281/zenodo.10833170" httpUri="https://zenodo.org/record/10833170/files/figure.png" pageId="6" pageNumber="171">Fig. 5B</figureCitation>
). Twelve transverse rows on underside of head (
<emphasis id="B97CEAAD12638200FA98FD19FAE6FD04" box="[1303,1310,654,677]" italics="true" pageId="6" pageNumber="171">i</emphasis>
), with each row aligned on a ridge. Operculum with an elongated transverse row (
<emphasis id="B97CEAAD12638200FC20FD47FC3AFD46" box="[943,962,720,743]" italics="true" pageId="6" pageNumber="171">ot</emphasis>
) along preopercular margin, connecting an oculoscapular transverse row (
<emphasis id="B97CEAAD12638200FBE3FD67FB8FFCA6" box="[1132,1143,752,775]" italics="true" pageId="6" pageNumber="171">z</emphasis>
) around pore M; an oblique row (
<emphasis id="B97CEAAD12638200FCE0FC86FC7BFC89" box="[879,899,785,808]" italics="true" pageId="6" pageNumber="171">os</emphasis>
) and a longitudinal row (
<emphasis id="B97CEAAD12638200FB12FC86FB48FC89" box="[1181,1200,785,808]" italics="true" pageId="6" pageNumber="171">oi</emphasis>
); with their posterior ends not joined.
</paragraph>
<paragraph id="8BB736BF12638200FCC1FCC6FBD8FB6E" blockId="6.[820,1432,198,1589]" pageId="6" pageNumber="171">
Approximately 18 small transverse rows (
<emphasis id="B97CEAAD12638200FA96FCC6FACBFCC8" box="[1305,1331,849,873]" italics="true" pageId="6" pageNumber="171">lm</emphasis>
) of sensory papillae aligned along the lateral midline of the body (
<figureCitation id="13332A3A12638200FCB3FC05FC80FC0B" box="[828,888,914,938]" captionStart="Fig" captionStartId="6.[156,187,2056,2078]" captionTargetBox="[207,1380,1657,2016]" captionTargetId="figure-589@6.[206,1381,1656,2017]" captionTargetPageId="6" captionText="Fig. 6. Neuromast rows (red dots with lowercase notations) of trunk and caudal fin in Paloa villadolidi (NSMT-P 143769, 55.4mm SL)." figureDoi="http://doi.org/10.5281/zenodo.10833174" httpUri="https://zenodo.org/record/10833174/files/figure.png" pageId="6" pageNumber="171">Fig. 6</figureCitation>
). Two transverse and three longitudinal rows (
<emphasis id="B97CEAAD12638200FAD5FC05FA95FC0B" box="[1370,1389,914,938]" italics="true" pageId="6" pageNumber="171">ld</emphasis>
) on dorsal to dorsolateral side of the body: anteriormost row (
<emphasis id="B97CEAAD12638200FCB2FC44FCA0FC40" box="[829,856,979,1003]" italics="true" pageId="6" pageNumber="171">
ld
<superScript id="7C7D9BF712638200FCDFFC44FCA0FC40" attach="left" box="[848,856,979,993]" fontSize="6" pageId="6" pageNumber="171">1</superScript>
</emphasis>
) below the base of the third spine of the first dorsal fin; posteriormost row (
<emphasis id="B97CEAAD12638200FB8CFC63FBE6FBA3" box="[1027,1054,1012,1036]" italics="true" pageId="6" pageNumber="171">
ld
<superScript id="7C7D9BF712638200FB99FC63FBE6FBA3" attach="left" box="[1046,1054,1012,1026]" fontSize="6" pageId="6" pageNumber="171">5</superScript>
</emphasis>
) on the posterior part of caudal peduncle. Two transverse rows (
<emphasis id="B97CEAAD12638200FBE6FB83FB82FB8D" box="[1129,1146,1044,1068]" italics="true" pageId="6" pageNumber="171">lv</emphasis>
) on ventral to ventrolateral side of the body: anterior row (
<emphasis id="B97CEAAD12638200FBFFFBA2FB71FBE2" box="[1136,1161,1077,1101]" italics="true" pageId="6" pageNumber="171">
lv
<superScript id="7C7D9BF712638200FB0EFBA2FB71FBE2" attach="left" box="[1153,1161,1077,1091]" fontSize="6" pageId="6" pageNumber="171">1</superScript>
</emphasis>
) at middle between posterior end of the pelvic-fin base to origin of anal fin; posterior row (
<emphasis id="B97CEAAD12638200FCE5FBE1FC7BFB25" box="[874,899,1142,1166]" italics="true" pageId="6" pageNumber="171">
lv
<superScript id="7C7D9BF712638200FCF4FBE1FC7BFB25" attach="left" box="[891,899,1142,1156]" fontSize="6" pageId="6" pageNumber="171">2</superScript>
</emphasis>
) above the base of third soft ray of the anal fin. Caudal fin with one transverse row (
<emphasis id="B97CEAAD12638200FB06FB01FB59FB0F" box="[1161,1185,1174,1198]" italics="true" pageId="6" pageNumber="171">lct</emphasis>
) and three longitudinal rows (
<emphasis id="B97CEAAD12638200FCFCFB20FC68FB6E" box="[883,912,1207,1231]" italics="true" pageId="6" pageNumber="171">lcd</emphasis>
,
<emphasis id="B97CEAAD12638200FC14FB20FC47FB6E" box="[923,959,1207,1231]" italics="true" pageId="6" pageNumber="171">lcm</emphasis>
, and
<emphasis id="B97CEAAD12638200FC79FB20FBE9FB6E" box="[1014,1041,1207,1231]" italics="true" pageId="6" pageNumber="171">lcv</emphasis>
).
</paragraph>
<paragraph id="8BB736BF12638201FCC1FB40FF35FAAE" blockId="6.[820,1432,198,1589]" lastBlockId="7.[156,768,882,2077]" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171">
<emphasis id="B97CEAAD12638200FCC1FB40FB32FB4E" bold="true" box="[846,1226,1239,1264]" pageId="6" pageNumber="171">Color of living and fresh specimens.</emphasis>
In
<specimenCount id="9D0EFD3612638200FB70FB40FAA5FB4E" box="[1279,1373,1239,1263]" count="5" pageId="6" pageNumber="171" type="adult">five adult</specimenCount>
specimens (41.075.0 mm SL), body and head pinkish brown laterally, and orange dorsally and dorso-laterally (
<figureCitation id="13332A3A12638200FAC5FA8FFA73FA90" box="[1354,1419,1304,1329]" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="6" pageNumber="171">Fig. 7</figureCitation>
). Throat, breast, and belly white except for a region between the bases of the left and right pelvic fins where brown pigments were distributed. Three obscure, red longitudinal stripes running dorsally, laterally, and ventrally on the body of
<specimenCount id="9D0EFD3612638200FCDBFA2BFC1DFA72" box="[852,997,1467,1491]" count="1" pageId="6" pageNumber="171" type="generic">one specimen</specimenCount>
(NSMT-P 143769:
<figureCitation id="13332A3A12638200FB35FA2CFAF4FA72" box="[1210,1292,1467,1491]" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="6" pageNumber="171">Fig. 7A</figureCitation>
), two stripes running dorsally and laterally of
<specimenCount id="9D0EFD3612638200FB07FA4BFAD4FA55" box="[1160,1324,1500,1524]" count="3" pageId="6" pageNumber="171" type="generic">three specimens</specimenCount>
(NSMT-P 145828145830:
<figureCitation id="13332A3A12638200FC6CFA6BFBCCF9B5" box="[995,1076,1532,1556]" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="6" pageNumber="171">Fig. 7D</figureCitation>
), and single stripe running along the lateral midline of another adult (URM-P 49659:
<figureCitation id="13332A3A12638200FACCF98AFA74F994" box="[1347,1420,1565,1589]" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="6" pageNumber="171">Fig. 7E</figureCitation>
). Iris brown with an internal gold edge. Two transverse, dark, broad bars may be on cheek radiating from the eye (
<figureCitation id="13332A3A12628201FD58FC05FF44FC6A" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="7" pageNumber="172">Fig. 7C</figureCitation>
). First and second dorsal and anal fins translucent with orange or brown spines and soft rays, but proximal parts of the membrane on first (between first and second spines) and second dorsal fins (from origin to end) and anal fin (between spine and first soft ray) brown, and spine of anal fin translucent in
<specimenCount id="9D0EFD3612628201FEBDFBC1FE3AFBCC" box="[306,450,1109,1133]" count="1" pageId="7" pageNumber="172" type="generic">one specimen</specimenCount>
. Caudal and pectoral fins with orange or brown rays; membrane brown proximally and translucent distally. Pelvic fins translucent. In smaller adult specimen (
<quantity id="4CF09B5A12628201FE80FB20FE9FFB6F" box="[271,359,1207,1231]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="7" pageNumber="172" unit="mm" value="31.1">31.1 mm</quantity>
SL), body cream with small melanophores scattered laterally on the body; fins almost translucent.
</paragraph>
<caption id="DF77663712638200FF13FA76FD0FF9EA" ID-DOI="http://doi.org/10.5281/zenodo.10833170" ID-Zenodo-Dep="10833170" httpUri="https://zenodo.org/record/10833170/files/figure.png" pageId="6" pageNumber="171" startId="6.[156,187,1505,1527]" targetBox="[159,765,479,1467]" targetPageId="6" targetType="figure">
<paragraph id="8BB736BF12638200FF13FA76FD0FF9EA" blockId="6.[156,767,1505,1611]" pageId="6" pageNumber="171">
Fig. 5. Sensory pores (open circles with uppercase notations) and neuromast rows (red dots with lowercase notations) in the head of
<taxonomicName id="4C084D3C12638200FF12F98EFED5F98F" box="[157,301,1561,1582]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12638200FF12F98EFED5F98F" box="[157,301,1561,1582]" italics="true" pageId="6" pageNumber="171">Paloa villadolidi</emphasis>
</taxonomicName>
(NSMT-P 143769, 55.4 mm SL). A, Dorsal, B, lateral, and C, ventral views. AN, anterior nostril; PN, posterior nostril.
</paragraph>
</caption>
<caption id="DF77663712638200FF13F79FFA89F7BC" ID-DOI="http://doi.org/10.5281/zenodo.10833174" ID-Zenodo-Dep="10833174" box="[156,1393,2056,2078]" httpUri="https://zenodo.org/record/10833174/files/figure.png" pageId="6" pageNumber="171" startId="6.[156,187,2056,2078]" targetBox="[207,1380,1657,2016]" targetPageId="6" targetType="figure">
<paragraph id="8BB736BF12638200FF13F79FFA89F7BC" blockId="6.[156,1393,2056,2078]" box="[156,1393,2056,2078]" pageId="6" pageNumber="171">
Fig. 6. Neuromast rows (red dots with lowercase notations) of trunk and caudal fin in
<taxonomicName id="4C084D3C12638200FC35F79FFBB2F7BC" box="[954,1098,2056,2077]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12638200FC35F79FFBB2F7BC" box="[954,1098,2056,2077]" italics="true" pageId="6" pageNumber="171">Paloa villadolidi</emphasis>
</taxonomicName>
(NSMT-P 143769, 55.4 mm SL).
</paragraph>
</caption>
<caption id="DF77663712628201FF13FD6BFE8FFCEB" ID-DOI="http://doi.org/10.5281/zenodo.10833176" ID-Zenodo-Dep="10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="7" pageNumber="172" startId="7.[156,187,764,786]" targetBox="[207,1380,206,725]" targetPageId="7" targetType="figure">
<paragraph id="8BB736BF12628201FF13FD6BFE8FFCEB" blockId="7.[156,1431,764,842]" pageId="7" pageNumber="172">
Fig. 7. Live specimens of
<taxonomicName id="4C084D3C12628201FE16FD6BFDD3FCB0" box="[409,555,764,785]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12628201FE16FD6BFDD3FCB0" box="[409,555,764,785]" italics="true" pageId="7" pageNumber="172">Paloa villadolidi</emphasis>
</taxonomicName>
from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1 mm SL); F, URM-P 49661 (ca. 10 mm SL soon after collection). White arrows indicate stripes on the body.
</paragraph>
</caption>
<paragraph id="8BB736BF12628201FF3AFA8EFD52F9D7" blockId="7.[156,768,882,2077]" pageId="7" pageNumber="172">
In a juvenile observed alive in a tank (
<figureCitation id="13332A3A12628201FDEAFA8FFD4DFA90" box="[613,693,1304,1329]" captionStart="Fig" captionStartId="7.[156,187,764,786]" captionTargetBox="[207,1380,206,725]" captionTargetId="figure-554@7.[206,1381,205,726]" captionTargetPageId="7" captionText="Fig. 7. Live specimens of Paloa villadolidi from the Ryukyu Archipelago, Japan. AC, NSMT-P 143769 (55.4mm SL); D, NSMT-P 145829 (41.0mm SL, photo by R. Ueda); E, URM-P 49659 (55.1mm SL); F, URM-P 49661 (ca. 10mm SL soon after collection). White arrows indicate stripes on the body." figureDoi="http://doi.org/10.5281/zenodo.10833176" httpUri="https://zenodo.org/record/10833176/files/figure.png" pageId="7" pageNumber="172">Fig. 7F</figureCitation>
), body translucent with orange dorsally on head and body, around snout and jaws, and on caudal- and pectoral-fin bases; melanophores scattered on the head and body surfaces. Internal melanophores along the ventral midline from anal-fin base to caudal peduncle. Red blood visible in some internal parts of the head, abdominal cavity, and vertebrae. Fins translucent, but second dorsal and caudal fins having orange or brown pigments. After rearing for approximately seven months in a tank, the transparency of the juvenile body decreased and the internal melanophores disappeared.
</paragraph>
<caption id="DF77663712628201FCB9FAF8FA99FA78" ID-DOI="http://doi.org/10.5281/zenodo.10833178" ID-Zenodo-Dep="10833178" httpUri="https://zenodo.org/record/10833178/files/figure.png" pageId="7" pageNumber="172" startId="7.[822,853,1391,1413]" targetBox="[823,1429,901,1351]" targetPageId="7" targetType="figure">
<paragraph id="8BB736BF12628201FCB9FAF8FA99FA78" blockId="7.[821,1433,1391,1497]" pageId="7" pageNumber="172">
Fig. 8. Urogenital papilla of
<taxonomicName id="4C084D3C12628201FBDBFAF8FB10FA25" box="[1108,1256,1391,1412]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12628201FBDBFAF8FB10FA25" box="[1108,1256,1391,1412]" italics="true" pageId="7" pageNumber="172">Paloa villadolidi</emphasis>
</taxonomicName>
. A, Male (NSMTP 143769, 55.4mm SL); B, female (NSMT-P 145829, 75.0 mm SL). Broken lines indicate the outer edge of urogenital papilla; white arrows indicate the anus; red circles indicate the anal-fin origin.
</paragraph>
</caption>
<paragraph id="8BB736BF12628201FF3AF9E9FE92F87D" blockId="7.[156,768,882,2077]" pageId="7" pageNumber="172">
<emphasis id="B97CEAAD12628201FF3AF9E9FE86F937" bold="true" box="[181,382,1662,1686]" pageId="7" pageNumber="172">Color in ethanol.</emphasis>
Head and body brown laterally, dark brown dorsally, and white or pale gray ventrally.
<specimenCount id="9D0EFD3612628201FD5CF908FF04F976" count="1" pageId="7" pageNumber="172" type="generic">One specimen</specimenCount>
(NSMT-P 143769) with a faint brown longitudinal stripe on the posterior half of the lateral midline of the body (
<figureCitation id="13332A3A12628201FF51F897FED2F8B9" box="[222,298,1792,1816]" captionStart="Fig" captionStartId="4.[156,187,1887,1909]" captionTargetBox="[207,1380,224,1751]" captionTargetId="figure-75@4.[206,1381,205,1848]" captionTargetPageId="4" captionText="Fig. 4. Specimens of Paloa villadolidi from Japan (A: URM-P 49659, 55.1mm SL; BE: NSMT-P 143769, 55.4mm SL). AD, Specimens immediately after fixation; E, preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.10833168" httpUri="https://zenodo.org/record/10833168/files/figure.png" pageId="7" pageNumber="172">Fig. 4E</figureCitation>
), while the other specimens lacked the stripe. Opercular region white edged. Markings of the fins similar to those of living specimens, but the orange and brown colors turned to gray or dusky brown. On urogenital papilla, males with a stellate black spot at center of papilla (
<figureCitation id="13332A3A12628201FD2AF815FD0BF83A" box="[677,755,1922,1947]" captionStart="Fig" captionStartId="7.[822,853,1391,1413]" captionTargetBox="[823,1429,901,1351]" captionTargetId="figure-508@7.[822,1430,900,1352]" captionTargetPageId="7" captionText="Fig. 8. Urogenital papilla of Paloa villadolidi. A, Male (NSMT- P 143769, 55.4mm SL); B, female (NSMT-P 145829, 75.0mm SL). Broken lines indicate the outer edge of urogenital papilla; white arrows indicate the anus; red circles indicate the anal-fin origin." figureDoi="http://doi.org/10.5281/zenodo.10833178" httpUri="https://zenodo.org/record/10833178/files/figure.png" pageId="7" pageNumber="172">Fig. 8A</figureCitation>
), females with a U shaped distinct line along the outer rim of the papilla (
<figureCitation id="13332A3A12628201FE9CF854FEA6F87D" box="[275,350,1987,2012]" captionStart="Fig" captionStartId="7.[822,853,1391,1413]" captionTargetBox="[823,1429,901,1351]" captionTargetId="figure-508@7.[822,1430,900,1352]" captionTargetPageId="7" captionText="Fig. 8. Urogenital papilla of Paloa villadolidi. A, Male (NSMT- P 143769, 55.4mm SL); B, female (NSMT-P 145829, 75.0mm SL). Broken lines indicate the outer edge of urogenital papilla; white arrows indicate the anus; red circles indicate the anal-fin origin." figureDoi="http://doi.org/10.5281/zenodo.10833178" httpUri="https://zenodo.org/record/10833178/files/figure.png" pageId="7" pageNumber="172">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31265341262820EFF3AF873FDA3FAD0" lastPageId="8" lastPageNumber="173" pageId="7" pageNumber="172" type="distribution">
<paragraph id="8BB736BF12628201FF3AF873FADAF994" blockId="7.[156,768,882,2077]" lastBlockId="7.[820,1432,1532,2076]" pageId="7" pageNumber="172">
<emphasis id="B97CEAAD12628201FF3AF873FEB9F85D" bold="true" box="[181,321,2020,2044]" pageId="7" pageNumber="172">Distribution.</emphasis>
<taxonomicName id="4C084D3C12628201FED5F873FDF9F85D" box="[346,513,2020,2044]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD12628201FED5F873FDF9F85D" box="[346,513,2020,2044]" italics="true" pageId="7" pageNumber="172">Paloa villadolidi</emphasis>
</taxonomicName>
is distributed in Luzon in the
<collectingCountry id="F31F762F12628201FF68F792FEA1F7BC" box="[231,345,2053,2077]" name="Philippines" pageId="7" pageNumber="172">Philippines</collectingCountry>
(
<bibRefCitation id="EF994B4E12628201FEE4F792FD98F7BC" author="Roxas, H. A. &amp; Ablan, G. L." box="[363,608,2053,2077]" pageId="7" pageNumber="172" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan 1940</bibRefCitation>
) and
<collectingRegion id="49CCF85D12628201FD2BF792FD07F7BC" box="[676,767,2053,2077]" country="Japan" name="Okinawa" pageId="7" pageNumber="172">Okinawa</collectingRegion>
and Ishigaki islands in
<collectingCountry id="F31F762F12628201FBAFFA6AFBA0F9B5" box="[1056,1112,1533,1556]" name="Japan" pageId="7" pageNumber="172">Japan</collectingCountry>
(this study).
<collectingRegion id="49CCF85D12628201FB6EFA6BFAC4F9B5" box="[1249,1340,1532,1556]" country="Japan" name="Okinawa" pageId="7" pageNumber="172">Okinawa</collectingRegion>
Island is the northernmost habitat known for this species.
</paragraph>
<paragraph id="8BB736BF1262820EFCC1F9AAFD0AFBEC" blockId="7.[820,1432,1532,2076]" lastBlockId="8.[156,767,979,1394]" lastPageId="8" lastPageNumber="173" pageId="7" pageNumber="172">
<emphasis id="B97CEAAD12628201FCC1F9AAFC5CF9F4" bold="true" box="[846,932,1597,1621]" pageId="7" pageNumber="172">Habitat.</emphasis>
All Japanese specimens were found in the tidal zone within or around mangroves in estuaries. Four of the
<specimenCount id="9D0EFD3612628201FCBBF9E8FC19F937" box="[820,993,1662,1686]" count="7" pageId="7" pageNumber="172" type="generic">seven specimens</specimenCount>
(NSMT-P 143769, 145829145831) were found hovering in small tidepools around the roots of mangrove trees [
<taxonomicName id="4C084D3C12628201FC39F957FB60F976" box="[950,1176,1727,1751]" class="Magnoliopsida" family="Rhizophoraceae" genus="Bruguiera" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="172" phylum="Tracheophyta" rank="species" species="gymnorhiza">
<emphasis id="B97CEAAD12628201FC39F957FB60F976" box="[950,1176,1727,1751]" italics="true" pageId="7" pageNumber="172">Bruguiera gymnorhiza</emphasis>
</taxonomicName>
(L.) Savigny] within the forest at night (
<figureCitation id="13332A3A12628201FC53F977FBE4F959" box="[988,1052,1760,1784]" captionStart="Fig" captionStartId="8.[156,187,906,928]" captionTargetBox="[208,1379,207,866]" captionTargetId="figure-616@8.[206,1381,205,868]" captionTargetPageId="8" captionText="Fig. 9. A tide pool in mangrove of Okinawa Island in Japan, where four specimens of Paloa villadolidi were collected." figureDoi="http://doi.org/10.5281/zenodo.10833180" httpUri="https://zenodo.org/record/10833180/files/figure.png" pageId="7" pageNumber="172">Fig. 9</figureCitation>
). The substrate of the tidepool was mud, and a thalassinid mud lobster,
<taxonomicName id="4C084D3C12628201FB45F897FC31F898" authority="(Herbst, 1804)" baseAuthorityName="Herbst" baseAuthorityYear="1804" class="Malacostraca" family="Thalassinidae" genus="Thalassina" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="172" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B97CEAAD12628201FB45F897FA6FF8B9" box="[1226,1431,1792,1816]" italics="true" pageId="7" pageNumber="172">Thalassina anomala</emphasis>
(Herbst, 1804)
</taxonomicName>
made a burrow system around the tidepool. Other fish species, including
<taxonomicName id="4C084D3C12628201FBECF8D6FC97F8DB" authority="(Lacepede, 1801)" baseAuthorityName="Lacepede" baseAuthorityYear="1801" family="Eleotridae" genus="Bostrychus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B97CEAAD12628201FBECF8D6FADAF8F8" box="[1123,1314,1857,1881]" italics="true" pageId="7" pageNumber="172">Bostrychus sinensis</emphasis>
(Lacepède, 1801)
</taxonomicName>
,
<taxonomicName id="4C084D3C12628201FC0BF8F5FB06F8DB" authority="Bleeker, 1853" authorityName="Bleeker" authorityYear="1853" box="[900,1278,1890,1914]" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="melanosoma">
<emphasis id="B97CEAAD12628201FC0BF8F5FBA3F8DB" box="[900,1115,1890,1914]" italics="true" pageId="7" pageNumber="172">Eleotris melanosoma</emphasis>
<bibRefCitation id="EF994B4E12628201FBE3F8F5FB06F8DB" author="Bleeker, P." box="[1132,1278,1890,1914]" pageId="7" pageNumber="172" pagination="1 - 166" refId="ref8056" refString="Bleeker, P. 1853. Nalezingen op de ichthyologische fauna van Bengalen en Hindostan. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 25 (8): 1 - 166." type="journal article" year="1853">Bleeker, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C084D3C12628201FA9DF8F5FBEEF83A" authority="(Smith, 1932)" baseAuthorityName="Smith" baseAuthorityYear="1932" family="Gobiidae" genus="Mugilogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="chulae">
<emphasis id="B97CEAAD12628201FA9DF8F5FC8CF83B" italics="true" pageId="7" pageNumber="172">Mugilogobius chulae</emphasis>
(Smith, 1932)
</taxonomicName>
, and
<taxonomicName id="4C084D3C12628201FBD3F815FC96F81A" authority="(Hamilton, 1822)" baseAuthorityName="Hamilton" baseAuthorityYear="1822" family="Ophichthidae" genus="Pisodonophis" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="boro">
<emphasis id="B97CEAAD12628201FBD3F815FAE2F83B" box="[1116,1306,1922,1946]" italics="true" pageId="7" pageNumber="172">Pisodonophis boro</emphasis>
(Hamilton, 1822)
</taxonomicName>
were observed in that tidepool. A juvenile specimen (URM-P 49661) was found in a similar tidepool on Ishigaki Island,
<collectingCountry id="F31F762F12628201FC12F873FC20F85A" box="[925,984,2020,2043]" name="Japan" pageId="7" pageNumber="172">Japan</collectingCountry>
. URM-P 49569 was found inside a polyvinyl chloride pipe buried in the muddy gravel bottom along the main course of the stream during the daytime. This site was approximately
<quantity id="4CF09B5A126D820EFEEEFC63FE69FBAA" box="[353,401,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="173" unit="m" value="10.0">10 m</quantity>
from the mangrove. URM-P 49660 was collected from the burrow of an unknown builder on a muddy substrate using a suction pump during the daytime.
</paragraph>
<caption id="DF776637126D820EFF13FC1DFB37FC01" ID-DOI="http://doi.org/10.5281/zenodo.10833180" ID-Zenodo-Dep="10833180" box="[156,1231,906,928]" httpUri="https://zenodo.org/record/10833180/files/figure.png" pageId="8" pageNumber="173" startId="8.[156,187,906,928]" targetBox="[208,1379,207,866]" targetPageId="8" targetType="figure">
<paragraph id="8BB736BF126D820EFF13FC1DFB37FC01" blockId="8.[156,1231,906,928]" box="[156,1231,906,928]" pageId="8" pageNumber="173">
Fig. 9. A tide pool in mangrove of Okinawa Island in Japan, where four specimens of
<taxonomicName id="4C084D3C126D820EFC3CFC1DFBBBFC3E" box="[947,1091,906,927]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="173" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD126D820EFC3CFC1DFBBBFC3E" box="[947,1091,906,927]" italics="true" pageId="8" pageNumber="173">Paloa villadolidi</emphasis>
</taxonomicName>
were collected.
</paragraph>
</caption>
<paragraph id="8BB736BF126D820EFF3AFBC1FDCAFB0F" blockId="8.[156,767,979,1394]" pageId="8" pageNumber="173">
<bibRefCitation id="EF994B4E126D820EFF3AFBC1FE54FBCC" author="Roxas, H. A. &amp; Ablan, G. L." box="[181,428,1109,1133]" pageId="8" pageNumber="173" pagination="301 - 311" refId="ref8948" refString="Roxas, H. A. and Ablan, G. L. 1940. New Philippine gobioid fishes. Philippine Journal of Science 73: 301 - 311, pls 1 - 8." type="journal article" year="1940">Roxas and Ablan (1940)</bibRefCitation>
did not report the habitat of this species. Therefore, no information is available regarding the habitat of this species in the
<collectingCountry id="F31F762F126D820EFE35FB01FDD7FB0F" box="[442,559,1174,1198]" name="Philippines" pageId="8" pageNumber="173">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BB736BF126D820EFF3AFB20FDA3FAD0" blockId="8.[156,767,979,1394]" pageId="8" pageNumber="173">
<emphasis id="B97CEAAD126D820EFF3AFB20FEA0FB6E" bold="true" box="[181,344,1207,1231]" pageId="8" pageNumber="173">Japanese name.</emphasis>
A new standard Japanese name “Shīsā-haze” is proposed for
<taxonomicName id="4C084D3C126D820EFEF9FB4FFE08FB4E" box="[374,496,1239,1263]" family="Eleotridae" genus="Paloa" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="173" phylum="Chordata" rank="species" species="villadolidi">
<emphasis id="B97CEAAD126D820EFEF9FB4FFE08FB4E" box="[374,496,1239,1263]" italics="true" pageId="8" pageNumber="173">P. villadolidi</emphasis>
</taxonomicName>
based on a specimen from
<collectingRegion id="49CCF85D126D820EFF13FB6FFF0FFAB1" box="[156,247,1272,1296]" country="Japan" name="Okinawa" pageId="8" pageNumber="173">Okinawa</collectingRegion>
Island (NSMT-P 143769). “Shīsā” is a traditional Ryukyuan artifact originating from Chinese guardian lion which has characteristic enlarged canine teeth like those of this species. “Haze” means goby in Japanese.
</paragraph>
</subSubSection>
</treatment>
</document>