treatments-xml/data/4B/60/C8/4B60C8363D1CFFACFF5894211125FC6D.xml

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<mods:title id="B0EF15375E64F53C43DB3122E5464CF7">Taito adrik, a new harvestman species from the Área de Conservación Privada Panguana, Peruvian Amazonia (Opiliones: Laniatores: Cosmetidae)</mods:title>
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(
<figureCitation id="5BF265A53D1CFFAAFF509406123DF9C2" box="[159,265,1589,1616]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1970,1995]" captionStartId-1="4.[151,250,1917,1942]" captionStartId-2="5.[151,250,1942,1967]" captionStartId-3="6.[151,250,1576,1601]" captionStartId-4="7.[151,250,1544,1569]" captionStartId-5="8.[151,250,1976,2001]" captionStartId-6="9.[151,250,1331,1356]" captionTargetBox-0="[151,1435,402,1850]" captionTargetBox-1="[159,1423,219,1807]" captionTargetBox-2="[166,1427,175,1911]" captionTargetBox-3="[151,1435,174,1556]" captionTargetBox-4="[158,1427,1045,1518]" captionTargetBox-5="[184,1402,1276,1948]" captionTargetBox-6="[151,1436,174,1312]" captionTargetId-0="figure@3.[151,1435,402,1850]" captionTargetId-1="figure@4.[151,1435,206,1821]" captionTargetId-2="figure@5.[151,1436,166,1923]" captionTargetId-3="figure@6.[151,1435,174,1556]" captionTargetId-4="figure@7.[151,1436,1037,1525]" captionTargetId-5="figure@8.[180,1408,1273,1956]" captionTargetId-6="figure@9.[151,1436,174,1312]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." captionText-1="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." captionText-2="FIGURE 3. Taito adrik sp. nov.: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm." captionText-3="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." captionText-4="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." captionText-5="FIGURE 6. Taito adrik sp. nov., representation of male genitalia, showing distal part of truncus, ventral plate with macrosetae, and glans: A. Ventral view; B. Lateral view; C. Dorsal view. Scale bar = 100 μm." captionText-6="FIGURE 7. Taito adrik sp. nov., S.E.M. micrographs of distal part of penis of two male paratypes (A–D, H: paratype ZSMA20190291, E–G: paratype ZSMA20190292):A. Ventral plate and distal part of truncus, ventral view; B. Lateral view; C. Glans and distal part of truncus, dorsal view (arrow shows small and straight D1, see also figure 7H); D. Dorsolateral view of glans; E./F. Lateral (E) and ventral (F) views of the specimen with large and curved D1 on one side and small and straight D1 on the other side (arrow, see also figure 7G); G. Detail of small and straight D1 in figure F; H. Detail of small and straight D1 in figure C. Scale bars: A–F = 20 µm; G, H = 2 µm." figureDoi-0="http://doi.org/10.5281/zenodo.3629058" figureDoi-1="http://doi.org/10.5281/zenodo.3629060" figureDoi-2="http://doi.org/10.5281/zenodo.3629062" figureDoi-3="http://doi.org/10.5281/zenodo.3629064" figureDoi-4="http://doi.org/10.5281/zenodo.3629066" figureDoi-5="http://doi.org/10.5281/zenodo.3629068" figureDoi-6="http://doi.org/10.5281/zenodo.3629070" httpUri-0="https://zenodo.org/record/3629058/files/figure.png" httpUri-1="https://zenodo.org/record/3629060/files/figure.png" httpUri-2="https://zenodo.org/record/3629062/files/figure.png" httpUri-3="https://zenodo.org/record/3629064/files/figure.png" httpUri-4="https://zenodo.org/record/3629066/files/figure.png" httpUri-5="https://zenodo.org/record/3629068/files/figure.png" httpUri-6="https://zenodo.org/record/3629070/files/figure.png" pageId="1" pageNumber="106">Figs. 1–7</figureCitation>
)
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<emphasis id="F1BDA5323D1CFFAAFF589491122AF92E" bold="true" box="[151,286,1698,1724]" pageId="1" pageNumber="106">Etymology.</emphasis>
The specific name is an indeclinable noun in apposition, given in honour of the outstanding arachnologist Adriano B. Kury (nickname “Adrik”), who established the genus 
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, together with Carla M.L. Barros, in 2014.As professor and senior curator of arachnids in the Museu Nacional/UFRJ, 
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, he lost most of his invaluable collections and fruits of his many field trips through the fire catastrophe in 
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.
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<emphasis id="F1BDA5323D1CFFAAFF089501124EF8DE" bold="true" box="[199,378,1841,1868]" pageId="1" pageNumber="106">Type material.</emphasis>
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; 
<materialsCitation id="73A1737D3D1CFFAAFC16954917F0F82A" ID-GBIF-Occurrence="2560338089" collectingDate="2013-09-20" collectingDateMax="2013-10-07" collectingDateMin="2013-09-20" collectionCode="MUSM, ZSMA" collectorName="S. Friedrich &amp; Wachtel" country="Peru" pageId="1" pageNumber="106" specimenCode="MUSM 0513706, ZSMA20190294, ZMH-A0002240" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1CFFAAFC1695491711F801" box="[985,1061,1913,1940]" pageId="1" pageNumber="106" type="male">3 male</specimenCount>
and 
<specimenCount id="D5CFB2A93D1CFFAAFB90954917F5F801" box="[1119,1217,1913,1940]" pageId="1" pageNumber="106" type="female">1 female</specimenCount>
<typeStatus id="1C72C7823D1CFFAAFB0595491601F806" box="[1226,1333,1914,1940]" pageId="1" pageNumber="106" type="paratype">paratypes</typeStatus>
(
<specimenCode id="936FD15B3D1CFFAAFA8A954913CFF82A" collectionCode="MUSM" pageId="1" pageNumber="106">MUSM 0513706</specimenCode>
/707, 
<specimenCode id="936FD15B3D1CFFAAFEF895AD12CCF82A" box="[311,504,1950,1976]" collectionCode="ZSMA" pageId="1" pageNumber="106">ZSMA20190294</specimenCode>
, 
<specimenCode id="936FD15B3D1CFFAAFDCE95AD11F7F82A" box="[513,707,1950,1976]" collectionCode="ZMH-A" pageId="1" pageNumber="106">ZMH-A0002240</specimenCode>
), 
<date id="B7775FE03D1CFFAAFD1B95AE10A6F825" box="[724,914,1949,1975]" pageId="1" pageNumber="106" value="2013-09-20" valueMax="2013-10-07" valueMin="2013-09-20">
<collectingDate id="A733A6083D1CFFAAFD1B95AE10A6F825" box="[724,914,1949,1975]" pageId="1" pageNumber="106" value="2013-09-20" valueMax="2013-10-07" valueMin="2013-09-20">20.ix.–07.x.2013</collectingDate>
</date>
(
<collectorName id="6E3C1CF63D1CFFAAFC6E95AD1717F825" box="[929,1059,1949,1976]" pageId="1" pageNumber="106">S. Friedrich</collectorName>
&amp; F. 
<collectorName id="6E3C1CF63D1CFFAAFB9495AE1788F82A" box="[1115,1212,1949,1976]" pageId="1" pageNumber="106">Wachtel</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="73A1737D3D1CFFAAFB0295AD173DF84E" ID-GBIF-Occurrence="2560338091" collectingDate="2015-05-01" collectingDateMax="2015-05-21" collectingDateMin="2015-05-01" collectionCode="ZSMA" collectorName="S. Friedrich &amp; F. Wachtel &amp; Steinherr" country="Peru" pageId="1" pageNumber="106" specimenCode="ZSMA20190289" specimenCount="2" specimenCount-female="2" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1CFFAAFB0295AD1618F825" box="[1229,1324,1949,1976]" pageId="1" pageNumber="106" type="female">2 female</specimenCount>
<typeStatus id="1C72C7823D1CFFAAFAFE95AD16A8F82A" box="[1329,1436,1950,1976]" pageId="1" pageNumber="106" type="paratype">paratypes</typeStatus>
(
<specimenCode id="936FD15B3D1CFFAAFF5095F11255F84E" box="[159,353,1986,2012]" collectionCode="ZSMA" pageId="1" pageNumber="106">ZSMA20190289</specimenCode>
/290), 
<date id="B7775FE03D1CFFAAFE6A95F21170F84E" box="[421,580,1985,2012]" pageId="1" pageNumber="106" value="2015-05-01" valueMax="2015-05-21" valueMin="2015-05-01">
<collectingDate id="A733A6083D1CFFAAFE6A95F21170F84E" box="[421,580,1985,2012]" pageId="1" pageNumber="106" value="2015-05-01" valueMax="2015-05-21" valueMin="2015-05-01">01.–21.v.2015</collectingDate>
</date>
(
<collectorName id="6E3C1CF63D1CFFAAFD9D95F111E3F849" box="[594,727,1985,2012]" pageId="1" pageNumber="106">S. Friedrich</collectorName>
, 
<collectorName id="6E3C1CF63D1CFFAAFD1095F11066F849" box="[735,850,1985,2012]" pageId="1" pageNumber="106">F.Wachtel</collectorName>
&amp; M. 
<collectorName id="6E3C1CF63D1CFFAAFC5A95F21735F84E" box="[917,1025,1985,2012]" pageId="1" pageNumber="106">Steinherr</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="73A1737D3D1CFFA9FBDE95F11731FF26" ID-GBIF-Occurrence="2560338090" collectingDate="2016-04-23" collectingDateMax="2016-05-09" collectingDateMin="2016-04-23" collectionCode="ZMH-A" collectorName="S. Friedrich &amp; F. Wachtel &amp; Hauth" country="Peru" lastPageId="2" lastPageNumber="107" pageId="1" pageNumber="106" specimenCode="ZMH-A0002241" specimenCount="2" specimenCount-male="2" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1CFFAAFBDE95F1176EF849" box="[1041,1114,1985,2012]" pageId="1" pageNumber="106" type="male">2 male</specimenCount>
<typeStatus id="1C72C7823D1CFFAAFB9195F117FDF84E" box="[1118,1225,1986,2012]" pageId="1" pageNumber="106" type="paratype">paratypes</typeStatus>
(
<specimenCode id="936FD15B3D1CFFAAFB1995F116ACF84E" box="[1238,1432,1986,2012]" collectionCode="ZMH-A" pageId="1" pageNumber="106">ZMH-A0002241</specimenCode>
, MHNG-ARTO-18641), 
<date id="B7775FE03D1FFFA9FE6A92AA116BFF21" box="[421,607,153,179]" pageId="2" pageNumber="107" value="2016-04-23" valueMax="2016-05-09" valueMin="2016-04-23">
<collectingDate id="A733A6083D1FFFA9FE6A92AA116BFF21" box="[421,607,153,179]" pageId="2" pageNumber="107" value="2016-04-23" valueMax="2016-05-09" valueMin="2016-04-23">23.iv.–09.v.2016</collectingDate>
</date>
(
<collectorName id="6E3C1CF63D1FFFA9FDA092AA11C1FF21" box="[623,757,153,179]" pageId="2" pageNumber="107">S. Friedrich</collectorName>
, 
<collectorName id="6E3C1CF63D1FFFA9FD3192AA1046FF21" box="[766,882,153,179]" pageId="2" pageNumber="107">F. Wachtel</collectorName>
&amp; D. 
<collectorName id="6E3C1CF63D1FFFA9FC7B92AA10C9FF26" box="[948,1021,153,180]" pageId="2" pageNumber="107">Hauth</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="73A1737D3D1FFFA9FBC092AA109DFF4A" ID-GBIF-Occurrence="2560338092" collectingDate="2017-09-22" collectingDateMax="2017-10-10" collectingDateMin="2017-09-22" collectionCode="T" collectorName="S. Friedrich &amp; F. Wachtel &amp; Hauth &amp; Lehmann" country="Peru" pageId="2" pageNumber="107" specimenCode="ZSMA20190296" specimenCount="1" specimenCount-male="1" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1FFFA9FBC092AA176DFF21" box="[1039,1113,153,179]" pageId="2" pageNumber="107" type="male">1 male</specimenCount>
<typeStatus id="1C72C7823D1FFFA9FB9092AA178BFF21" box="[1119,1215,153,179]" pageId="2" pageNumber="107" type="paratype">paratype</typeStatus>
(
<specimenCode id="936FD15B3D1FFFA9FB0392AA16A4FF26" box="[1228,1424,153,180]" collectionCode="ZSMA" pageId="2" pageNumber="107">ZSMA20190296</specimenCode>
), 
<date id="B7775FE03D1FFFA9FF58928E1261FF45" box="[151,341,189,215]" pageId="2" pageNumber="107" value="2017-09-22" valueMax="2017-10-10" valueMin="2017-09-22">
<collectingDate id="A733A6083D1FFFA9FF58928E1261FF45" box="[151,341,189,215]" pageId="2" pageNumber="107" value="2017-09-22" valueMax="2017-10-10" valueMin="2017-09-22">22.ix.–10.x.2017</collectingDate>
</date>
(
<collectorName id="6E3C1CF63D1FFFA9FEA7928E12C4FF45" box="[360,496,189,215]" pageId="2" pageNumber="107">S. Friedrich</collectorName>
, 
<collectorName id="6E3C1CF63D1FFFA9FE32928E1143FF45" box="[509,631,189,215]" pageId="2" pageNumber="107">F. Wachtel</collectorName>
, D. 
<collectorName id="6E3C1CF63D1FFFA9FD67928E11D8FF45" box="[680,748,189,215]" pageId="2" pageNumber="107">Hauth</collectorName>
&amp; 
<collectionCode id="A5D8E1E53D1FFFA9FCDC928E1011FF45" box="[787,805,189,215]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="107">T</collectionCode>
. 
<collectorName id="6E3C1CF63D1FFFA9FCFD928E1095FF4A" box="[818,929,189,216]" pageId="2" pageNumber="107">Lehmann</collectorName>
)
</materialsCitation>
; 
<specimenCount id="D5CFB2A93D1FFFA9FC79928E172CFF45" box="[950,1048,189,215]" pageId="2" pageNumber="107" type="female">1 female</specimenCount>
<typeStatus id="1C72C7823D1FFFA9FBEE928E17B5FF45" box="[1057,1153,189,215]" pageId="2" pageNumber="107" type="paratype">paratype</typeStatus>
(MHNG-ARTO-18642), 
<date id="B7775FE03D1FFFA9FF5892D21261FF69" box="[151,341,225,251]" pageId="2" pageNumber="107" value="2017-09-22" valueMax="2017-10-10" valueMin="2017-09-22">22.ix.–10.x.2017</date>
(E.-G. Burmeister); 
<materialsCitation id="73A1737D3D1FFFA9FD8F92D212B5FEB2" ID-GBIF-Occurrence="2560338095" collectingDate="2018-04-21" collectingDateMax="2018-05-04" collectingDateMin="2018-04-21" collectionCode="MNRJ, ZSMA" collectorName="Diller" country="Peru" pageId="2" pageNumber="107" specimenCode="MNRJ 6033-036, ZSMA20190297" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1FFFA9FD8F92D211BFFF69" box="[576,651,225,251]" pageId="2" pageNumber="107" type="male">2 male</specimenCount>
and 
<specimenCount id="D5CFB2A93D1FFFA9FD0C92D21011FF69" box="[707,805,225,251]" pageId="2" pageNumber="107" type="female">3 female</specimenCount>
<typeStatus id="1C72C7823D1FFFA9FCE292D210ACFF69" box="[813,920,225,251]" pageId="2" pageNumber="107" type="paratype">paratypes</typeStatus>
(
<specimenCode id="936FD15B3D1FFFA9FC6892D2175AFF69" box="[935,1134,225,251]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="107">MNRJ 6033–036</specimenCode>
, 
<specimenCode id="936FD15B3D1FFFA9FBB592D21609FF69" box="[1146,1341,225,251]" collectionCode="ZSMA" pageId="2" pageNumber="107">ZSMA20190297</specimenCode>
), 
<date id="B7775FE03D1FFFA9FA9F92D21232FE8D" pageId="2" pageNumber="107" value="2018-04-21" valueMax="2018-05-04" valueMin="2018-04-21">
<collectingDate id="A733A6083D1FFFA9FA9F92D21232FE8D" pageId="2" pageNumber="107" value="2018-04-21" valueMax="2018-05-04" valueMin="2018-04-21">21.iv.– 04.v.2018</collectingDate>
</date>
(E. 
<collectorName id="6E3C1CF63D1FFFA9FEFC9336124DFEB2" box="[307,377,261,288]" pageId="2" pageNumber="107">Diller</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="73A1737D3D1FFFA9FE4393361253FEF5" ID-GBIF-Occurrence="2560338094" collectingDate="2018-08-24" collectingDateMax="2018-09-06" collectingDateMin="2018-08-24" collectionCode="ZSMA, MUSM" collectorName="E. Diller &amp; All" country="Peru" pageId="2" pageNumber="107" specimenCode="ZSMA20190291-293, MUSM 0513701-705" specimenCount="18" specimenCount-female="11" specimenCount-male="7" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="D5CFB2A93D1FFFA9FE43933612E2FE8D" box="[396,470,261,287]" pageId="2" pageNumber="107" type="male">7 male</specimenCount>
and 
<specimenCount id="D5CFB2A93D1FFFA9FDC49336114DFE8D" box="[523,633,261,287]" pageId="2" pageNumber="107" type="female">11 female</specimenCount>
<typeStatus id="1C72C7823D1FFFA9FDB0933611DEFE8D" box="[639,746,261,287]" pageId="2" pageNumber="107" type="paratype">paratypes</typeStatus>
(
<specimenCode id="936FD15B3D1FFFA9FD37933610C5FE8D" box="[760,1009,261,287]" collectionCode="ZSMA" pageId="2" pageNumber="107">ZSMA20190291–293</specimenCode>
/298–304, 
<specimenCode id="936FD15B3D1FFFA9FBA893361668FE8D" box="[1127,1372,260,287]" collectionCode="MUSM" pageId="2" pageNumber="107">MUSM 0513701–705</specimenCode>
/708– 710), 
<date id="B7775FE03D1FFFA9FF13931A1284FED1" box="[220,432,297,323]" pageId="2" pageNumber="107" value="2018-08-24" valueMax="2018-09-06" valueMin="2018-08-24">
<collectingDate id="A733A6083D1FFFA9FF13931A1284FED1" box="[220,432,297,323]" pageId="2" pageNumber="107" value="2018-08-24" valueMax="2018-09-06" valueMin="2018-08-24">24.viii.–06.ix.2018</collectingDate>
</date>
(
<collectorName id="6E3C1CF63D1FFFA9FE0B931A1119FED6" box="[452,557,297,324]" pageId="2" pageNumber="107">E. Diller</collectorName>
). 
<collectorName id="6E3C1CF63D1FFFA9FD8E931A1151FED1" box="[577,613,297,323]" pageId="2" pageNumber="107">All</collectorName>
specimens were collected at night on the ground in primary evergreen lowland rainforest
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BD32AAB3D1FFFA9FF0893431189FD15" pageId="2" pageNumber="107" type="diagnosis">
<paragraph id="C37679203D1FFFA9FF0893431189FD15" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0893431276FE19" bold="true" box="[199,322,368,395]" pageId="2" pageNumber="107">Diagnosis.</emphasis>
Dorsal scutum sturdy (average ratio length/width 
<emphasis id="F1BDA5323D1FFFA9FC4893411095FE19" box="[903,929,370,395]" italics="true" pageId="2" pageNumber="107">ca</emphasis>
. 
<quantity id="0431D4C53D1FFFA9FC7D93421737FE19" box="[946,1027,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="2.9972" pageId="2" pageNumber="107" unit="in" value="1.18">1.18 in</quantity>
females, 
<emphasis id="F1BDA5323D1FFFA9FBBB934117BAFE19" box="[1140,1166,370,395]" italics="true" pageId="2" pageNumber="107">ca</emphasis>
. 
<quantity id="0431D4C53D1FFFA9FB50934217C4FE19" box="[1183,1264,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="2" pageNumber="107" unit="in" value="1.21">1.21 in</quantity>
males), β-type (normal β) (
<bibRefCitation id="A75804D13D1FFFA9FED293A612ECFE3D" author="Kury, A. B. &amp; Villarreal M. &amp; Sampaio, C." box="[285,472,405,431]" pageId="2" pageNumber="107" pagination="325 - 333" refId="ref5548" refString="Kury, A. B., Villarreal M., O. &amp; Sampaio, C. (2007) Redescription of the type species of Cynorta (Arachnida, Opiliones, Cosmetidae). The Journal of Arachnology, 35 (2), 325 - 333. https: // doi. org / 10.1636 / H 06 - 35.1" type="journal article" year="2007">
Kury 
<emphasis id="F1BDA5323D1FFFA9FE9093A512A3FE3D" box="[351,407,405,431]" italics="true" pageId="2" pageNumber="107">et al.</emphasis>
2007
</bibRefCitation>
) and not βL-type (elongate β) (
<bibRefCitation id="A75804D13D1FFFA9FCFA93A6170EFE22" author="Medrano, M. A. &amp; Kury, A. B." box="[821,1082,405,432]" pageId="2" pageNumber="107" pagination="52 - 62" refId="ref5888" refString="Medrano, M. A. &amp; Kury, A. B. (2016) Characterization of Platymessa with redescription of the type species and a new generic synonymy (Arachnida, Opiliones, Cosmetidae). Zootaxa, 4085 (1), 52 - 62. https: // doi. org / 10.11646 / zootaxa. 4085.1.2" type="journal article" year="2016">Kury &amp; Medrano 2016</bibRefCitation>
) (
<figureCitation id="5BF265A53D1FFFA9FB9F93A61784FE3D" box="[1104,1200,405,431]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Fig. 2 A</figureCitation>
). Equuleus butterflyshaped with discrete feet and arms, and elongated, slim horns (
<figureCitation id="5BF265A53D1FFFA9FCAC938A10FBFE41" box="[867,975,441,467]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1995]" captionTargetBox="[151,1435,402,1850]" captionTargetId="figure@3.[151,1435,402,1850]" captionTargetPageId="3" captionText="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="2" pageNumber="107">Figs. 1 A</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FC13938A171FFE41" box="[988,1067,441,467]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">2 A, C</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FBF8938A1772FE41" box="[1079,1094,441,467]" captionStart="FIGURE 3" captionStartId="5.[151,250,1942,1967]" captionTargetBox="[166,1427,175,1911]" captionTargetId="figure@5.[151,1436,166,1923]" captionTargetPageId="5" captionText="FIGURE 3. Taito adrik sp. nov.: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629062" httpUri="https://zenodo.org/record/3629062/files/figure.png" pageId="2" pageNumber="107">3</figureCitation>
); two whitish blots on minute tubercles at posterior margin of scutal area III (in some specimens two smaller additional blots beside) (
<figureCitation id="5BF265A53D1FFFA9FAE393EE16A3FE65" box="[1324,1431,477,503]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1995]" captionTargetBox="[151,1435,402,1850]" captionTargetId="figure@3.[151,1435,402,1850]" captionTargetPageId="3" captionText="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="2" pageNumber="107">Figs. 1 A</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FF58903213D1FD89" box="[151,229,513,539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">2 A, C</figureCitation>
). Groin warts (
<bibRefCitation id="A75804D13D1FFFA9FE5F9032114AFD89" author="Kury, A. B. &amp; Barros, C. M." box="[400,638,513,539]" pageId="2" pageNumber="107" pagination="1 - 46" refId="ref5656" refString="Kury, A. B. &amp; Barros, C. M. (2014) A new genus and eight new species of Amazonian cosmetines (Opiliones, Laniatores, Cosmetidae). Zoological Studies, 53 (24), 1 - 46. https: // doi. org / 10.1186 / s 40555 - 014 - 0024 - 4" type="journal article" year="2014">Kury &amp; Barros 2014</bibRefCitation>
) on dorso-proximal part of coxa IV (
<figureCitation id="5BF265A53D1FFFA9FBEB90321792FD89" box="[1060,1190,513,539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Fig. 2 A, B</figureCitation>
), coxa IV with dorsal triangular apophysis (
<figureCitation id="5BF265A53D1FFFA9FE45901612D2FDAD" box="[394,486,549,575]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Fig. 2 F</figureCitation>
). Leg IV of male: femur IV gently curved with comb of five to six curved apophyses on distal-prolateral side; patella IV with coarse granules and one single apophysis proximal-retrolateral; tibia IV unarmed, covered with granules (
<figureCitation id="5BF265A53D1FFFA9FDC1905E11A0FD15" box="[526,660,621,647]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Figs. 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FD50905F119BFD15" box="[671,687,620,647]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="2" pageNumber="107">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BD32AAB3D1FFFA9FF0890A310E9FC85" pageId="2" pageNumber="107" type="distribution">
<paragraph id="C37679203D1FFFA9FF0890A310E9FC85" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0890A31255FD39" bold="true" box="[199,353,656,683]" pageId="2" pageNumber="107">Distribution.</emphasis>
Only known from the 
<typeStatus id="1C72C7823D1FFFA9FDA790A211ACFD39" box="[616,664,657,683]" pageId="2" pageNumber="107">type</typeStatus>
locality (
<figureCitation id="5BF265A53D1FFFA9FCCA90A21053FD39" box="[773,871,657,683]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1995]" captionTargetBox="[151,1435,402,1850]" captionTargetId="figure@3.[151,1435,402,1850]" captionTargetPageId="3" captionText="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="2" pageNumber="107">Fig. 1 B</figureCitation>
). This extends the distribution range of the genus 
<taxonomicName id="04C902A33D1FFFA9FF58908513E4FD5D" authorityName="Kury &amp; Barros" authorityYear="2014" box="[151,208,694,719]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1FFFA9FF58908513E4FD5D" box="[151,208,694,719]" italics="true" pageId="2" pageNumber="107">Taito</emphasis>
</taxonomicName>
to the south-west. So far, the genus was known from the Upper Amazon Basin up the eastern slope of the Andes. Now, the distribution reaches to a territory west of the 
<collectingRegion id="010DB7C23D1FFFA9FCE190EA10F8FD61" box="[814,972,729,755]" country="Peru" name="Ucayali" pageId="2" pageNumber="107">Ucayali River</collectingRegion>
, into the Pachitea Basin, which is located between the eastern slopes of the Andes and the Sira Mountains (
<figureCitation id="5BF265A53D1FFFA9FCBF90CE10FBFC85" box="[880,975,765,791]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1995]" captionTargetBox="[151,1435,402,1850]" captionTargetId="figure@3.[151,1435,402,1850]" captionTargetPageId="3" captionText="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="2" pageNumber="107">Fig. 1 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8BD32AAB3D1FFFACFF0891131125FC6D" lastPageId="7" lastPageNumber="112" pageId="2" pageNumber="107" type="description">
<paragraph id="C37679203D1FFFA9FF08911311EEFCA9" blockId="2.[151,1437,153,1670]" box="[199,730,800,827]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF089113126DFCA9" bold="true" box="[199,345,800,827]" pageId="2" pageNumber="107">Description.</emphasis>
Male 
<typeStatus id="1C72C7823D1FFFA9FE6F91121136FCA9" box="[416,514,801,827]" pageId="2" pageNumber="107" type="holotype">holotype</typeStatus>
(MUSM 0513700)
</paragraph>
<paragraph id="C37679203D1FFFA9FF0891761070FC3A" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF089176123CFCCD" box="[199,264,837,863]" italics="true" pageId="2" pageNumber="107">Color</emphasis>
(in ethanol) (
<figureCitation id="5BF265A53D1FFFA9FE68917612DAFCCD" box="[423,494,837,863]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Fig. 2</figureCitation>
). Body medium brown, densely covered with small, lighter orange-brown islands. Appendages also showing this pattern, but a little bit lighter. Equuleus on dorsal scutum and interrupted stripe on coda nearly white. Shape of equuleus very consistent (
<figureCitation id="5BF265A53D1FFFA9FD3691BE100CFC3A" box="[761,824,909,936]" captionStart="FIGURE 3" captionStartId="5.[151,250,1942,1967]" captionTargetBox="[166,1427,175,1911]" captionTargetId="figure@5.[151,1436,166,1923]" captionTargetPageId="5" captionText="FIGURE 3. Taito adrik sp. nov.: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629062" httpUri="https://zenodo.org/record/3629062/files/figure.png" pageId="2" pageNumber="107">Fig.3</figureCitation>
).
</paragraph>
<paragraph id="C37679203D1FFFA9FF0891811700FC59" blockId="2.[151,1437,153,1670]" box="[199,1076,944,971]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0891811245FC59" box="[199,369,946,971]" italics="true" pageId="2" pageNumber="107">Measurements.</emphasis>
CL = 2.3, CW = 3.6, AL = 4.2, AW = 5.7; legs I to IV: 
<tableCitation id="8E4B4C9B3D1FFFA9FC1491821705FC59" box="[987,1073,945,971]" captionStart="TABLE 2" captionStartId="2.[151,239,1717,1741]" captionTargetPageId="2" captionText="TABLE 2. Taito adrik sp. nov., male holotype (MUSM 0513700): Articles measurements of left legs (in mm)." httpUri="http://table.plazi.org/id/97B629A83D1FFFA9FF5894861606F95F" pageId="2" pageNumber="107" tableUuid="97B629A83D1FFFA9FF5894861606F95F">Table 2</tableCitation>
.
</paragraph>
<paragraph id="C37679203D1FFFA9FF0891E51041FBED" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0891E51215FC7D" box="[199,289,982,1007]" italics="true" pageId="2" pageNumber="107">Dorsum</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FEFE91E61288FC7D" box="[305,444,981,1007]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Figs. 2 A, D</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FE0791E612C4FC7D" box="[456,496,981,1007]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="2" pageNumber="107">4 F</figureCitation>
). Dorsal scutum sturdy (ratio 
<emphasis id="F1BDA5323D1FFFA9FCF291E5106BFC7D" box="[829,863,982,1007]" italics="true" pageId="2" pageNumber="107">ca.</emphasis>
1.19), maximum width at scutal area III, thickened laterals with granulate surface; two minute tubercles on area III. Posterior margin of scutum sub-straight in dorsal view. Posterior margin of free tergite I also sub-straight, that of free tergite II weekly convex, and of free tergite III strongly convex (
<figureCitation id="5BF265A53D1FFFA9FE9496721288FBC9" box="[347,444,1089,1115]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Fig. 2 A</figureCitation>
). All free tergites with a row of transverse granules. Anal operculum unarmed, with two dorso-lateral indentations, covered with granules (
<figureCitation id="5BF265A53D1FFFA9FD0696561000FBED" box="[713,820,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Figs. 2 D</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FC8F9656105CFBED" box="[832,872,1125,1151]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="2" pageNumber="107">4 F</figureCitation>
).
</paragraph>
<paragraph id="C37679203D1FFFA9FF0896B910DEFA82" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0896B9123BFB31" box="[199,271,1162,1187]" italics="true" pageId="2" pageNumber="107">Venter</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FED096BA12BFFB31" box="[287,395,1161,1187]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Figs. 2 B</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FE5696BA12F7FB31" box="[409,451,1161,1187]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="2" pageNumber="107">4 F</figureCitation>
). Coxae I–II nearly transverse, parallel and sub-equal in size. Coxa III larger. Coxa IV much stronger and slanted backwards. Anterior part of genital operculum elliptical, posterior margin truncated. Stigmatic area y-shaped with large elliptical stigmata. Stigmatic area deeply concave relative to the greatly bulging coxa IV. Free sternites I–VI strongly concave, with a dense row of granules.
</paragraph>
<paragraph id="C37679203D1FFFA9FF08972F120EFA12" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF08972F1276FAA4" box="[199,322,1308,1334]" italics="true" pageId="2" pageNumber="107">Chelicerae</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FE9E97281299FAA4" box="[337,429,1307,1334]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="2" pageNumber="107">Fig. 4 E</figureCitation>
). Basichelicerite with an ectal row of six tubercles; three anterior tubercles large and connected, and a posterior row of 12 granulate tubercles with nearly equal size. One single large and rounded anteromesal process.
</paragraph>
<paragraph id="C37679203D1FFFA9FF0897BF1671FA59" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0897BF1203FA34" box="[199,311,1420,1446]" italics="true" pageId="2" pageNumber="107">Pedipalps</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FE8997B812B3FA34" box="[326,391,1419,1446]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="2" pageNumber="107">Fig. 4</figureCitation>
A–D). Trochanter with a single ventral spine. Femur with dorsal proximal keel with two anterior teeth, ventral row of 17 teeth, the middle one largest. Tibia with an apophysis at distal end, near tarsus.
</paragraph>
<paragraph id="C37679203D1FFFA9FF0897E4124CF914" blockId="2.[151,1437,153,1670]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF0897E413C8FA62" box="[199,252,1495,1520]" italics="true" pageId="2" pageNumber="107">Legs</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FEC297E512AEFA63" box="[269,410,1494,1521]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="2" pageNumber="107">Figs. 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1FFFA9FE6797E6128CFA62" box="[424,440,1493,1520]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="2" pageNumber="107">5</figureCitation>
). Legs I–III granulous and unarmed. Femur IV gently curved both laterally and dorsoventrally, finely granulous; it is armed with a prolateral comb of six posteriorly curved spine-shaped apophyses, at distal end. Patella IV covered with coarse granulation and armed with a single retrolateral apophysis at the proximal end. Tibia IV unarmed with granulous surface. Tarsal count: 6–6/14–14/7–7/9–9. Measurements of leg segments provided in 
<tableCitation id="8E4B4C9B3D1FFFA9FED194581240F914" box="[286,372,1643,1670]" captionStart="TABLE 2" captionStartId="2.[151,239,1717,1741]" captionTargetPageId="2" captionText="TABLE 2. Taito adrik sp. nov., male holotype (MUSM 0513700): Articles measurements of left legs (in mm)." httpUri="http://table.plazi.org/id/97B629A83D1FFFA9FF5894861606F95F" pageId="2" pageNumber="107" tableUuid="97B629A83D1FFFA9FF5894861606F95F">Table 2</tableCitation>
.
</paragraph>
<caption id="97B629A83D1FFFA9FF5894861606F95F" ID-Table-UUID="97B629A83D1FFFA9FF5894861606F95F" box="[151,1330,1717,1741]" httpUri="http://table.plazi.org/id/97B629A83D1FFFA9FF5894861606F95F" pageId="2" pageNumber="107" targetBox="[166,1339,1753,1929]" targetIsTable="true" targetPageId="2">
<paragraph id="C37679203D1FFFA9FF5894861606F95F" blockId="2.[151,1330,1717,1741]" box="[151,1330,1717,1741]" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF589486123EF95F" bold="true" box="[151,266,1717,1741]" pageId="2" pageNumber="107">TABLE 2.</emphasis>
<taxonomicName id="04C902A33D1FFFA9FEDE948512B1F95F" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[273,389,1717,1741]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1FFFA9FEDE948512B1F95F" box="[273,389,1717,1741]" italics="true" pageId="2" pageNumber="107">Taito adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1FFFA9FE43948612D6F95F" bold="true" box="[396,482,1717,1741]" pageId="2" pageNumber="107">
<taxonomicNameLabel id="EA8E18493D1FFFA9FE43948612D6F95F" box="[396,482,1717,1741]" pageId="2" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (MUSM 0513700): Articles measurements of left legs (in mm).
</paragraph>
</caption>
<paragraph id="C37679203D1FFFA9FF6394EA162DF81B" pageId="2" pageNumber="107">
<table id="B1C98B803D1F0054FF6994EA160FF81B" box="[166,1339,1753,1929]" gridcols="7" gridrows="5" pageId="2" pageNumber="107">
<tr id="7DF97B623D1F0054FF6994EA160FF960" box="[166,1339,1753,1778]" gridrow="0" pageId="2" pageNumber="107">
<th id="3E28121E3D1F0054FF6994EA13E1F960" box="[166,213,1753,1778]" gridcol="0" gridrow="0" pageId="2" pageNumber="107">legs</th>
<th id="3E28121E3D1F0054FEA794EA12E2F960" box="[360,470,1753,1778]" gridcol="1" gridrow="0" pageId="2" pageNumber="107">Trochanter</th>
<th id="3E28121E3D1F0054FD8894EA11BDF960" box="[583,649,1753,1778]" gridcol="2" gridrow="0" pageId="2" pageNumber="107">Femur</th>
<th id="3E28121E3D1F0054FD3694EA100AF960" box="[761,830,1753,1778]" gridcol="3" gridrow="0" pageId="2" pageNumber="107">Patella</th>
<th id="3E28121E3D1F0054FC6494EA10D4F960" box="[939,992,1753,1778]" gridcol="4" gridrow="0" pageId="2" pageNumber="107">Tibia</th>
<th id="3E28121E3D1F0054FBFE94EA1794F960" box="[1073,1184,1753,1778]" gridcol="5" gridrow="0" pageId="2" pageNumber="107">Metatarsus</th>
<th id="3E28121E3D1F0054FB3694EA160FF960" box="[1273,1339,1753,1778]" gridcol="6" gridrow="0" pageId="2" pageNumber="107">Tarsus</th>
</tr>
<tr id="7DF97B623D1F0054FF699533160FF88A" box="[166,1339,1792,1816]" gridrow="1" pageId="2" pageNumber="107">
<th id="3E28121E3D1F0054FF69953313E1F88A" box="[166,213,1792,1816]" gridcol="0" gridrow="1" pageId="2" pageNumber="107">I</th>
<td id="3E28121E3D1F0054FEA7953312E2F88A" box="[360,470,1792,1816]" gridcol="1" gridrow="1" pageId="2" pageNumber="107">1.0</td>
<td id="3E28121E3D1F0054FD88953311BDF88A" box="[583,649,1792,1816]" gridcol="2" gridrow="1" pageId="2" pageNumber="107">4.8</td>
<td id="3E28121E3D1F0054FD369533100AF88A" box="[761,830,1792,1816]" gridcol="3" gridrow="1" pageId="2" pageNumber="107">1.2</td>
<td id="3E28121E3D1F0054FC64953310D4F88A" box="[939,992,1792,1816]" gridcol="4" gridrow="1" pageId="2" pageNumber="107">3.0</td>
<td id="3E28121E3D1F0054FBFE95331794F88A" box="[1073,1184,1792,1816]" gridcol="5" gridrow="1" pageId="2" pageNumber="107">4.8</td>
<td id="3E28121E3D1F0054FB369533160FF88A" box="[1273,1339,1792,1816]" gridcol="6" gridrow="1" pageId="2" pageNumber="107">2.9</td>
</tr>
<tr id="7DF97B623D1F0054FF699516160FF8AC" box="[166,1339,1829,1854]" gridrow="2" pageId="2" pageNumber="107">
<th id="3E28121E3D1F0054FF69951613E1F8AC" box="[166,213,1829,1854]" gridcol="0" gridrow="2" pageId="2" pageNumber="107">II</th>
<td id="3E28121E3D1F0054FEA7951612E2F8AC" box="[360,470,1829,1854]" gridcol="1" gridrow="2" pageId="2" pageNumber="107">1.4</td>
<td id="3E28121E3D1F0054FD88951611BDF8AC" box="[583,649,1829,1854]" gridcol="2" gridrow="2" pageId="2" pageNumber="107">9.7</td>
<td id="3E28121E3D1F0054FD369516100AF8AC" box="[761,830,1829,1854]" gridcol="3" gridrow="2" pageId="2" pageNumber="107">1.5</td>
<td id="3E28121E3D1F0054FC64951610D4F8AC" box="[939,992,1829,1854]" gridcol="4" gridrow="2" pageId="2" pageNumber="107">7.9</td>
<td id="3E28121E3D1F0054FBFE95161794F8AC" box="[1073,1184,1829,1854]" gridcol="5" gridrow="2" pageId="2" pageNumber="107">10.1</td>
<td id="3E28121E3D1F0054FB369516160FF8AC" box="[1273,1339,1829,1854]" gridcol="6" gridrow="2" pageId="2" pageNumber="107">5.9</td>
</tr>
<tr id="7DF97B623D1F0054FF699579160FF8F1" box="[166,1339,1866,1891]" gridrow="3" pageId="2" pageNumber="107">
<th id="3E28121E3D1F0054FF69957913E1F8F1" box="[166,213,1866,1891]" gridcol="0" gridrow="3" pageId="2" pageNumber="107">III</th>
<td id="3E28121E3D1F0054FEA7957912E2F8F1" box="[360,470,1866,1891]" gridcol="1" gridrow="3" pageId="2" pageNumber="107">1.3</td>
<td id="3E28121E3D1F0054FD88957911BDF8F1" box="[583,649,1866,1891]" gridcol="2" gridrow="3" pageId="2" pageNumber="107">6.9</td>
<td id="3E28121E3D1F0054FD369579100AF8F1" box="[761,830,1866,1891]" gridcol="3" gridrow="3" pageId="2" pageNumber="107">1.6</td>
<td id="3E28121E3D1F0054FC64957910D4F8F1" box="[939,992,1866,1891]" gridcol="4" gridrow="3" pageId="2" pageNumber="107">3.8</td>
<td id="3E28121E3D1F0054FBFE95791794F8F1" box="[1073,1184,1866,1891]" gridcol="5" gridrow="3" pageId="2" pageNumber="107">6.2</td>
<td id="3E28121E3D1F0054FB369579160FF8F1" box="[1273,1339,1866,1891]" gridcol="6" gridrow="3" pageId="2" pageNumber="107">3.3</td>
</tr>
<tr id="7DF97B623D1F0054FF699543160FF81B" box="[166,1339,1904,1929]" gridrow="4" pageId="2" pageNumber="107">
<th id="3E28121E3D1F0054FF69954313E1F81B" box="[166,213,1904,1929]" gridcol="0" gridrow="4" pageId="2" pageNumber="107">IV</th>
<td id="3E28121E3D1F0054FEA7954312E2F81B" box="[360,470,1904,1929]" gridcol="1" gridrow="4" pageId="2" pageNumber="107">1.5</td>
<td id="3E28121E3D1F0054FD88954311BDF81B" box="[583,649,1904,1929]" gridcol="2" gridrow="4" pageId="2" pageNumber="107">9.0</td>
<td id="3E28121E3D1F0054FD369543100AF81B" box="[761,830,1904,1929]" gridcol="3" gridrow="4" pageId="2" pageNumber="107">1.8</td>
<td id="3E28121E3D1F0054FC64954310D4F81B" box="[939,992,1904,1929]" gridcol="4" gridrow="4" pageId="2" pageNumber="107">5.6</td>
<td id="3E28121E3D1F0054FBFE95431794F81B" box="[1073,1184,1904,1929]" gridcol="5" gridrow="4" pageId="2" pageNumber="107">9.2</td>
<td id="3E28121E3D1F0054FB369543160FF81B" box="[1273,1339,1904,1929]" gridcol="6" gridrow="4" pageId="2" pageNumber="107">4.2</td>
</tr>
</table>
</paragraph>
<paragraph id="C37679203D1FFFA8FF08958912B8FE8D" blockId="2.[151,1436,1978,2040]" lastBlockId="3.[151,1437,153,287]" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107">
<emphasis id="F1BDA5323D1FFFA9FF089589125FF846" box="[199,363,1978,2004]" italics="true" pageId="2" pageNumber="107">Male genitalia</emphasis>
(
<figureCitation id="5BF265A53D1FFFA9FEB6958912F2F846" box="[377,454,1978,2004]" captionStart="FIGURE 6" captionStartId="8.[151,250,1976,2001]" captionTargetBox="[184,1402,1276,1948]" captionTargetId="figure@8.[180,1408,1273,1956]" captionTargetPageId="8" captionText="FIGURE 6. Taito adrik sp. nov., representation of male genitalia, showing distal part of truncus, ventral plate with macrosetae, and glans: A. Ventral view; B. Lateral view; C. Dorsal view. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3629068" httpUri="https://zenodo.org/record/3629068/files/figure.png" pageId="2" pageNumber="107">Figs. 6</figureCitation>
A–C; 7 A–H). Penis: slender trunk with distal bulge. Dorsal process of glans thumb-like and smooth. Glans sac with cellulite texture. Stylus covered with spiny tubercles on its ventral border. Ventral plate with macrosetae C1 and C2 large, curved and spatulate. D1 short and straight, or large and curved in a 
<typeStatus id="1C72C7823D1EFFA8FAC692AA165DFF21" box="[1289,1385,153,179]" pageId="3" pageNumber="108" type="paratype">paratype</typeStatus>
(see Discussion). D2 reduced. Only one pair of macrosetae A and B. A small and straight, B reduced, E1 and E2 reduced. Microsetae (ms) on ventral plate with a wide central gap and made up of type 4 ms distally, which gradually become type 1 ms proximally.
</paragraph>
<caption id="97B629A83D1EFFA8FF58958110A1F87D" ID-DOI="http://doi.org/10.5281/zenodo.3629058" ID-Zenodo-Dep="3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="3" pageNumber="108" startId="3.[151,250,1970,1995]" targetBox="[151,1435,402,1850]" targetPageId="3">
<paragraph id="C37679203D1EFFA8FF58958110A1F87D" blockId="3.[151,1437,1970,2031]" pageId="3" pageNumber="108">
<emphasis id="F1BDA5323D1EFFA8FF5895811227F859" bold="true" box="[151,275,1970,1995]" pageId="3" pageNumber="108">FIGURE 1.</emphasis>
A. Living female paratype (ZSMA20190293) of 
<taxonomicName id="04C902A33D1EFFA8FCC995801040F858" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[774,884,1970,1994]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1EFFA8FCC995801040F858" box="[774,884,1970,1994]" italics="true" pageId="3" pageNumber="108">Taito adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1EFFA8FCB4958110FFF858" bold="true" box="[891,971,1970,1994]" pageId="3" pageNumber="108">
<taxonomicNameLabel id="EA8E18493D1EFFA8FCB4958110FFF858" box="[891,971,1970,1994]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality.
</paragraph>
</caption>
<caption id="97B629A83D19FFAFFF58954E100EF84F" ID-DOI="http://doi.org/10.5281/zenodo.3629060" ID-Zenodo-Dep="3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="4" pageNumber="109" startId="4.[151,250,1917,1942]" targetBox="[159,1423,219,1807]" targetPageId="4">
<paragraph id="C37679203D19FFAFFF58954E100EF84F" blockId="4.[151,1437,1917,2014]" pageId="4" pageNumber="109">
<emphasis id="F1BDA5323D19FFAFFF58954E1227F804" bold="true" box="[151,275,1917,1942]" pageId="4" pageNumber="109">FIGURE 2.</emphasis>
<taxonomicName id="04C902A33D19FFAFFED6954D12B3F807" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[281,391,1917,1941]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D19FFAFFED6954D12B3F807" box="[281,391,1917,1941]" italics="true" pageId="4" pageNumber="109">Taito adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D19FFAFFE41954E12EAF807" bold="true" box="[398,478,1917,1941]" pageId="4" pageNumber="109">
<taxonomicNameLabel id="EA8E18493D19FFAFFE41954E12EAF807" box="[398,478,1917,1941]" pageId="4" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm.
</paragraph>
</caption>
<caption id="97B629A83D18FFAEFF5895A51259F865" ID-DOI="http://doi.org/10.5281/zenodo.3629062" ID-Zenodo-Dep="3629062" httpUri="https://zenodo.org/record/3629062/files/figure.png" pageId="5" pageNumber="110" startId="5.[151,250,1942,1967]" targetBox="[166,1427,175,1911]" targetPageId="5">
<paragraph id="C37679203D18FFAEFF5895A51259F865" blockId="5.[151,1437,1942,2039]" pageId="5" pageNumber="110">
<emphasis id="F1BDA5323D18FFAEFF5895A51220F83D" bold="true" box="[151,276,1942,1967]" pageId="5" pageNumber="110">FIGURE 3.</emphasis>
<taxonomicName id="04C902A33D18FFAEFED495AB12BEF83D" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[283,394,1943,1967]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="110" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D18FFAEFED495AB12BEF83D" box="[283,394,1943,1967]" italics="true" pageId="5" pageNumber="110">Taito adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D18FFAEFE5E95A412D6F83D" bold="true" box="[401,482,1943,1967]" pageId="5" pageNumber="110">
<taxonomicNameLabel id="EA8E18493D18FFAEFE5E95A412D6F83D" box="[401,482,1943,1967]" pageId="5" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm.
</paragraph>
</caption>
<caption id="97B629A83D1BFFADFF58941B17C4F91B" ID-DOI="http://doi.org/10.5281/zenodo.3629064" ID-Zenodo-Dep="3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="6" pageNumber="111" startId="6.[151,250,1576,1601]" targetBox="[151,1435,174,1556]" targetPageId="6">
<paragraph id="C37679203D1BFFADFF58941B17C4F91B" blockId="6.[151,1437,1576,1673]" pageId="6" pageNumber="111">
<emphasis id="F1BDA5323D1BFFADFF58941B1226F9D3" bold="true" box="[151,274,1576,1601]" pageId="6" pageNumber="111">FIGURE 4.</emphasis>
<taxonomicName id="04C902A33D1BFFADFED8941A12B1F9D3" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[279,389,1577,1601]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFED8941A12B1F9D3" box="[279,389,1577,1601]" italics="true" pageId="6" pageNumber="111">Taito adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFE44941A12EEF9D3" bold="true" box="[395,474,1577,1601]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFE44941A12EEF9D3" box="[395,474,1577,1601]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (MUSM 0513700): A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm.
</paragraph>
</caption>
<paragraph id="C37679203D1BFFACFF089485163FFDF1" blockId="6.[151,1437,1718,2034]" lastBlockId="7.[151,1437,153,683]" lastPageId="7" lastPageNumber="112" pageId="6" pageNumber="111">
<emphasis id="F1BDA5323D1BFFADFF0894851259F943" bold="true" box="[199,365,1718,1745]" pageId="6" pageNumber="111">Comparisons.</emphasis>
In several 
<taxonomicName id="04C902A33D1BFFADFE25948B1117F943" authorityName="Kury &amp; Barros" authorityYear="2014" box="[490,547,1720,1745]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1BFFADFE25948B1117F943" box="[490,547,1720,1745]" italics="true" pageId="6" pageNumber="111">Taito</emphasis>
</taxonomicName>
species, femur IV is armed with a row/comb of a few distal spines; in 
<taxonomicName id="04C902A33D1BFFADFA8E948B138BF967" authorityName="Kury &amp; Barros" authorityYear="2014" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="medinae">
<emphasis id="F1BDA5323D1BFFADFA8E948B138BF967" italics="true" pageId="6" pageNumber="111">T. medinae</emphasis>
</taxonomicName>
, there are eight spines, while in 
<taxonomicName id="04C902A33D1BFFADFDF994EF11C7F967" authorityName="Kury &amp; Barros" authorityYear="2014" box="[566,755,1755,1781]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="spaceinvaders">
<emphasis id="F1BDA5323D1BFFADFDF994EF11C7F967" box="[566,755,1755,1781]" italics="true" pageId="6" pageNumber="111">T. spaceinvaders</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFCCD94EF105AF967" authorityName="Kury &amp; Barros" authorityYear="2014" box="[770,878,1756,1781]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="osmari">
<emphasis id="F1BDA5323D1BFFADFCCD94EF105AF967" box="[770,878,1756,1781]" italics="true" pageId="6" pageNumber="111">T. osmari</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFCB294EF173BF967" baseAuthorityName="Soares" baseAuthorityYear="1970" box="[893,1039,1756,1781]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="insperatus">
<emphasis id="F1BDA5323D1BFFADFCB294EF173BF967" box="[893,1039,1756,1781]" italics="true" pageId="6" pageNumber="111">T. insperatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1BFFADFB8094EF17DCF967" box="[1103,1256,1755,1781]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="oblongatus">
<emphasis id="F1BDA5323D1BFFADFB8094EF17DCF967" box="[1103,1256,1755,1781]" italics="true" pageId="6" pageNumber="111">T. oblongatus</emphasis>
</taxonomicName>
there are seven spines. Only in 
<taxonomicName id="04C902A33D1BFFADFE89953312A8F88B" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[326,412,1791,1817]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFE89953312A8F88B" box="[326,412,1791,1817]" italics="true" pageId="6" pageNumber="111">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFE6C94CC12C9F88B" bold="true" box="[419,509,1791,1817]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFE6C94CC12C9F88B" box="[419,509,1791,1817]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the number of distal spines is six (
<figureCitation id="5BF265A53D1BFFADFC4F94CC10CFF888" box="[896,1019,1791,1818]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="6" pageNumber="111">Fig. 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1BFFADFBC994CD1722F88B" box="[1030,1046,1790,1817]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="6" pageNumber="111">5</figureCitation>
). Generally, the armature of leg IV in 
<taxonomicName id="04C902A33D1BFFADFF7C9517123DF8AF" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[179,265,1827,1853]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFF7C9517123DF8AF" box="[179,265,1827,1853]" italics="true" pageId="6" pageNumber="111">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFEDF9510125DF8AF" bold="true" box="[272,361,1827,1853]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFEDF9510125DF8AF" box="[272,361,1827,1853]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to two closely related species, 
<taxonomicName id="04C902A33D1BFFADFCE09517108BF8AF" baseAuthorityName="Soares" baseAuthorityYear="1970" box="[815,959,1828,1853]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="insperatus">
<emphasis id="F1BDA5323D1BFFADFCE09517108BF8AF" box="[815,959,1828,1853]" italics="true" pageId="6" pageNumber="111">T. insperatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1BFFADFC3B951717BFF8AF" box="[1012,1163,1827,1853]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="oblongatus">
<emphasis id="F1BDA5323D1BFFADFC3B951717BFF8AF" box="[1012,1163,1827,1853]" italics="true" pageId="6" pageNumber="111">T. oblongatus</emphasis>
</taxonomicName>
. However, these species differ in having dorsal scutum elongated, while it is sturdy in 
<taxonomicName id="04C902A33D1BFFADFC96957B1085F8F3" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[857,945,1863,1889]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFC96957B1085F8F3" box="[857,945,1863,1889]" italics="true" pageId="6" pageNumber="111">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFC7695741720F8F3" bold="true" box="[953,1044,1863,1889]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFC7695741720F8F3" box="[953,1044,1863,1889]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, a feature only shared with 
<taxonomicName id="04C902A33D1BFFADFA9B957B13DFF817" class="Arachnida" family="Cosmetidae" genus="Cynortula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="serriperna">
<emphasis id="F1BDA5323D1BFFADFA9B957B13DFF817" italics="true" pageId="6" pageNumber="111">T. serriperna</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1BFFADFEEF955F1291F817" authorityName="Kury &amp; Barros" authorityYear="2014" box="[288,421,1899,1925]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="kawaiikei">
<emphasis id="F1BDA5323D1BFFADFEEF955F1291F817" box="[288,421,1899,1925]" italics="true" pageId="6" pageNumber="111">T. kawaiikei</emphasis>
</taxonomicName>
. These species in turn are easily separated from 
<taxonomicName id="04C902A33D1BFFADFC71955F1727F817" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[958,1043,1899,1925]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFC71955F1727F817" box="[958,1043,1899,1925]" italics="true" pageId="6" pageNumber="111">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFBD695581746F817" bold="true" box="[1049,1138,1899,1925]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFBD695581746F817" box="[1049,1138,1899,1925]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
especially by the armature of leg IV (apophyses on coxa, femur, and patella). With respect to coxa IV, in 
<taxonomicName id="04C902A33D1BFFADFBCE95A31763F83B" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[1025,1111,1935,1961]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1BFFADFBCE95A31763F83B" box="[1025,1111,1935,1961]" italics="true" pageId="6" pageNumber="111">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1BFFADFB9195BC178CF83B" bold="true" box="[1118,1208,1935,1961]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="EA8E18493D1BFFADFB9195BC178CF83B" box="[1118,1208,1935,1961]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it has a single small apophysis (
<figureCitation id="5BF265A53D1BFFADFED9958012A4F85C" box="[278,400,1971,1998]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="6" pageNumber="111">Fig. 2 A, F</figureCitation>
), just like in 
<taxonomicName id="04C902A33D1BFFADFDEF95871184F85F" baseAuthorityName="Soares" baseAuthorityYear="1970" box="[544,688,1972,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="insperatus">
<emphasis id="F1BDA5323D1BFFADFDEF95871184F85F" box="[544,688,1972,1997]" italics="true" pageId="6" pageNumber="111">T. insperatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFD7295871010F85F" authorityName="Kury &amp; Barros" authorityYear="2014" box="[701,804,1971,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="kakera">
<emphasis id="F1BDA5323D1BFFADFD7295871010F85F" box="[701,804,1971,1997]" italics="true" pageId="6" pageNumber="111">T. kakera</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFCFE9587109FF85F" authorityName="Kury &amp; Barros" authorityYear="2014" box="[817,939,1971,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="medinae">
<emphasis id="F1BDA5323D1BFFADFCFE9587109FF85F" box="[817,939,1971,1997]" italics="true" pageId="6" pageNumber="111">T. medinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFC779587177BF85F" box="[952,1103,1971,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="oblongatus">
<emphasis id="F1BDA5323D1BFFADFC779587177BF85F" box="[952,1103,1971,1997]" italics="true" pageId="6" pageNumber="111">T. oblongatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFB92958717F2F85F" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1117,1222,1972,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="osmari">
<emphasis id="F1BDA5323D1BFFADFB92958717F2F85F" box="[1117,1222,1972,1997]" italics="true" pageId="6" pageNumber="111">T. osmari</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFB1C95871652F85F" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1235,1382,1971,1997]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="rorschachi">
<emphasis id="F1BDA5323D1BFFADFB1C95871652F85F" box="[1235,1382,1971,1997]" italics="true" pageId="6" pageNumber="111">T. rorschachi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1BFFADFF5895EB1267F863" authorityName="Kury &amp; Barros" authorityYear="2014" box="[151,339,2007,2033]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="spaceinvaders">
<emphasis id="F1BDA5323D1BFFADFF5895EB1267F863" box="[151,339,2007,2033]" italics="true" pageId="6" pageNumber="111">T. spaceinvaders</emphasis>
</taxonomicName>
; in contrast, 
<taxonomicName id="04C902A33D1BFFADFE2795EB1166F863" authorityName="Kury &amp; Barros" authorityYear="2014" box="[488,594,2007,2033]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1BFFADFE2795EB1166F863" box="[488,594,2007,2033]" italics="true" pageId="6" pageNumber="111">T. galaga</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1BFFADFD4695EB11D8F863" authorityName="Kury &amp; Barros" authorityYear="2014" box="[649,748,2007,2033]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="honda">
<emphasis id="F1BDA5323D1BFFADFD4695EB11D8F863" box="[649,748,2007,2033]" italics="true" pageId="6" pageNumber="111">T. honda</emphasis>
</taxonomicName>
have a single large apophysis, and 
<taxonomicName id="04C902A33D1BFFADFBB395EB17CBF863" box="[1148,1279,2008,2033]" class="Arachnida" family="Cosmetidae" genus="Eucynortella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="juruensis">
<emphasis id="F1BDA5323D1BFFADFBB395EB17CBF863" box="[1148,1279,2008,2033]" italics="true" pageId="6" pageNumber="111">T. juruensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1BFFADFAC195EB16A1F863" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1294,1429,2007,2033]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="kawaiikei">
<emphasis id="F1BDA5323D1BFFADFAC195EB16A1F863" box="[1294,1429,2007,2033]" italics="true" pageId="6" pageNumber="111">T. kawaiikei</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1AFFACFF0792A9126FFF21" box="[200,347,154,179]" class="Arachnida" family="Cosmetidae" genus="Cynortula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="serriperna">
<emphasis id="F1BDA5323D1AFFACFF0792A9126FFF21" box="[200,347,154,179]" italics="true" pageId="7" pageNumber="112">T. serriperna</emphasis>
</taxonomicName>
have a double apophysis. Patella IV is unarmed in most species; in a few species, this article is covered with acuminate/setiferous tubercles (
<taxonomicName id="04C902A33D1AFFACFD57928D1037FF45" authorityName="Kury &amp; Barros" authorityYear="2014" box="[664,771,189,215]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFD57928D1037FF45" box="[664,771,189,215]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFCDD928D104FFF45" authorityName="Kury &amp; Barros" authorityYear="2014" box="[786,891,189,215]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="kakera">
<emphasis id="F1BDA5323D1AFFACFCDD928D104FFF45" box="[786,891,189,215]" italics="true" pageId="7" pageNumber="112">T. kakera</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFC45928D10C1FF45" authorityName="Kury &amp; Barros" authorityYear="2014" box="[906,1013,190,215]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="osmari">
<emphasis id="F1BDA5323D1AFFACFC45928D10C1FF45" box="[906,1013,190,215]" italics="true" pageId="7" pageNumber="112">T. osmari</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1AFFACFBFB928D17FDFF45" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1076,1225,189,215]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="rorschachi">
<emphasis id="F1BDA5323D1AFFACFBFB928D17FDFF45" box="[1076,1225,189,215]" italics="true" pageId="7" pageNumber="112">T. rorschachi</emphasis>
</taxonomicName>
). 
<taxonomicName id="04C902A33D1AFFACFB2D928D160DFF45" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[1250,1337,189,215]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFB2D928D160DFF45" box="[1250,1337,189,215]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFA8D928E16A8FF45" bold="true" box="[1346,1436,189,215]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFA8D928E16A8FF45" box="[1346,1436,189,215]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only species with a single retrolateral conspicuous apophysis (
<figureCitation id="5BF265A53D1AFFACFC5C92D2173BFF69" box="[915,1039,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig. 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFBD592D3171EFF69" box="[1050,1066,224,251]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="7" pageNumber="112">5</figureCitation>
). 
<taxonomicName id="04C902A33D1AFFACFB8F92D1179DFF69" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1088,1193,225,251]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFB8F92D1179DFF69" box="[1088,1193,225,251]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1AFFACFB1092D11647FF69" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1247,1395,225,251]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="rorschachi">
<emphasis id="F1BDA5323D1AFFACFB1092D11647FF69" box="[1247,1395,225,251]" italics="true" pageId="7" pageNumber="112">T. rorschachi</emphasis>
</taxonomicName>
are the only species that share a butterfly-shaped equuleus with the new species. In 
<taxonomicName id="04C902A33D1AFFACFBD093351742FE8D" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[1055,1142,261,287]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFBD093351742FE8D" box="[1055,1142,261,287]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFBB1933617ECFE8D" bold="true" box="[1150,1240,261,287]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFBB1933617ECFE8D" box="[1150,1240,261,287]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the design of the butterfly is most similar to 
<taxonomicName id="04C902A33D1AFFACFE0E9319111DFED1" authorityName="Kury &amp; Barros" authorityYear="2014" box="[449,553,297,323]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFE0E9319111DFED1" box="[449,553,297,323]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
, but only in the new species the equuleus has elongated, slim horns reaching the ocularium (
<figureCitation id="5BF265A53D1AFFACFED8937E1241FEF5" box="[279,373,333,359]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1995]" captionTargetBox="[151,1435,402,1850]" captionTargetId="figure@3.[151,1435,402,1850]" captionTargetPageId="3" captionText="FIGURE 1. A. Living female paratype (ZSMA20190293) of Taito adrik sp. nov. from the type locality, ACP Panguana (photo: Konrad Wothe); B. Location of the type locality; C. Habitat at type locality." figureDoi="http://doi.org/10.5281/zenodo.3629058" httpUri="https://zenodo.org/record/3629058/files/figure.png" pageId="7" pageNumber="112">Fig. 1 A</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFE4F937E12FEFEF5" box="[384,458,333,359]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">2 A, C</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFE1A937E12D1FEF5" box="[469,485,333,359]" captionStart="FIGURE 3" captionStartId="5.[151,250,1942,1967]" captionTargetBox="[166,1427,175,1911]" captionTargetId="figure@5.[151,1436,166,1923]" captionTargetPageId="5" captionText="FIGURE 3. Taito adrik sp. nov.: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629062" httpUri="https://zenodo.org/record/3629062/files/figure.png" pageId="7" pageNumber="112">3</figureCitation>
). Moreover, 
<taxonomicName id="04C902A33D1AFFACFDBB937D11FEFEF5" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[628,714,333,359]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFDBB937D11FEFEF5" box="[628,714,333,359]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFD1F937E101DFEF5" bold="true" box="[720,809,333,359]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFD1F937E101DFEF5" box="[720,809,333,359]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="04C902A33D1AFFACFC75937D1717FEF5" authorityName="Kury &amp; Barros" authorityYear="2014" box="[954,1059,333,359]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFC75937D1717FEF5" box="[954,1059,333,359]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1AFFACFB97937D17DFFEF5" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1112,1259,333,359]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="rorschachi">
<emphasis id="F1BDA5323D1AFFACFB97937D17DFFEF5" box="[1112,1259,333,359]" italics="true" pageId="7" pageNumber="112">T. rorschachi</emphasis>
</taxonomicName>
in several other features (scutal area III, anal operculum, and armature of leg IV). In the genus 
<taxonomicName id="04C902A33D1AFFACFBC193411773FE19" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1038,1095,370,395]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1AFFACFBC193411773FE19" box="[1038,1095,370,395]" italics="true" pageId="7" pageNumber="112">Taito</emphasis>
</taxonomicName>
, scutal area III has either high spines (
<taxonomicName id="04C902A33D1AFFACFF2093A5126EFE3D" authorityName="Kury &amp; Barros" authorityYear="2014" box="[239,346,405,431]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFF2093A5126EFE3D" box="[239,346,405,431]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
and 
<taxonomicName id="04C902A33D1AFFACFE5C93A512C2FE3D" authorityName="Kury &amp; Barros" authorityYear="2014" box="[403,502,405,431]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="honda">
<emphasis id="F1BDA5323D1AFFACFE5C93A512C2FE3D" box="[403,502,405,431]" italics="true" pageId="7" pageNumber="112">T. honda</emphasis>
</taxonomicName>
) or small granules/minute tubercles (all other species, including 
<taxonomicName id="04C902A33D1AFFACFB1593A51606FE3D" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[1242,1330,405,431]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFB1593A51606FE3D" box="[1242,1330,405,431]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFAF593A616A1FE3D" bold="true" box="[1338,1429,405,431]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFAF593A616A1FE3D" box="[1338,1429,405,431]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="5BF265A53D1AFFACFF58938A120DFE41" box="[151,313,441,467]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig. 2 A, C, D</figureCitation>
). Finally, the dorsal anal operculum is either unarmed (
<taxonomicName id="04C902A33D1AFFACFC69938910C8FE41" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[934,1020,441,467]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFC69938910C8FE41" box="[934,1020,441,467]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFBCB938A1769FE41" bold="true" box="[1028,1117,441,467]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFBCB938A1769FE41" box="[1028,1117,441,467]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="5BF265A53D1AFFACFBA4938A17FFFE41" box="[1131,1227,441,467]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig. 2 D</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFB18938A17CAFE41" box="[1239,1278,441,467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1576,1601]" captionTargetBox="[151,1435,174,1556]" captionTargetId="figure@6.[151,1435,174,1556]" captionTargetPageId="6" captionText="FIGURE 4. Taito adrik sp. nov., male holotype (MUSM 0513700):A. Left pedipalp: mesal view; B. ectal view; C. patella-tibia, dorsal view and femur lateral view; D. tibia-tarsus, dorsal view and femur-patella, lateral view; E. Left chelicera: dorsal view of basichelicerite; F. Anal operculum and free sternites, posterior view. Scale bars: A–D = 1 mm; E, F = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.3629064" httpUri="https://zenodo.org/record/3629064/files/figure.png" pageId="7" pageNumber="112">4 F</figureCitation>
), as well as 
<taxonomicName id="04C902A33D1AFFACFA47938913D7FE65" authorityName="Kury &amp; Barros" authorityYear="2014" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="kakera">
<emphasis id="F1BDA5323D1AFFACFA47938913D7FE65" italics="true" pageId="7" pageNumber="112">T. kakera</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFF3F93ED1265FE65" authorityName="Kury &amp; Barros" authorityYear="2014" box="[240,337,477,503]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="honda">
<emphasis id="F1BDA5323D1AFFACFF3F93ED1265FE65" box="[240,337,477,503]" italics="true" pageId="7" pageNumber="112">T. honda</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFE9193ED12DAFE65" baseAuthorityName="Soares" baseAuthorityYear="1970" box="[350,494,478,503]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="insperatus">
<emphasis id="F1BDA5323D1AFFACFE9193ED12DAFE65" box="[350,494,478,503]" italics="true" pageId="7" pageNumber="112">T. insperatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFE3493ED11A5FE65" box="[507,657,477,503]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="oblongatus">
<emphasis id="F1BDA5323D1AFFACFE3493ED11A5FE65" box="[507,657,477,503]" italics="true" pageId="7" pageNumber="112">T. oblongatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFD5093ED1010FE65" authorityName="Kury &amp; Barros" authorityYear="2014" box="[671,804,477,503]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="kawaiikei">
<emphasis id="F1BDA5323D1AFFACFD5093ED1010FE65" box="[671,804,477,503]" italics="true" pageId="7" pageNumber="112">T. kawaiikei</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFCFD93ED1087FE65" box="[818,947,478,503]" class="Arachnida" family="Cosmetidae" genus="Eucynortella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="juruensis">
<emphasis id="F1BDA5323D1AFFACFCFD93ED1087FE65" box="[818,947,478,503]" italics="true" pageId="7" pageNumber="112">T. juruensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1AFFACFC2093ED174BFE65" box="[1007,1151,478,503]" class="Arachnida" family="Cosmetidae" genus="Cynortula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="serriperna">
<emphasis id="F1BDA5323D1AFFACFC2093ED174BFE65" box="[1007,1151,478,503]" italics="true" pageId="7" pageNumber="112">T. serriperna</emphasis>
</taxonomicName>
), or armed with a median spine, a blunt protuberance or a crown of tubercles (
<taxonomicName id="04C902A33D1AFFACFD2690311092FD89" authorityName="Kury &amp; Barros" authorityYear="2014" box="[745,934,513,539]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="spaceinvaders">
<emphasis id="F1BDA5323D1AFFACFD2690311092FD89" box="[745,934,513,539]" italics="true" pageId="7" pageNumber="112">T. spaceinvaders</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFC7A90311714FD89" authorityName="Kury &amp; Barros" authorityYear="2014" box="[949,1056,513,539]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="galaga">
<emphasis id="F1BDA5323D1AFFACFC7A90311714FD89" box="[949,1056,513,539]" italics="true" pageId="7" pageNumber="112">T. galaga</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFBE0903117A5FD89" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1071,1169,513,539]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="honda">
<emphasis id="F1BDA5323D1AFFACFBE0903117A5FD89" box="[1071,1169,513,539]" italics="true" pageId="7" pageNumber="112">T. honda</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFB6F90311628FD89" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1184,1308,513,539]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="medinae">
<emphasis id="F1BDA5323D1AFFACFB6F90311628FD89" box="[1184,1308,513,539]" italics="true" pageId="7" pageNumber="112">T. medinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFAE4903116A1FD89" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1323,1429,514,539]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="osmari">
<emphasis id="F1BDA5323D1AFFACFAE4903116A1FD89" box="[1323,1429,514,539]" italics="true" pageId="7" pageNumber="112">T. osmari</emphasis>
</taxonomicName>
, 
<taxonomicName id="04C902A33D1AFFACFF5890151218FDAD" authorityName="Kury &amp; Barros" authorityYear="2014" box="[151,300,549,575]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="rorschachi">
<emphasis id="F1BDA5323D1AFFACFF5890151218FDAD" box="[151,300,549,575]" italics="true" pageId="7" pageNumber="112">T. rorschachi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="04C902A33D1AFFACFEA39015111DFDAD" authorityName="Kury &amp; Barros" authorityYear="2014" box="[364,553,549,575]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="spaceinvaders">
<emphasis id="F1BDA5323D1AFFACFEA39015111DFDAD" box="[364,553,549,575]" italics="true" pageId="7" pageNumber="112">T. spaceinvaders</emphasis>
</taxonomicName>
); however, only in 
<taxonomicName id="04C902A33D1AFFACFCC590151056FDAD" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[778,866,549,575]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFCC590151056FDAD" box="[778,866,549,575]" italics="true" pageId="7" pageNumber="112">T. adrik</emphasis>
</taxonomicName>
<emphasis id="F1BDA5323D1AFFACFCA5901610F1FDAD" bold="true" box="[874,965,549,575]" pageId="7" pageNumber="112">
<taxonomicNameLabel id="EA8E18493D1AFFACFCA5901610F1FDAD" box="[874,965,549,575]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the anal operculum has two dorso-lateral indentations, which can be seen with the SEM and were not described in all other 
<taxonomicName id="04C902A33D1AFFACFBE190791753FDF1" authorityName="Kury &amp; Barros" authorityYear="2014" box="[1070,1127,586,611]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1AFFACFBE190791753FDF1" box="[1070,1127,586,611]" italics="true" pageId="7" pageNumber="112">Taito</emphasis>
</taxonomicName>
species so far.
</paragraph>
<paragraph id="C37679203D1AFFACFF08905E17BCFD39" blockId="7.[151,1437,153,683]" pageId="7" pageNumber="112">
The new species can be inserted in the key of 
<bibRefCitation id="A75804D13D1AFFACFD01905E10DAFD15" author="Kury, A. B. &amp; Barros, C. M." box="[718,1006,621,647]" pageId="7" pageNumber="112" pagination="1 - 46" refId="ref5656" refString="Kury, A. B. &amp; Barros, C. M. (2014) A new genus and eight new species of Amazonian cosmetines (Opiliones, Laniatores, Cosmetidae). Zoological Studies, 53 (24), 1 - 46. https: // doi. org / 10.1186 / s 40555 - 014 - 0024 - 4" type="journal article" year="2014">Kury &amp; Barros (2014:20)</bibRefCitation>
, at the dichotomy 7(2) (then adjusting the numbering of dichotomies by changing the former 7(2) to 8(7), and so on), as follows:
</paragraph>
<paragraph id="C37679203D1AFFACFF5890E516A8FCB7" blockId="7.[151,1436,725,806]" pageId="7" pageNumber="112">
7(2) Equuleus butterfly-shaped with elongated, slim horns (
<figureCitation id="5BF265A53D1AFFACFD3D90E51077FD79" box="[754,835,726,748]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig. 2 A</figureCitation>
, 
<figureCitation id="5BF265A53D1AFFACFC8190E5106EFD79" box="[846,858,726,747]" captionStart="FIGURE 3" captionStartId="5.[151,250,1942,1967]" captionTargetBox="[166,1427,175,1911]" captionTargetId="figure@5.[151,1436,166,1923]" captionTargetPageId="5" captionText="FIGURE 3. Taito adrik sp. nov.: Outline of equulei of all specimens of the type series (aa: male holotype (MUSM 0513700), ab–ao: male paratypes, ba–bs: female paratypes, c: juvenile male paratype ZSMA20190294), showing minimal variation. Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629062" httpUri="https://zenodo.org/record/3629062/files/figure.png" pageId="7" pageNumber="112">3</figureCitation>
aa–bs); femur IV armed with a comb of 6 distal spines (
<figureCitation id="5BF265A53D1AFFACFAB490E5122BFC9A" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFEE690C01203FC9B" box="[297,311,755,777]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="7" pageNumber="112">5</figureCitation>
); patella IV armed with a single retrolateral conspicuous apophysis (
<figureCitation id="5BF265A53D1AFFACFC0690C01719FC9A" box="[969,1069,755,777]" captionStart="FIGURE 2" captionStartId="4.[151,250,1917,1942]" captionTargetBox="[159,1423,219,1807]" captionTargetId="figure@4.[151,1435,206,1821]" captionTargetPageId="4" captionText="FIGURE 2. Taito adrik sp. nov., holotype male (MUSM 0513700): A. Dorsal view; B. Ventral view; C. Left side, lateral view; D. Posterior view; E. Left leg IV, dorsal view. F. Left leg IV, prolateral view. (c: coda, co: coxa IV, e: equuleus, fe: femur IV, pa: patella IV, ti: tibia IV, tr: trochanter IV). Overall scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629060" httpUri="https://zenodo.org/record/3629060/files/figure.png" pageId="7" pageNumber="112">Fig 2 E, F</figureCitation>
; 
<figureCitation id="5BF265A53D1AFFACFBF890C01777FC9B" box="[1079,1091,755,777]" captionStart="FIGURE 5" captionStartId="7.[151,250,1544,1569]" captionTargetBox="[158,1427,1045,1518]" captionTargetId="figure@7.[151,1436,1037,1525]" captionTargetPageId="7" captionText="FIGURE 5. Taito adrik sp. nov., left leg IV of male holotype (MUSM 0513700), dorsal view: Patella, distal part of femur, and proximal part of tibia. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3629066" httpUri="https://zenodo.org/record/3629066/files/figure.png" pageId="7" pageNumber="112">5</figureCitation>
)............ 
<emphasis id="F1BDA5323D1AFFACFB1190C016A8FC9A" bold="true" box="[1246,1436,755,777]" pageId="7" pageNumber="112">
<taxonomicName id="04C902A33D1AFFACFB1190C0167EFC9B" authority="Friedrich &amp; Lehmann, 2020" authorityName="Friedrich &amp; Lehmann" authorityYear="2020" box="[1246,1354,755,777]" class="Arachnida" family="Cosmetidae" genus="Taito" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="adrik" status="sp. nov.">
<emphasis id="F1BDA5323D1AFFACFB1190C0167EFC9B" bold="true" box="[1246,1354,755,777]" italics="true" pageId="7" pageNumber="112">Taito adrik</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA8E18493D1AFFACFA9F90C016A8FC9A" box="[1360,1436,755,776]" pageId="7" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Equuleus H- or easel-shaped; femur IV with a comb of 7 or 8 distal spines; patella IV unarmed....................... 8
</paragraph>
<paragraph id="C37679203D1AFFACFF0891671125FC6D" blockId="7.[151,1437,852,1024]" pageId="7" pageNumber="112">
<emphasis id="F1BDA5323D1AFFACFF08916712B2FCFD" bold="true" box="[199,390,852,879]" pageId="7" pageNumber="112">COI barcoding.</emphasis>
Specimen data and DNA sequences of the studied species are available from BOLD (
<bibRefCitation id="A75804D13D1AFFACFAA5916612E4FC01" author="Ratnasingham, S. &amp; Hebert, P. D." pageId="7" pageNumber="112" pagination="355 - 364" refId="ref6129" refString="Ratnasingham, S. &amp; Hebert, P. D. (2007) BOLD: The Barcode of Life Data System (http: // www. barcodinglife. org). Molecular Ecology Resources, 7 (3), 355 - 364. https: // doi. org / 10.1111 / j. 1471 - 8286.2007.01678. x" type="journal article" year="2007">Ratnasingham &amp; Hebert 2007</bibRefCitation>
). The genetic divergences between the six sequenced specimens (
<typeStatus id="1C72C7823D1AFFACFB00914A1607FC01" box="[1231,1331,889,915]" pageId="7" pageNumber="112" type="holotype">holotype</typeStatus>
and five 
<typeStatus id="1C72C7823D1AFFACFF5891AE1230FC25" box="[151,260,925,951]" pageId="7" pageNumber="112" type="paratype">paratypes</typeStatus>
) range between 0.0%, and 2.0% (average 0.8%, see 
<tableCitation id="8E4B4C9B3D1AFFACFC9591AE1098FC25" box="[858,940,925,951]" captionStart="TABLE 1" captionStartId="1.[151,239,550,574]" captionTargetPageId="1" captionText="TABLE 1. COI p-distances for Taito adrik sp. nov. specimens sequenced in this study (* holotype MUSM 0513700)." httpUri="http://table.plazi.org/id/97B629A83D1CFFAAFF5890151649FDAC" pageId="7" pageNumber="112" tableUuid="97B629A83D1CFFAAFF5890151649FDAC">Table 1</tableCitation>
). A search in BOLD revealed specimens of the cosmetids 
<taxonomicName id="04C902A33D1AFFACFEF191F212AEFC49" authorityName="Koch" authorityYear="1839" box="[318,410,961,987]" class="Arachnida" family="Cosmetidae" genus="Cynorta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1AFFACFEF191F212AEFC49" box="[318,410,961,987]" italics="true" pageId="7" pageNumber="112">Cynorta</emphasis>
</taxonomicName>
sp. (86.49% similarity), 
<taxonomicName id="04C902A33D1AFFACFD7091F11028FC49" authorityName="Simon" authorityYear="1879" box="[703,796,962,987]" class="Arachnida" family="Cosmetidae" genus="Vonones" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1AFFACFD7091F11028FC49" box="[703,796,962,987]" italics="true" pageId="7" pageNumber="112">Vonones</emphasis>
</taxonomicName>
sp. (85.71% similarity), and 
<taxonomicName id="04C902A33D1AFFACFBBB91F1163CFC49" authorityName="Pickard-Cambridge" authorityYear="1904" box="[1140,1288,962,987]" class="Arachnida" family="Cosmetidae" genus="Paravonones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F1BDA5323D1AFFACFBBB91F1163CFC49" box="[1140,1288,962,987]" italics="true" pageId="7" pageNumber="112">Paravonones</emphasis>
</taxonomicName>
sp. (85.65% similarity) as the closest matches.
</paragraph>
</subSubSection>
</treatment>
</document>