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<document id="9C5373115B9B533111E6302E16002FD5" ID-DOI="10.5281/zenodo.158599" ID-GBIF-Dataset="cd0c4151-d8fc-439c-8a83-ba18ee2f20bf" ID-ISSN="1175­5326" ID-Zenodo-Dep="158599" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032444118" checkinUser="plazi" docAuthor="Rödel, Mark-Oliver, Schmitz, Andreas, Pauwels, Olivier S. G. &amp; Böhme, Wolfgang" docDate="2004" docId="7B388787E24DFFFCFEE43269891EFADA" docLanguage="en" docName="zt00720.pdf" docOrigin="Zootaxa 720" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Werneria" docType="key" docVersion="8" lastPageNumber="22" masterDocId="8701FFFFE258FFE9FFEC30328C35FF9A" masterDocTitle="Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="22" updateTime="1698573783727" updateUser="plazi">
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<mods:title id="5ABE2BBCA2C459FA084CA7B0DFD868E4">Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)</mods:title>
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<mods:namePart id="C7F07B7232EE2341687282CCF4144970">Rödel, Mark-Oliver</mods:namePart>
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<mods:namePart id="BA1AE84ECC30B59401945B7437C5FE5C">Schmitz, Andreas</mods:namePart>
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<mods:namePart id="E888CC47C310B7110CBBC9C790D94D4D">Pauwels, Olivier S. G.</mods:namePart>
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<mods:roleTerm id="609B7DB69804CD108495FFDD605C948B">Author</mods:roleTerm>
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<mods:namePart id="8449AA2D7D2D60AA393564677F0801B5">Böhme, Wolfgang</mods:namePart>
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<mods:title id="7DD99304A7F9E75A62DAD9FAC42F7720">Zootaxa</mods:title>
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<mods:date id="69AD6B36C889F17F1482074BFBDD53AD">2004</mods:date>
<mods:detail id="8AD4CC861BC790A03063326343C2EF2B" type="volume">
<mods:number id="E7CBE46C547366523559F3AE41A7B490">720</mods:number>
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<mods:classification id="F676DFFE77DFCA2A3C8200BD8AA198C5">journal article</mods:classification>
<mods:identifier id="414CBFA2D7278FB851E4EBED5E07BFDD" type="DOI">10.5281/zenodo.158599</mods:identifier>
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<mods:identifier id="33BE0EA467826202EB370528E445132B" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="1C989BB251803D1AE18F2AE8AB7F9793" type="Zenodo-Dep">158599</mods:identifier>
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<treatment id="7B388787E24DFFFCFEE43269891EFADA" ID-DOI="http://doi.org/10.5281/zenodo.5610969" ID-GBIF-Taxon="119608459" ID-Zenodo-Dep="5610969" LSID="urn:lsid:plazi:treatment:7B388787E24DFFFCFEE43269891EFADA" httpUri="http://treatment.plazi.org/id/7B388787E24DFFFCFEE43269891EFADA" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="BB8B651AE24DFFFCFEE432698E1AFDEF" box="[264,559,603,630]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="F32E3691E24DFFFCFEE432698E1AFDEF" blockId="21.[264,559,603,630]" box="[264,559,603,630]" pageId="21" pageNumber="22">
<heading id="A86681FDE24DFFFCFEE432698E1AFDEF" bold="true" box="[264,559,603,630]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="C1E5EA83E24DFFFCFEE432698E1AFDEF" bold="true" box="[264,559,603,630]" pageId="21" pageNumber="22">
Key to
<taxonomicName id="34914D12E24DFFFCFE8E326E8DE5FDEC" ID-CoL="87T9" box="[354,464,604,630]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="C1E5EA83E24DFFFCFE8E326E8DE5FDEC" bold="true" box="[354,464,604,630]" italics="true" pageId="21" pageNumber="22">Werneria</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BB8B651AE24DFFFCFEE43295891EFADA" pageId="21" pageNumber="22" type="key">
<keyStep id="48652E34E24DFFFCFEE432958919FD73" pageId="21" pageNumber="22">
<paragraph id="F32E3691E24DFFFCFEE432958919FD5B" blockId="21.[264,1324,679,1344]" box="[264,1324,679,705]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE432958919FD5B" box="[264,1324,679,705]" pageId="21" pageNumber="22">
1 Belly with larger (&gt;
<quantity id="34699B74E24DFFFCFDCA32958EBCFD5B" box="[550,649,679,705]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="21" pageNumber="22" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
) white or yellow spots ................................................... 2
</keyLead>
</paragraph>
<paragraph id="F32E3691E24DFFFCFEE432FD8919FD73" blockId="21.[264,1324,679,1344]" box="[264,1324,719,745]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE432FD8919FD73" box="[264,1324,719,745]" pageId="21" pageNumber="22">
1' Belly with only very small (&lt;
<quantity id="34699B74E24DFFFCFD7432FD8EE8FD73" box="[664,733,719,745]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="mm" value="1.0">1 mm</quantity>
) white points ...................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="48652E34E24DFFFCFEE432C5891EFC2B" pageId="21" pageNumber="22">
<paragraph id="F32E3691E24DFFFCFEE432C5891FFCFA" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE432C5891FFCFA" pageId="21" pageNumber="22">
2
<collectingCountry id="8B867601E24DFFFCFED432C58D46FC8B" box="[312,371,759,785]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet moderately webbed, at least web not extending middle of fourth toe; back clear brown or reddish brown; two dark bars on thighs; urostyle bordered by black stripes
<emphasis id="C1E5EA83E24DFFFCFED73375891FFCFA" box="[315,1322,839,864]" italics="true" pageId="21" pageNumber="22">
......................................................................................................
<taxonomicName id="34914D12E24DFFFCFBC63375891FFCFA" ID-CoL="5BXH2" box="[1066,1322,839,864]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="mertensiana">Werneria mertensiana</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F32E3691E24DFFFCFEE4335D891EFC2B" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE4335D891EFC2B" pageId="21" pageNumber="22">
2'
<collectingCountry id="8B867601E24DFFFCFED4335D8D46FC13" box="[312,371,879,905]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet with webbing that extends at least to the middle of the medial side of the fourth toe ...................................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="48652E34E24DFFFCFEE4338D891EFBCB" pageId="21" pageNumber="22">
<paragraph id="F32E3691E24DFFFCFEE4338D891EFB9B" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE4338D891EFB9B" pageId="21" pageNumber="22">
3
<collectingCountry id="8B867601E24DFFFCFED4338D8D46FC43" box="[312,371,959,985]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet completely webbed; very thin dorsolateral line of equal width along body; flanks uniform black
<emphasis id="C1E5EA83E24DFFFCFDD733D588FEFB9A" box="[571,1227,999,1024]" italics="true" pageId="21" pageNumber="22">
............................................................
<taxonomicName id="34914D12E24DFFFCFC1433D588FEFB9A" ID-CoL="5BXGK" box="[1016,1227,999,1024]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="iboundji">Werneria iboundji</taxonomicName>
</emphasis>
nov. sp.
</keyLead>
</paragraph>
<paragraph id="F32E3691E24DFFFCFEE4343D891EFBCB" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE4343D891EFBCB" pageId="21" pageNumber="22">
3' At least fourth toe of hind foot incompletely webbed; dorsolateral line extending posteriorly; lower flanks with clear spots
<emphasis id="C1E5EA83E24DFFFCFD15340588FCFBCA" box="[761,1225,1079,1104]" italics="true" pageId="21" pageNumber="22">
............................
<taxonomicName id="34914D12E24DFFFCFC25340588FCFBCA" ID-CoL="5BXJ4" box="[969,1225,1079,1104]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="submontana">Werneria submontana</taxonomicName>
</emphasis>
nov. sp.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="48652E34E24DFFFCFEE4346D891EFB3B" pageId="21" pageNumber="22">
<paragraph id="F32E3691E24DFFFCFEE4346D891EFBE2" blockId="21.[264,1324,679,1344]" box="[264,1323,1119,1145]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE4346D891EFBE2" box="[264,1323,1119,1145]" pageId="21" pageNumber="22">
4
<collectingCountry id="8B867601E24DFFFCFED4346D8D47FBE3" box="[312,370,1119,1145]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet with only traces of webbing
<emphasis id="C1E5EA83E24DFFFCFCED346D891EFBE2" box="[769,1323,1119,1144]" italics="true" pageId="21" pageNumber="22">
.................................................
<taxonomicName id="34914D12E24DFFFCFB83346D891EFBE2" ID-CoL="5BXJ5" box="[1135,1323,1119,1144]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="tandyi">Werneria tandyi</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F32E3691E24DFFFCFEE434B5891EFB3B" blockId="21.[264,1324,679,1344]" box="[264,1323,1159,1185]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE434B5891EFB3B" box="[264,1323,1159,1185]" pageId="21" pageNumber="22">
4'
<collectingCountry id="8B867601E24DFFFCFED434B58D47FB3B" box="[312,370,1159,1185]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet webbed ......................................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="48652E34E24DFFFCFEE4349D891EFADA" pageId="21" pageNumber="22">
<paragraph id="F32E3691E24DFFFCFEE4349D891FFB6A" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE4349D891FFB6A" pageId="21" pageNumber="22">
5
<collectingCountry id="8B867601E24DFFFCFED4349D8D46FB53" box="[312,371,1199,1225]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet not completely webbed; young males and some females with distinct, clear dorsolateral lines
<emphasis id="C1E5EA83E24DFFFCFDE334E5891FFB6A" box="[527,1322,1239,1264]" italics="true" pageId="21" pageNumber="22">
.................................................................................
<taxonomicName id="34914D12E24DFFFCFB8834E5891FFB6A" ID-CoL="5BXHG" box="[1124,1322,1239,1264]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="preussi">Werneria preussi</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F32E3691E24DFFFCFEE434CD891EFADA" blockId="21.[264,1324,679,1344]" pageId="21" pageNumber="22">
<keyLead id="486095A4E24DFFFCFEE434CD891EFADA" pageId="21" pageNumber="22">
5'
<collectingCountry id="8B867601E24DFFFCFED434CD8D46FA83" box="[312,371,1279,1305]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
feet completely webbed; neither sex with distinct, clear dorsolateral lines ..........
<emphasis id="C1E5EA83E24DFFFCFED73515891EFADA" box="[315,1323,1319,1344]" italics="true" pageId="21" pageNumber="22">
.....................................................................................................
<taxonomicName id="34914D12E24DFFFCFBCA3515891EFADA" ID-CoL="7FW2Z" box="[1062,1323,1319,1344]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="bambutensis">Werneria bambutensis</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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