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<document id="0A4F2AECF868240230222C85FDC53AB5" ID-DOI="10.5281/zenodo.158599" ID-GBIF-Dataset="cd0c4151-d8fc-439c-8a83-ba18ee2f20bf" ID-ISSN="1175­5326" ID-Zenodo-Dep="158599" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032444118" checkinUser="plazi" docAuthor="Rödel, Mark-Oliver, Schmitz, Andreas, Pauwels, Olivier S. G. &amp; Böhme, Wolfgang" docDate="2004" docId="7B388787E248FFFAFEE432FE8D08FDD4" docLanguage="en" docName="zt00720.pdf" docOrigin="Zootaxa 720" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Werneria submontana" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="8701FFFFE258FFE9FFEC30328C35FF9A" masterDocTitle="Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="17" updateTime="1698573783727" updateUser="plazi">
<mods:mods id="5D1A70C016326E41F5743F1B4FEB1E55" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A12374424A7386146E56AAA0BC57A35F">
<mods:title id="E8447BCB092589B1A6B7FFB6C94D62E2">Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)</mods:title>
</mods:titleInfo>
<mods:name id="6C90912A7DDA82CE108A6AD8C653890C" type="personal">
<mods:role id="43452C4C415C7F162C2C6809BD553761">
<mods:roleTerm id="4791FDF592150E7DDAA295E28933BB0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58BF0EDD14BF8080D0BB100C4577CACB">Rödel, Mark-Oliver</mods:namePart>
</mods:name>
<mods:name id="B28D20E1D761ABC6369A36F836C99D0D" type="personal">
<mods:role id="5A019218D5D747D0A68000C2E5076711">
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</mods:role>
<mods:namePart id="DE832B1C79AF58BD628A0020196EA3EF">Schmitz, Andreas</mods:namePart>
</mods:name>
<mods:name id="0FECF59AB03A918B5B20E873F336A098" type="personal">
<mods:role id="144EE9FFB9C40F5EE4C9E71893A92A80">
<mods:roleTerm id="7A6908DEDFD4F993CB054AC76B0787C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26633F12DF17A438E138D2CC181EC28E">Pauwels, Olivier S. G.</mods:namePart>
</mods:name>
<mods:name id="4B16600BE9EC94ABDB072DA3AE710328" type="personal">
<mods:role id="3E0DF21EA3458ECD5F3741E041CEA2B4">
<mods:roleTerm id="F5AD58F3485078CC71272D9D9B67717D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C42A8C76D4BAEFB304186EF3A5CC3C39">Böhme, Wolfgang</mods:namePart>
</mods:name>
<mods:typeOfResource id="CD2E34E6EA69BB2DEAE4B63FAE5EF3D8">text</mods:typeOfResource>
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<mods:titleInfo id="D67174FA7E754E83FFF77C155CAD0A06">
<mods:title id="AC824CD3F7F61D53A3442C3EF2B0B765">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AF3923C2A641273F0CC1D65FFFBF5136">
<mods:date id="8D4B247DA37C909A9DC977C8F035C482">2004</mods:date>
<mods:detail id="8EAD2D5E24892ED0197329682E127DE5" type="volume">
<mods:number id="EBE2CF271DBCE015B32AE8E18A91EC01">720</mods:number>
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<mods:start id="9EB3D060B5D66E0F42EAB4FF272CB85A">1</mods:start>
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<mods:classification id="1A790E84018DD83DCD15285AB960D500">journal article</mods:classification>
<mods:identifier id="40B78C892BC9F4C80F9A547816798F49" type="DOI">10.5281/zenodo.158599</mods:identifier>
<mods:identifier id="7F337D09C2FE12719741885B34447897" type="GBIF-Dataset">cd0c4151-d8fc-439c-8a83-ba18ee2f20bf</mods:identifier>
<mods:identifier id="A528DEB609167913FF40BBD2CE9B873C" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="0D1241B7F974D0CAC4B7BB3744CC12F3" type="Zenodo-Dep">158599</mods:identifier>
<mods:identifier id="1E2617FD09CCE1AA12038994107176AF" type="ZooBank">6D085854-E586-4291-B968-F3C59885BF2E</mods:identifier>
</mods:mods>
<treatment id="7B388787E248FFFAFEE432FE8D08FDD4" ID-DOI="http://doi.org/10.5281/zenodo.5610965" ID-GBIF-Taxon="119608466" ID-Zenodo-Dep="5610965" LSID="urn:lsid:plazi:treatment:7B388787E248FFFAFEE432FE8D08FDD4" httpUri="http://treatment.plazi.org/id/7B388787E248FFFAFEE432FE8D08FDD4" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="BB8B651AE248FFF9FEE432FE8E7BFC93" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="F32E3691E248FFF9FEE432FE8E4FFD7C" blockId="16.[264,634,716,777]" box="[264,634,716,742]" pageId="16" pageNumber="17">
<heading id="A86681FDE248FFF9FEE432FE8E4FFD7C" bold="true" box="[264,634,716,742]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="C1E5EA83E248FFF9FEE432FE8E4FFD7C" bold="true" box="[264,634,716,742]" pageId="16" pageNumber="17">
<taxonomicName id="34914D12E248FFF9FEE432FE8E20FD7C" ID-CoL="5BXJ4" box="[264,533,716,742]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E248FFF9FEE432FE8E20FD7C" bold="true" box="[264,533,716,742]" italics="true" pageId="16" pageNumber="17">Werneria submontana</emphasis>
</taxonomicName>
nov. sp.
</emphasis>
</heading>
</paragraph>
<paragraph id="F32E3691E248FFF9FEE432DD8E7BFC93" blockId="16.[264,634,716,777]" box="[264,590,751,777]" pageId="16" pageNumber="17">
<figureCitation id="6BAA2A14E248FFF9FEE432DD8D60FC93" box="[264,341,751,777]" captionStart="FIGURE 1" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[450,1136,1036,1715]" captionTargetId="figure@4.[450,1136,1036,1717]" captionTargetPageId="4" captionText="FIGURE 1: Dorsal view of Werneria species; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/169501/files/figure.png" pageId="16" pageNumber="17">Figs. 1</figureCitation>
d, 2d, 3d, 4d, 8b, 10a
</paragraph>
</subSubSection>
<subSubSection id="BB8B651AE248FFF9FEE4330388CDFB29" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="F32E3691E248FFF9FEE433038FB8FCE9" blockId="16.[264,1323,817,1883]" pageId="16" pageNumber="17">
<emphasis id="C1E5EA83E248FFF9FEE433038D46FCD0" box="[264,371,817,842]" italics="true" pageId="16" pageNumber="17">
<typeStatus id="2C2A8833E248FFF9FEE433038D46FCD0" box="[264,371,817,842]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
.—
<collectionCode id="9580AE54E248FFF9FE4E33038DC1FCD1" LSID="urn:lsid:biocol.org:col:34613" box="[418,500,817,843]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
69699, adult male, Bakossi Mts., Edib Hills, Lake Edib,
<geoCoordinate id="96A55056E248FFF9FB443303893FFCD1" box="[1192,1290,817,843]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="4.95">N 4°57’</geoCoordinate>
,
<geoCoordinate id="96A55056E248FFF9FAF533038D7EFCE9" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="9.65">E 9°39’</geoCoordinate>
,
<quantity id="34699B74E248FFF9FEB4336B8D8DFCE9" box="[344,440,857,883]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="17" unit="m" value="1100.0">1,100 m</quantity>
a.s.l.,
<collectingCountry id="8B867601E248FFF9FDE9336B8EB7FCE9" box="[517,642,857,883]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
,
<date id="872F1051E248FFF9FD61336B8F28FCE9" box="[653,797,857,883]" pageId="16" pageNumber="17" value="1998-05-05" valueMax="1998-05-07" valueMin="1998-05-05">5.–7.V.1998</date>
, C. Wild.
</paragraph>
<paragraph id="F32E3691E248FFF9FED433B388CDFB29" blockId="16.[264,1323,817,1883]" pageId="16" pageNumber="17">
<emphasis id="C1E5EA83E248FFF9FED433B38D9AFC00" box="[312,431,897,922]" italics="true" pageId="16" pageNumber="17">
<typeStatus id="2C2A8833E248FFF9FED433B38D9AFC00" box="[312,431,897,922]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
.—
<collectionCode id="9580AE54E248FFF9FE3133B38E1BFC01" LSID="urn:lsid:biocol.org:col:34613" box="[477,558,897,923]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
67271, juvenile,
<quantity id="34699B74E248FFF9FCEC33B38F57FC01" box="[768,866,897,923]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="16" pageNumber="17" unit="m" value="1250.0">1,250 m</quantity>
a.s.l.,
<date id="872F1051E248FFF9FC5F33B38810FC01" box="[947,1061,897,923]" pageId="16" pageNumber="17" value="1998-01-14">14.I.1998</date>
,
<collectingCountry id="8B867601E248FFF9FBD833B38879FC01" box="[1076,1100,897,923]" name="Iceland" pageId="16" pageNumber="17">O</collectingCountry>
. Euskirchen, other data as
<typeStatus id="2C2A8833E248FFF9FE8D339B8DFEFC59" box="[353,459,937,963]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<collectionCode id="9580AE54E248FFF9FE3B339B8E1CFC59" LSID="urn:lsid:biocol.org:col:34613" box="[471,553,937,963]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
69747, adult male, same data as
<typeStatus id="2C2A8833E248FFF9FC5E339B8829FC59" box="[946,1052,937,963]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
;
<collectionCode id="9580AE54E248FFF9FBC4339B884FFC59" LSID="urn:lsid:biocol.org:col:34613" box="[1064,1146,937,963]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
69698, 69700, adult female and male, Bakossi Mts., Edib Hills, Elabah Camp,
<quantity id="34699B74E248FFF9FC1733E38869FC71" box="[1019,1116,977,1003]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="17" unit="m" value="1100.0">1,100 m</quantity>
a.s.l.,
<collectingCountry id="8B867601E248FFF9FB4533E38913FC71" box="[1193,1318,977,1003]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
,
<date id="872F1051E248FFF9FEE433CB8DADFB89" box="[264,408,1017,1043]" pageId="16" pageNumber="17" value="1998-05-05" valueMax="1998-05-07" valueMin="1998-05-05">5.–7.V.1998</date>
, C. Wild;
<collectionCode id="9580AE54E248FFF9FDF633CB8E5EFB89" LSID="urn:lsid:biocol.org:col:34613" box="[538,619,1017,1043]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
67272, 67781­785, six juveniles, Bakossi Mts., Mwendelengo Mts., Kodmin,
<quantity id="34699B74E248FFF9FDC834138EBAFBA1" box="[548,655,1057,1083]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="16" pageNumber="17" unit="m" value="1070.0">1,070 m</quantity>
a.s.l.,
<collectingCountry id="8B867601E248FFF9FD1D34138F5BFBA1" box="[753,878,1057,1083]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
,
<date id="872F1051E248FFF9FC6934138872FBA1" box="[901,1095,1057,1083]" pageId="16" pageNumber="17" value="1997-12-18" valueMax="1997-12-25" valueMin="1997-12-18">18.–25.XII.1997</date>
&amp;
<date id="872F1051E248FFF9FB6F341388C3FBA1" box="[1155,1270,1057,1083]" pageId="16" pageNumber="17" value="1998-01-24">24.I.1998</date>
,
<collectingCountry id="8B867601E248FFF9FAE134138910FBA1" box="[1293,1317,1057,1083]" name="Iceland" pageId="16" pageNumber="17">O</collectingCountry>
. Euskirchen &amp; A. Schmitz;
<collectionCode id="9580AE54E248FFF9FDAA347B8EA2FBF9" LSID="urn:lsid:biocol.org:col:34613" box="[582,663,1097,1123]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
67786, adult male, Meked,
<collectingCountry id="8B867601E248FFF9FC0D347B886BFBF9" box="[993,1118,1097,1123]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
,
<date id="872F1051E248FFF9FB86347B8937FBF9" box="[1130,1282,1097,1123]" pageId="16" pageNumber="17" value="1998-01-08" valueMax="1998-01-10" valueMin="1998-01-08">8.–10.I.1998</date>
, A. Schmitz;
<collectionCode id="9580AE54E248FFF9FE9434438DFFFB11" LSID="urn:lsid:biocol.org:col:34613" box="[376,458,1137,1163]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
67787­796, ten juveniles, Mt. Kupe, south­west side (north­west flank in catalogue of
<collectionCode id="9580AE54E248FFF9FE7334AB8DC0FB29" LSID="urn:lsid:biocol.org:col:34613" box="[415,501,1177,1203]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
), Nyasoso,
<quantity id="34699B74E248FFF9FD9034AB8EF3FB29" box="[636,710,1177,1203]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="16" pageNumber="17" unit="m" value="910.0">910 m</quantity>
a.s.l.,
<collectingCountry id="8B867601E248FFF9FCFF34AB8FA5FB29" box="[787,912,1177,1203]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
,
<date id="872F1051E248FFF9FC7734AB8855FB29" box="[923,1120,1177,1203]" pageId="16" pageNumber="17" value="1997-12-18" valueMax="1997-12-25" valueMin="1997-12-18">18.–25.XII.1997</date>
, A. Schmitz.
</paragraph>
</subSubSection>
<subSubSection id="BB8B651AE248FFF9FED434F38848F979" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="F32E3691E248FFF9FED434F38848F979" blockId="16.[264,1323,817,1883]" pageId="16" pageNumber="17">
<emphasis id="C1E5EA83E248FFF9FED434F38D9AFB40" box="[312,431,1217,1242]" italics="true" pageId="16" pageNumber="17">Diagnosis</emphasis>
.— A medium­sized, slender but compact
<taxonomicName id="34914D12E248FFF9FC4834F3883BFB40" box="[932,1038,1217,1242]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="C1E5EA83E248FFF9FC4834F3883BFB40" box="[932,1038,1217,1242]" italics="true" pageId="16" pageNumber="17">Werneria</emphasis>
</taxonomicName>
with short and rounded snout; skin of flanks and throat wrinkled, at least in adult males, otherwise skin smooth; a distinct, broad white, yellowish or reddish dorsolateral line occurs on the dark brown dorsum; urostyle bordered by a pair of longitudinal black lines; lower flanks with large clear spots; greyish venter with large clear spots, toes nearly fully webbed. Superficially this species is most similar to
<taxonomicName id="34914D12E248FFF9FDD235BB8EC0FA38" box="[574,757,1417,1442]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="mertensiana">
<emphasis id="C1E5EA83E248FFF9FDD235BB8EC0FA38" box="[574,757,1417,1442]" italics="true" pageId="16" pageNumber="17">W. mertensiana</emphasis>
</taxonomicName>
,
<taxonomicName id="34914D12E248FFF9FCE935BB8F4AFA38" box="[773,895,1417,1442]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="preussi">
<emphasis id="C1E5EA83E248FFF9FCE935BB8F4AFA38" box="[773,895,1417,1442]" italics="true" pageId="16" pageNumber="17">W. preussi</emphasis>
</taxonomicName>
and the new Gabonese species (see below). From the Gabonese species and
<emphasis id="C1E5EA83E248FFF9FD1C35838844FA50" box="[752,1137,1457,1482]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="34914D12E248FFF9FD1C35838F99FA50" box="[752,940,1457,1482]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="bambutensis">W. bambutensis</taxonomicName>
,
<taxonomicName id="34914D12E248FFF9FC5635838844FA50" box="[954,1137,1457,1482]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="submontana">W. submontana</taxonomicName>
</emphasis>
nov. sp. differs by its less­developed webbing and from
<taxonomicName id="34914D12E248FFF9FD1C35EB8F6AFA68" box="[752,863,1497,1522]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="tandyi">
<emphasis id="C1E5EA83E248FFF9FD1C35EB8F6AFA68" box="[752,863,1497,1522]" italics="true" pageId="16" pageNumber="17">W. tandyi</emphasis>
</taxonomicName>
,
<taxonomicName id="34914D12E248FFF9FC9C35EB881DFA68" box="[880,1064,1497,1522]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="mertensiana">
<emphasis id="C1E5EA83E248FFF9FC9C35EB881DFA68" box="[880,1064,1497,1522]" italics="true" pageId="16" pageNumber="17">W. mertensiana</emphasis>
</taxonomicName>
and
<taxonomicName id="34914D12E248FFF9FB8935EB88E9FA68" box="[1125,1244,1497,1522]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="preussi">
<emphasis id="C1E5EA83E248FFF9FB8935EB88E9FA68" box="[1125,1244,1497,1522]" italics="true" pageId="16" pageNumber="17">W. preussi</emphasis>
</taxonomicName>
by its more extensive webbing. It differs from the Gabonese species by a different body shape and much broader dorsolateral lines. With
<taxonomicName id="34914D12E248FFF9FD13361B8F80F9D8" box="[767,949,1577,1602]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="mertensiana">
<emphasis id="C1E5EA83E248FFF9FD13361B8F80F9D8" box="[767,949,1577,1602]" italics="true" pageId="16" pageNumber="17">W. mertensiana</emphasis>
</taxonomicName>
it shares the black bars on hind legs and the black borders of the urostyle, but differs by its wrinkled flanks that bear clear spots on the ventral part, the more distinctly dorsal border of the dorsolateral lines and head shape. From
<taxonomicName id="34914D12E248FFF9FE0F36938E68F920" box="[483,605,1697,1722]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="preussi">
<emphasis id="C1E5EA83E248FFF9FE0F36938E68F920" box="[483,605,1697,1722]" italics="true" pageId="16" pageNumber="17">W. preussi</emphasis>
</taxonomicName>
it differs by its dorsal and ventral coloration. It differs from all
<taxonomicName id="34914D12E248FFF9FEC136FB8DACF978" box="[301,409,1737,1762]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="C1E5EA83E248FFF9FEC136FB8DACF978" box="[301,409,1737,1762]" italics="true" pageId="16" pageNumber="17">Werneria</emphasis>
</taxonomicName>
species by the wrinkled skin on throat (at least in adult males).
</paragraph>
</subSubSection>
<subSubSection id="BB8B651AE248FFF8FED436C388D6F94C" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="F32E3691E248FFF8FED436C3881BFD74" blockId="16.[264,1323,817,1883]" lastBlockId="17.[264,1326,284,1750]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="C1E5EA83E248FFF9FED436C38E43F890" box="[312,630,1777,1802]" italics="true" pageId="16" pageNumber="17">
Description of the
<typeStatus id="2C2A8833E248FFF9FDFE36C38E43F890" box="[530,630,1777,1802]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
</emphasis>
.— The
<typeStatus id="2C2A8833E248FFF9FD3B36C38F0BF891" box="[727,830,1777,1803]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
is an adult male with slender but compact body shape (
<figureCitation id="6BAA2A14E248FFF9FE70372B8DEBF8A9" box="[412,478,1817,1843]" captionStart="FIGURE 1" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[450,1136,1036,1715]" captionTargetId="figure@4.[450,1136,1036,1717]" captionTargetPageId="4" captionText="FIGURE 1: Dorsal view of Werneria species; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/169501/files/figure.png" pageId="16" pageNumber="17">Fig. 1</figureCitation>
d);
<quantity id="34699B74E248FFF9FDE5372B8E44F8A9" box="[521,625,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="16" pageNumber="17" unit="mm" value="35.4">35.4 mm</quantity>
SVL; short, broad but rounded snout; head width
<quantity id="34699B74E248FFF9FB50372B8913F8A9" box="[1212,1318,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="16" pageNumber="17" unit="mm" value="11.3">11.3 mm</quantity>
, eye diameter 4.0 mm; interorbital distance
<quantity id="34699B74E248FFF9FD1337738F69F8C1" box="[767,860,1857,1883]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="16" pageNumber="17" unit="mm" value="4.6">4.6 mm</quantity>
; naris closer to snout tip (
<quantity id="34699B74E248FFF9FB66377388D2F8C1" box="[1162,1255,1857,1883]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="17" unit="mm" value="2.4">2.4 mm</quantity>
) than to eye (
<quantity id="34699B74E249FFF8FE8C312E8DF5FEAC" box="[352,448,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="17" pageNumber="18" unit="mm" value="3.1">3.1 mm</quantity>
); small warts at angle of mouth (
<figureCitation id="6BAA2A14E249FFF8FCAF312E8FBFFEAC" box="[835,906,284,310]" captionStart="FIGURE 4" captionStartId="7.[264,367,914,938]" captionTargetBox="[280,1307,294,888]" captionTargetId="figure@7.[280,1311,284,907]" captionTargetPageId="7" captionText="FIGURE 4: Heads of Werneria species, illustrating shape of snout, dorsolateral lines (stippled), warts and spines; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi female SMF 24159; g: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/158603/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
d), skin of back and venter smooth; broad (
<quantity id="34699B74E249FFF8FEB631768DABFEC4" box="[346,414,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="2.0">2 mm</quantity>
at mid­body) white dorsolateral lines that extend from snout to groins; dorsolateral line distinctly separated from back and flank coloration and broadened posteriorly; back dark grey; blackish flanks with granular skin, lower flanks with white spots; back uniform dark brown to black; urostyle bordered by black lines; dorsal surfaces of extremities dark brown with two black bars on thighs, lower leg and feet (
<figureCitation id="6BAA2A14E249FFF8FBE231D68865FE64" box="[1038,1104,484,510]" captionStart="FIGURE 1" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[450,1136,1036,1715]" captionTargetId="figure@4.[450,1136,1036,1717]" captionTargetPageId="4" captionText="FIGURE 1: Dorsal view of Werneria species; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/169501/files/figure.png" pageId="17" pageNumber="18">Fig. 1</figureCitation>
d); basic colour of venter brownish grey; posterior venter with large (&gt;
<quantity id="34699B74E249FFF8FC6B323E8FE4FDBC" box="[903,977,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="mm" value="4.0">4 mm</quantity>
) yellowish spots; breast and throat grey with a dark ventral figure on throat and few smaller yellowish spots; skin of throat wrinkled (
<figureCitation id="6BAA2A14E249FFF8FE3F326E8E1CFDEC" box="[467,553,604,630]" captionStart="FIGURE 2" captionStartId="5.[264,367,1184,1208]" captionTargetBox="[359,1226,310,1159]" captionTargetId="figure@5.[359,1228,309,1160]" captionTargetPageId="5" captionText="FIGURE 2: Ventral view of Werneria species; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/169502/files/figure.png" pageId="17" pageNumber="18">Figs. 2</figureCitation>
d, 10a); ventral surfaces of thighs and lower leg reddish brown with large yellowish spots on the whole surface; long, but broad, hind legs, femur length
<quantity id="34699B74E249FFF8FEE4329E8D4DFD5C" box="[264,376,684,710]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="17" pageNumber="18" unit="mm" value="16.7">16.7 mm</quantity>
, tibia
<quantity id="34699B74E249FFF8FE2B329E8E02FD5C" box="[455,567,684,710]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="17" pageNumber="18" unit="mm" value="17.2">17.2 mm</quantity>
, foot (incl. longest toe)
<quantity id="34699B74E249FFF8FC88329E8FE6FD5C" box="[868,979,684,710]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="17" pageNumber="18" unit="mm" value="25.2">25.2 mm</quantity>
; toes 2, 3 and 5 completely webbed, tip of toe 1 and external side of toe 4 without web (
<figureCitation id="6BAA2A14E249FFF8FC2532E6883AFD74" box="[969,1039,724,750]" captionStart="FIGURE 3" captionStartId="6.[264,367,1772,1796]" captionTargetBox="[371,1243,440,1760]" captionTargetId="figure@6.[335,1266,435,1767]" captionTargetPageId="6" captionText="FIGURE 3: Left hind feet of Werneria species, webs are stippled; a: W. iboundji nov. sp. female, holotype, IRSNB 1929; b: W. bambutensis male, MHNG 1453.17; c: W. tandyi male, MHNG 1453.21; d: W. submontana nov. sp. male, holotype, ZFMK 69699; e: W. mertensiana male, ZFMK 69137; f: W. preussi male, SMF 24180." httpUri="https://zenodo.org/record/158602/files/figure.png" pageId="17" pageNumber="18">Fig. 3</figureCitation>
d).
</paragraph>
<paragraph id="F32E3691E249FFF8FED432CE8D61FB9C" blockId="17.[264,1326,284,1750]" pageId="17" pageNumber="18">
<emphasis id="C1E5EA83E249FFF8FED432CE8E4DFC8F" box="[312,632,764,789]" italics="true" pageId="17" pageNumber="18">Variation and colour in life</emphasis>
.— Adult males SVL ranged from
<quantity id="34699B74E249FFF8FBFF32CE88F4FC8C" box="[1043,1217,764,790]" metricMagnitude="-2" metricUnit="m" metricValue="3.335" metricValueMax="3.54" metricValueMin="3.13" pageId="17" pageNumber="18" unit="mm" value="33.35" valueMax="35.4" valueMin="31.3">31.3–35.4 mm</quantity>
, the only adult female measured
<quantity id="34699B74E249FFF8FDF433168EB0FCA4" box="[536,645,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="17" pageNumber="18" unit="mm" value="44.5">44.5 mm</quantity>
. The smallest juvenile was
<quantity id="34699B74E249FFF8FC2A3316881BFCA4" box="[966,1070,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="17" pageNumber="18" unit="mm" value="10.5">10.5 mm</quantity>
long. Other measurements of
<typeStatus id="2C2A8833E249FFF8FE9B337E8DD2FCFC" box="[375,487,844,870]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
are summarized in
<tableCitation id="BE13032AE249FFF8FD3E337E8F63FCFC" box="[722,854,844,870]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="2.[264,350,923,947]" captionStartId-1="3.[264,350,283,307]" captionStartId-2="3.[264,350,1166,1190]" captionTargetBox-1="[264,1319,438,1092]" captionTargetBox-2="[264,1312,1291,1560]" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="TABLE 1. Summary of morphological measures (mm) of adult male Werneria. Given are minimum, maximum and mean values, including standard deviation. SVL = snout­vent­length, HW = head width, ED = eye diameter, IOD = interorbital distance, EN = distance eye nose, NS = distance nose snout­tip, FE = femur, TI = tibia, FO = foot including longest toe. For specimens investigated see respective species descriptions. Values from specimens kept in the research collection of J. ­ L. Amiet are given in parenthesis (data kindly provided by J. ­ L. Amiet)." captionText-1="TABLE 2. Summary of morphological measures (mm) of adult female Werneria. Given are minimum, maximum and mean values, including standard deviation. For specimens investigated see respective species descriptions. Values from specimens kept in the research collection of J. ­ L. Amiet are given in parenthesis (data kindly provided by J. ­ L. Amiet). For abbreviations see Tab. 1." captionText-2="TABLE 3. Summary of morphological measures (mm) of juvenile and subadult Werneria. Given are minimum and maximum values. For specimens investigated see respective species descriptions. For abbreviations see Tab. 1." httpUri-1="http://table.plazi.org/id/A7EE6619E25BFFEAFEE431298910FE09" httpUri-2="http://table.plazi.org/id/A7EE6619E25BFFEAFEE434BC8E0EFB71" pageId="17" pageNumber="18" tableUuid-1="A7EE6619E25BFFEAFEE431298910FE09" tableUuid-2="A7EE6619E25BFFEAFEE434BC8E0EFB71">Tables 1–3</tableCitation>
. Body dimensions and colouration generally were as in the
<typeStatus id="2C2A8833E249FFF8FDEF33468E58FC14" box="[515,621,884,910]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
. Other adult males also had a wrinkled skin on throat. The female’s throat was only faintly wrinkled. Warts at angle of mouth may be difficult to see or absent. Juveniles generally differ from adults by having smooth skin on flanks and throat.
</paragraph>
<paragraph id="F32E3691E249FFF8FED434268E34FA84" blockId="17.[264,1326,284,1750]" pageId="17" pageNumber="18">
In life these toads have a golden iris; dorsal colour dark brown to black; pale, whitish, pink or orange­red dorsolateral stripes (
<figureCitation id="6BAA2A14E249FFF8FD31340E8F16FBCC" box="[733,803,1084,1110]" captionStart="FIGURE 8" captionStartId="12.[264,367,1783,1807]" captionTargetBox="[313,1272,830,1764]" captionTargetId="figure@12.[313,1273,830,1767]" captionTargetPageId="12" captionText="FIGURE 8: a: W. preussi female from Buëa, Mt. Cameroon; b: W. submontana nov. sp. male, Bakossi Mts., Lake Edib; c: W. mertensiana male, Mt. Kupe (photo: K. ­ H. Jungfer); d: ventral view of Werneria sp., Monte Alén National Park, Equatorial Guinea (photo: I. de la Riva)." httpUri="https://zenodo.org/record/169504/files/figure.png" pageId="17" pageNumber="18">Fig. 8</figureCitation>
b), very faint in the one adult female; upper part of flanks dark brown to black, lower part of flanks with yellowish spots that may be hard to see in juveniles; belly dark grey with numerous cream speckles; belly generally with larger spots than breast and throat, however, in some specimens not exceeding
<quantity id="34699B74E249FFF8FB1334868D03FB6C" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="17" pageNumber="18" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
in size; femur, tibia, foot brownish with two black bars on each; forefeet and fingers with yellowish spots.
</paragraph>
<paragraph id="F32E3691E249FFF8FED4351E88D6F94C" blockId="17.[264,1326,284,1750]" pageId="17" pageNumber="18">
<emphasis id="C1E5EA83E249FFF8FED4351E8DDCFADF" box="[312,489,1324,1349]" italics="true" pageId="17" pageNumber="18">Natural history</emphasis>
.—
<taxonomicName id="34914D12E249FFF8FDF9351E8EFDFADF" box="[533,712,1324,1349]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E249FFF8FDF9351E8EFDFADF" box="[533,712,1324,1349]" italics="true" pageId="17" pageNumber="18">W. submontana</emphasis>
</taxonomicName>
occurs in an altitudinal range of about
<quantity id="34699B74E249FFF8FB63351E891CFADC" box="[1167,1321,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="17" pageNumber="18" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">800–1,200 m</quantity>
a.s.l., occasionally it may be found at lower altitudes (see below). This species was found in broad daylight either under rocks in a partly dried up river basin (Kodmin;
<figureCitation id="6BAA2A14E249FFF8FB74354E88DEFA0C" box="[1176,1259,1404,1430]" captionStart="FIGURE 11" captionStartId="18.[264,367,1558,1582]" captionTargetBox="[459,1095,310,1537]" captionTargetId="figure@18.[459,1095,309,1539]" captionTargetPageId="18" captionText="FIGURE 11: Habitats of Werneria submontana nov. sp. on the Bakossi Mts.; a: Edib Hills, surroundings of Lake Edib (type locality); b: small forest creek on Mwendelengo Mts., Kodmin, several of the paratypes have been collected beneath the stones on the creek’s bank; c: watercatchments at Mt. Kupe; the toads were living in the spray zone of the small, artificial waterfall." httpUri="https://zenodo.org/record/169506/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
b), or on stony ground between wet, very low vegetation which was lying in the spray area of a small, artificial waterfall (Mt. Kupe;
<figureCitation id="6BAA2A14E249FFF8FD5735FE8F3AFA7C" box="[699,783,1484,1510]" captionStart="FIGURE 11" captionStartId="18.[264,367,1558,1582]" captionTargetBox="[459,1095,310,1537]" captionTargetId="figure@18.[459,1095,309,1539]" captionTargetPageId="18" captionText="FIGURE 11: Habitats of Werneria submontana nov. sp. on the Bakossi Mts.; a: Edib Hills, surroundings of Lake Edib (type locality); b: small forest creek on Mwendelengo Mts., Kodmin, several of the paratypes have been collected beneath the stones on the creek’s bank; c: watercatchments at Mt. Kupe; the toads were living in the spray zone of the small, artificial waterfall." httpUri="https://zenodo.org/record/169506/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
c). The habitat at the base of Mt. Kupe, where this species seem to live in sympatry with
<taxonomicName id="34914D12E249FFF8FCEF35C68F4CF997" box="[771,889,1524,1549]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="preussi">
<emphasis id="C1E5EA83E249FFF8FCEF35C68F4CF997" box="[771,889,1524,1549]" italics="true" pageId="17" pageNumber="18">W. preussi</emphasis>
</taxonomicName>
(see above), was within a transition zone between good quality secondary forest, and more­or­less undisturbed primary forest at
<quantity id="34699B74E249FFF8FEC936768D5AF9C4" box="[293,367,1604,1630]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="17" pageNumber="18" unit="m" value="910.0">910 m</quantity>
a.s.l. M. Gartshore (pers. comm.) found these toads near the same locality a little lower at
<quantity id="34699B74E249FFF8FE82365E8D82F91C" box="[366,439,1644,1670]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="17" pageNumber="18" unit="m" value="850.0">850 m</quantity>
a.s.l., under rocks during the day along a stream on the grounds of the Nyasoso Girls Secondary School, above the school reservoir, just outside town. The very wrinkled throat of adult males may indicate that this species utters an advertisement call.
</paragraph>
</subSubSection>
<caption id="A7EE6619E24AFFFBFEE43624893DF90F" httpUri="https://zenodo.org/record/169506/files/figure.png" pageId="18" pageNumber="19" targetBox="[459,1095,310,1537]" targetPageId="18">
<paragraph id="F32E3691E24AFFFBFEE43624893DF90F" blockId="18.[264,1322,1558,1685]" pageId="18" pageNumber="19">
<emphasis id="C1E5EA83E24AFFFBFEE436248DA7F9B4" bold="true" box="[264,402,1558,1582]" pageId="18" pageNumber="19">FIGURE 11</emphasis>
: Habitats of
<taxonomicName id="34914D12E24AFFFBFDC836248F27F9B7" box="[548,786,1558,1581]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E24AFFFBFDC836248F27F9B7" box="[548,786,1558,1581]" italics="true" pageId="18" pageNumber="19">Werneria submontana</emphasis>
</taxonomicName>
nov. sp. on the Bakossi Mts.; a: Edib Hills, surroundings of Lake Edib (type locality); b: small forest creek on Mwendelengo Mts., Kodmin, several of the paratypes have been collected beneath the stones on the creek’s bank; c: watercatchments at Mt. Kupe; the toads were living in the spray zone of the small, artificial waterfall.
</paragraph>
</caption>
<subSubSection id="BB8B651AE24AFFFAFED436DE8FD1FE64" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="distribution">
<paragraph id="F32E3691E24AFFFAFED436DE8FD1FE64" blockId="18.[264,1324,1771,1877]" lastBlockId="19.[264,1323,284,590]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="C1E5EA83E24AFFFBFED436DE8DF2F89F" box="[312,455,1772,1797]" italics="true" pageId="18" pageNumber="19">Distribution</emphasis>
.— Known from the Bakossi Mts., Mwendelengo Mts. and Nyasoso at the base of Mt. Kupe (
<figureCitation id="6BAA2A14E24AFFFBFE0937218E1BF8B7" box="[485,558,1811,1837]" captionStart="FIGURE 5" captionStartId="8.[264,367,1482,1506]" captionTargetBox="[294,1293,510,1469]" captionTargetId="figure@8.[294,1294,509,1469]" captionTargetPageId="8" captionText="FIGURE 5: Distribution of known Werneria species; compare with text." httpUri="https://zenodo.org/record/169503/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
). Judging from pictures kindly provided by M. Gartshore, the
<specimenCount id="E597FD18E24AFFFBFAE137218D10F8CF" pageId="18" pageNumber="19" type="generic">
38
<taxonomicName id="34914D12E24AFFFBFEE4370E8D10F8CF" box="[264,293,1852,1877]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="mertensiana">
<emphasis id="C1E5EA83E24AFFFBFEE4370E8D10F8CF" box="[264,293,1852,1877]" italics="true" pageId="18" pageNumber="19">W.</emphasis>
</taxonomicName>
</specimenCount>
mertensiana specimens recorded by her in February and April in the streams around Nyasoso at
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on the western slope of Mt. Kupe (
<bibRefCitation id="97004B60E24BFFFAFC96312E8806FEAC" author="Gartshore" box="[890,1075,284,310]" pageId="19" pageNumber="26" refString="Gartshore, M. (1984) The status of the montane herpetofauna of the Cameroon highlands. In: Stuart, S. N. (Ed.) Conservation of Cameroon Montane Forests, International Council for Bird Preservation, London, 204 - 263." type="book chapter" year="1984">Gartshore 1984</bibRefCitation>
) are
<taxonomicName id="34914D12E24BFFFAFB83312E8914FEAF" box="[1135,1313,284,309]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E24BFFFAFB83312E8914FEAF" box="[1135,1313,284,309]" italics="true" pageId="19" pageNumber="20">W. submontana</emphasis>
</taxonomicName>
. Likewise a specimen collected by
<bibRefCitation id="97004B60E24BFFFAFD4231768F7CFEC4" author="Amiet" box="[686,841,324,350]" pageId="19" pageNumber="25" refString="Amiet, J. - L. (1975) Ecologie et distribution des Amphibiens Anoures de la region de Nkongsamba (Cameroun). Annales de la Faculte des Sciences du Cameroun, 20, 33 - 107." type="journal article" year="1975">Amiet (1975</bibRefCitation>
,
<bibRefCitation id="97004B60E24BFFFAFCB531768F91FEC4" author="Amiet" box="[857,932,324,350]" pageId="19" pageNumber="25" refString="Amiet, J. - L. (1976 b) Observations anatomiques et biologiques sur le genre Werneria Poche, 1903. Revue de Zoologie africaine, 90, 33 - 45." type="journal article" year="1976" yearSuffix="b">1976b</bibRefCitation>
) in the vicinity of Nkongsamba (Maholé,
<quantity id="34699B74E24BFFFAFE96315E8DF0FE1C" box="[378,453,364,390]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="km" value="10.0">10 km</quantity>
north­west of Tombel, forest reserve of Bakossi, primary rainforest,
<quantity id="34699B74E24BFFFAFB1C315E8D63FE34" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="19" pageNumber="20" unit="m" value="325.0" valueMax="350.0" valueMin="300.0">300– 350 m</quantity>
a.s.l.), may belong to this new species.
<taxonomicName id="34914D12E24BFFFAFCAA31A68FC8FE37" box="[838,1021,404,429]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E24BFFFAFCAA31A68FC8FE37" box="[838,1021,404,429]" italics="true" pageId="19" pageNumber="20">W. submontana</emphasis>
</taxonomicName>
might also occur on the Rumpi Hills, as characters of three preserved specimens from Mofako showed characters that best fit this new species (photos provided by J.­L. Amiet).
</paragraph>
</subSubSection>
<subSubSection id="BB8B651AE24BFFFAFED4323E8D08FDD4" pageId="19" pageNumber="20" type="etymology">
<paragraph id="F32E3691E24BFFFAFED4323E8D08FDD4" blockId="19.[264,1323,284,590]" pageId="19" pageNumber="20">
<emphasis id="C1E5EA83E24BFFFAFED4323E8D86FDBF" box="[312,435,524,549]" italics="true" pageId="19" pageNumber="20">Etymology</emphasis>
.— The name
<taxonomicName id="34914D12E24BFFFAFD8F323E8EC7FDBF" box="[611,754,524,549]" class="Amphibia" family="Bufonidae" genus="Werneria" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="submontana">
<emphasis id="C1E5EA83E24BFFFAFD8F323E8EC7FDBF" box="[611,754,524,549]" italics="true" pageId="19" pageNumber="20">submontana</emphasis>
</taxonomicName>
indicates the altitudinal range of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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