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403 lines
50 KiB
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<mods:title id="2910BC27432E0EA77F81A70DE344FFAB">First report of Early Eocene Decapods in Morocco: description of a new genus and a new species of Carpiliidae (Decapoda: Brachyura) with remarks on its paleobiogeography</mods:title>
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<mods:namePart id="6DBAF938C63F1183DDCF052FBDD956B5">Ossó, Àlex</mods:namePart>
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<mods:namePart id="AB2232006617C9D0EA822594F76B25B5">Gagnaison, Cyril</mods:namePart>
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<mods:namePart id="10CDBDAFF1BF3779F9C2BF13B594697C">Bailleul, Julien</mods:namePart>
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<mods:title id="BB3D217DABB01CC4B61B5B04D53EF439">Geodiversitas</mods:title>
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<mods:part id="D92CBBCEC301452E81F98C6431FA3AC2">
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<mods:date id="DB3AAD4D77F9FF6AB8C1F2E182489B6F">2020</mods:date>
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<mods:number id="56D5AA503FD055C6C802A70006388A06">2020-02-27</mods:number>
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<mods:number id="DBF3A607B0771CC76158AB42089502A9">42</mods:number>
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<mods:identifier id="3DD711DED28D7E69DC88AE5E19861E34" type="DOI">10.5252/geodiversitas2020v42a4</mods:identifier>
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<subSection id="E2C62F06FFF05975FED4A5D9FBE6F836" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="multiple">
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<treatment id="03E087FBFFF05975FED4A5D9FBE6F836" ID-GBIF-Taxon="162385043" LSID="urn:lsid:zoobank.org:act:4A10F6F4-A9CA-4C7A-B679-E10D5266A77C" httpUri="http://treatment.plazi.org/id/03E087FBFFF05975FED4A5D9FBE6F836" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
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<subSubSection id="C3536566FFF05976FED4A5D9FDFCFA6B" box="[295,613,1429,1456]" pageId="5" pageNumber="50" type="nomenclature">
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<paragraph id="8BF636EDFFF05976FED4A5D9FDFCFA6B" blockId="5.[295,613,1429,1456]" box="[295,613,1429,1456]" pageId="5" pageNumber="50">
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<heading id="D0BE8181FFF05976FED4A5D9FDFCFA6B" box="[295,613,1429,1456]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="50" reason="2">
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Genus
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<taxonomicName id="4C494D6EFFF05976FE87A5D9FDFCFA6B" ID-CoL="03388ab7-8bcf-4181-b8f5-1299eaae9a2a" authority="Ossó & Gagnaison & Bailleul, 2020" authorityName="Ossó & Gagnaison & Bailleul" authorityYear="2020" box="[372,613,1429,1456]" class="Malacostraca" family="Carpiliidae" genus="Maurocarpilius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="B93DEAFFFFF05976FE87A5D9FD8AFA74" box="[372,531,1429,1455]" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
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<taxonomicNameLabel id="A20E5784FFF05976FDE9A5DAFDF8FA6B" box="[538,609,1430,1456]" pageId="5" pageNumber="50" rank="genus">n. gen</taxonomicNameLabel>
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.
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3536566FFF05976FF77A581FC9FFA3A" box="[132,774,1485,1505]" pageId="5" pageNumber="50" type="description">
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<paragraph id="8BF636EDFFF05976FF77A581FC9FFA3A" blockId="5.[132,774,1485,1505]" box="[132,774,1485,1505]" pageId="5" pageNumber="50">
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urn:lsid:zoobank.org:act:
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<uuid id="FFEF0C38FFF05976FEAEA581FC9FFA3A" box="[349,774,1485,1505]" pageId="5" pageNumber="50">4A10F6F4-A9CA-4C7A-B679-E10D5266A77C</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<paragraph id="8BF636EDFFF05976FF77A64BFC9EF996" blockId="5.[132,774,1544,1567]" lastBlockId="5.[132,775,1589,1692]" pageId="5" pageNumber="50">
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<subSubSection id="C3536566FFF05976FF77A64BFC9FF9FB" box="[132,774,1543,1568]" pageId="5" pageNumber="50" type="materials_examined">
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<materialsCitation id="3B213CB0FFF05976FF77A64BFC9FF9FB" ID-GBIF-Occurrence="2573323301" box="[132,774,1543,1568]" pageId="5" pageNumber="50" typeStatus="holotype">
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<typeStatus id="54F2884FFFF05976FF77A64BFF2DF9C7" box="[132,180,1543,1567]" pageId="5" pageNumber="50">
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<collectionCode id="ED58AE28FFF05976FF77A64BFF2DF9C7" box="[132,180,1543,1567]" pageId="5" pageNumber="50">TYPE</collectionCode>
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</typeStatus>
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SPECIES. —
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<taxonomicName id="4C494D6EFFF05976FED0A64BFD9EF9C4" authority="Ossó & Gagnaison & Bailleul, 2020" authorityName="Ossó & Gagnaison & Bailleul" authorityYear="2020" box="[291,519,1543,1567]" class="Malacostraca" family="Carpiliidae" genus="Maurocarpilius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="binodosus" status="n. gen., n. sp.">
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<emphasis id="B93DEAFFFFF05976FED0A64BFD9EF9C4" box="[291,519,1543,1567]" italics="true" pageId="5" pageNumber="50">Maurocarpilius binodosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E5784FFF05976FDFFA644FD1EF9C4" box="[524,647,1544,1568]" pageId="5" pageNumber="50" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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by monotypy
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</materialsCitation>
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</subSubSection>
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<subSubSection id="C3536566FFF05976FF77A678FC9EF996" box="[132,775,1588,1613]" pageId="5" pageNumber="50" type="etymology">
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ETYMOLOGY. — From
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<emphasis id="B93DEAFFFFF05976FE99A679FE3DF997" box="[362,420,1589,1612]" italics="true" pageId="5" pageNumber="50">mauri</emphasis>
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, the name given to the people who
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</subSubSection>
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</paragraph>
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<paragraph id="8BF636EDFFF05976FF77A603FE35F947" blockId="5.[132,775,1589,1692]" pageId="5" pageNumber="50">
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lived in
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<collectingCountry id="F35E767DFFF05976FF20A603FED9F9BC" box="[211,320,1615,1639]" name="Mauritania" pageId="5" pageNumber="50">Mauritania</collectingCountry>
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Province, during the Roman Empire, which is the part of the current
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<collectingCountry id="F35E767DFFF05976FE92A626FE20F95A" box="[353,441,1642,1665]" name="Morocco" pageId="5" pageNumber="50">Morocco</collectingCountry>
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where the studied specimens were recovered. Gender: masculine.
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</paragraph>
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<subSubSection id="C3536566FFF05976FF77A6FDFE72F836" pageId="5" pageNumber="50" type="diagnosis">
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<paragraph id="8BF636EDFFF05976FF77A6FDFE72F836" blockId="5.[132,776,1713,2029]" pageId="5" pageNumber="50">DIAGNOSIS. — Carapace transversely ovate, wider than long, L/W ratio about 0.72, smooth. Vaulted in both senses, mainly anteriorly. Maximum width at half-length of carapace. Front slightly subtriangular, bilobed, strongly downturned. Orbits small, rounded, supraorbital margin entire, slightly rimmed. Anterolateral margin strongly convex, acute, bearing two small nodes, one at half-length level and a second at the angle with the posterolateral margin. Posterolateral margin concave, entire, acute in anterior half, starting about 60° respect to the axis. Posterior margin slightly convex. Regions not defined. Branchiocardiac grooves defined by muscle scars. Gastric pits visible. Ventral features and appendages no preserved.</paragraph>
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</subSubSection>
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<subSubSection id="C3536566FFF05975FCDEA095FBE6F836" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="discussion">
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||
<paragraph id="8BF636EDFFF05976FCDEA095FC08FF2A" blockId="5.[813,1456,217,2030]" box="[813,913,217,243]" pageId="5" pageNumber="50">REMARKS</paragraph>
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<paragraph id="8BF636EDFFF05976FCDEA0B5FC47FCCA" blockId="5.[813,1456,217,2030]" pageId="5" pageNumber="50">
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Arose in the early Eocene carpiliids are defined by a transversely ovate and broad carapace with well demarcated margins, a usually smooth surface of carapace, anterolateral margins entire or with blunt teeth or nodes, and from poorly to not defined regions (e.g.
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<bibRefCitation id="EFD84B1CFFF05976FBF4A135FAFFFE48" author="KARASAWA H. & SCHWEITZER C. E." box="[1031,1382,377,403]" pageId="5" pageNumber="50" pagination="23 - 73" refId="ref7751" refString="KARASAWA H. & SCHWEITZER C. E. 2006. - A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology 75 (1 - 2): 23 - 73. https: // doi. org / 10.1163 / 18759866 - 0750102002" type="journal article" year="2006">Karasawa & Schweitzer 2006: 43</bibRefCitation>
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;
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<bibRefCitation id="EFD84B1CFFF05976FA83A135FC46FE68" author="DAVIE P. J. F. & GUINOT D. & NG P. K. L." pageId="5" pageNumber="50" pagination="1049 - 1130" refId="ref6958" refString="DAVIE P. J. F., GUINOT D. & NG P. K. L. 2015. - Systematics and classification of Brachyura, in CASTRO P., DAVIE P. J. F., GUINOT D., SCHRAM F. & VON VAUPEL KLEIN C. (eds), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea. Vol. 9 (C) (I), Decapoda: Brachyura (Part 2): 1049 - 1130. https: // doi. org / 10.1163 / 9789004190832 _ 021" type="book chapter" year="2015">
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Davie
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<emphasis id="B93DEAFFFFF05976FCDEA1D5FCC7FE69" box="[813,862,408,434]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
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2015: 1074
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</bibRefCitation>
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||
;
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<bibRefCitation id="EFD84B1CFFF05976FC1AA1D4FB71FE68" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[1001,1256,408,435]" pageId="5" pageNumber="50" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
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Schweitzer
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<emphasis id="B93DEAFFFFF05976FBACA1D5FB09FE69" box="[1119,1168,408,434]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
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2018: 2
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</bibRefCitation>
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). The inexpressiveness of the dorsal carapaces of members of
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<taxonomicName id="4C494D6EFFF05976FB00A1F4FAF7FE09" authorityName="Ortmann" authorityYear="1893" box="[1267,1390,440,466]" class="Malacostraca" family="Carpiliidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="family">Carpiliidae</taxonomicName>
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, often leaves comparisons between their different genera and species reduced to subtle details, as the number of anterolateral teeth, if present, to length/width or fronto-orbital ratios, shape of them, or the angle degrees of posterior margins.
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<bibRefCitation id="EFD84B1CFFF05976FAB1A274FC3EFDA9" author="FELDMANN R. M. & SCHWEITZER C. E. & BENNETT O. & FRANtESCU O. & RESAR N. & TRUDEAU A." pageId="5" pageNumber="50" pagination="323 - 353" refId="ref7468" refString="FELDMANN R. M., SCHWEITZER C. E., BENNETT O., FRANtESCU O., RESAR N. & TRUDEAU A. 2011. - Decapod crustaceans from the Eocene of Egypt. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlung 262: 323 - 353. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
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Feldmann
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<emphasis id="B93DEAFFFFF05976FCDEA215FCC6FDA9" box="[813,863,600,626]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
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(2011
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</bibRefCitation>
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||
: t. 3) in the revision of
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<emphasis id="B93DEAFFFFF05976FB56A214FAA6FDA9" box="[1189,1343,600,626]" italics="true" pageId="5" pageNumber="50">Palaeocarpilius</emphasis>
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A. Milne-
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<bibRefCitation id="EFD84B1CFFF05976FCDEA234FC4AFD49" author="EDWARDS A." box="[813,979,632,658]" pageId="5" pageNumber="50" pagination="31 - 85" refId="ref8109" refString="MILNE- EDWARDS A. 1862. - Monographie des Crustaces fossiles de la famille des Canceriens. Annales des Sciences naturelles, Zoologie 4 (18): 31 - 85. https: // biodiversitylibrary. org / page / 29146016" type="journal article" year="1862">Edwards, 1862</bibRefCitation>
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, summarized the aforementioned characters of all the genera placed within
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<taxonomicName id="4C494D6EFFF05976FB7FA2D4FA93FD69" authorityName="Ortmann" authorityYear="1893" box="[1164,1290,664,690]" class="Malacostraca" family="Carpiliidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="family">Carpiliidae</taxonomicName>
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, which will be used in the following comparisons as well. Therefore,
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<emphasis id="B93DEAFFFFF05976FA87A2F4FC00FD2A" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
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n. gen. is compared with all the known carpiliid genera as follow.
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</paragraph>
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<paragraph id="8BF636EDFFF05975FCB0A35BFB0CF8F5" blockId="5.[813,1456,217,2030]" lastBlockId="6.[813,1456,1461,2030]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
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<emphasis id="B93DEAFFFFF05976FCB0A35BFC7BFCEA" box="[835,994,791,817]" italics="true" pageId="5" pageNumber="50">Braggicarpilius</emphasis>
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||
<bibRefCitation id="EFD84B1CFFF05976FC18A35BFAF5FCE9" author="BESCHIN C. & BUSULINI A. & TESSIER G." box="[1003,1388,791,818]" pageId="5" pageNumber="50" pagination="47 - 109" refId="ref6407" refString="BESCHIN C., BUSULINI A. & TESSIER G. 2015. - Nuova segnalazione di crostacei associate a coralli nell'Eocene inferior dei Lessini orientali (Vestenanova-Verona). Lavori Societa Veneziana di Scienze Naturali 40: 47 - 109." type="journal article" year="2015">Beschin, Busulini & Tessier, 2015</bibRefCitation>
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, from the Ypresian of northern
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<collectingCountry id="F35E767DFFF05976FBBBA37BFBE0FC8A" box="[1096,1145,823,849]" name="Italy" pageId="5" pageNumber="50">Italy</collectingCountry>
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differs from
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<emphasis id="B93DEAFFFFF05976FAFEA37BFA36FC8A" box="[1293,1455,823,849]" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
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n. gen. in having entire anterolateral margins, and convex posterolateral margins (see
|
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<bibRefCitation id="EFD84B1CFFF05976FBABA33BFAB2FC49" author="BESCHIN C. & BUSULINI A. & TESSIER G." box="[1112,1323,887,914]" pageId="5" pageNumber="50" pagination="47 - 109" refId="ref6407" refString="BESCHIN C., BUSULINI A. & TESSIER G. 2015. - Nuova segnalazione di crostacei associate a coralli nell'Eocene inferior dei Lessini orientali (Vestenanova-Verona). Lavori Societa Veneziana di Scienze Naturali 40: 47 - 109." type="journal article" year="2015">
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||
Beschin
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<emphasis id="B93DEAFFFFF05976FB46A334FB72FC4A" box="[1205,1259,887,913]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
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||
2015
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</bibRefCitation>
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||
;
|
||
<bibRefCitation id="EFD84B1CFFF05976FAC8A33BFC24FC6A" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." pageId="5" pageNumber="50" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
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Schweitzer
|
||
<emphasis id="B93DEAFFFFF05976FCDEA3DBFCF8FC6A" box="[813,865,919,945]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2018: 3
|
||
</bibRefCitation>
|
||
, fig. 3.1). An array of carpiliids, such as
|
||
<emphasis id="B93DEAFFFFF05976FA84A3DBFC1CFC0A" italics="true" pageId="5" pageNumber="50">Bryocarpilius</emphasis>
|
||
Feldmann, Schweitzer, Bennet, Frantescu, Resar & Trudeau, 2011, from the Eocene;
|
||
<taxonomicName id="4C494D6EFFF05976FB57A39AFA32FC2A" authority="Rathbun, 1929" authorityName="Rathbun" authorityYear="1929" box="[1188,1451,982,1009]" class="Malacostraca" family="Carpiliidae" genus="Ocalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEAFFFFF05976FB57A39AFB6EFC2B" box="[1188,1271,982,1008]" italics="true" pageId="5" pageNumber="50">Ocalina</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FAF2A39AFA32FC2A" author="RATHBUN M. J." box="[1281,1451,982,1009]" pageId="5" pageNumber="50" pagination="1 - 4" refId="ref8324" refString="RATHBUN M. J. 1929. - A new crab from the Eocene of Florida. Proceedings of the U. S. National Museum 75: 1 - 4. https: // doi. org / 10.5479 / si. 00963801.75 - 2786.1" type="journal article" year="1929">Rathbun, 1929</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Eocene;
|
||
<emphasis id="B93DEAFFFFF05976FC01A3BAFB15FBCB" box="[1010,1164,1014,1040]" italics="true" pageId="5" pageNumber="50">Palaeocarpilius</emphasis>
|
||
, from the Eocene to Miocene; and
|
||
<taxonomicName id="4C494D6EFFF05976FC65A45AFCF1FB8B" authority="Beschin, Busulini, De Angeli & Tessier 2007" authorityName="Beschin, Busulini, De Angeli & Tessier" authorityYear="2007" class="Malacostraca" family="Carpiliidae" genus="Paraocalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEAFFFFF05976FC65A45AFB89FBEB" box="[918,1040,1046,1072]" italics="true" pageId="5" pageNumber="50">Paraocalina</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FBE6A45AFCF1FB8B" author="BESCHIN C. & BUSULINI A. & DE ANGELI A. & TESSIER G." pageId="5" pageNumber="50" refId="ref6347" refString="BESCHIN C., BUSULINI A., DE ANGELI A. & TESSIER G. 2007. - I decapodi dell'Eocene inferiore di Contrada Gecchelina (Vicenza, Italia settentrionale) (Anomura e Brachiura). Museo di Archeologia e Scienze Naturali " G. Zannato, " Montecchio Maggiore, Vicenza, Italy, 76 p." type="book" year="2007">Beschin, Busulini, De Angeli & Tessier 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Eocene, all of them with wide Tethysian and also Atlantic distribution;
|
||
<emphasis id="B93DEAFFFFF05976FBE6A41AFB3DFBAB" box="[1045,1188,1110,1136]" italics="true" pageId="5" pageNumber="50">Proxicarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FB59A41AFA32FBAB" author="COLLINS J. S. H. & MORRIS S. F." box="[1194,1451,1110,1136]" pageId="5" pageNumber="50" pagination="957 - 981" refId="ref6712" refString="COLLINS J. S. H. & MORRIS S. F. 1978. - New Lower Tertiary crabs from Pakistan. Palaeontology 21: 957 - 981. http: // www. biodiversitylibrary. org / page / 49759017" type="journal article" year="1978">Collins & Morris, 1978</bibRefCitation>
|
||
, from the Eocene of
|
||
<collectingCountry id="F35E767DFFF05976FC08A43AFBCEFB4B" box="[1019,1111,1142,1168]" name="Pakistan" pageId="5" pageNumber="50">Pakistan</collectingCountry>
|
||
; and
|
||
<emphasis id="B93DEAFFFFF05976FB7EA43AFABBFB4B" box="[1165,1314,1142,1168]" italics="true" pageId="5" pageNumber="50">Tethyscarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FADBA43BFC25FB6B" author="DE ANGELI A. & ALBERTI R." pageId="5" pageNumber="50" pagination="121 - 128" refId="ref7063" refString="DE ANGELI A. & ALBERTI R. 2016. - Tethyscarpilius bericus n. gen., n. sp. (Decapoda, Brachyura, Carpiliidae) dell'Eocene superiore dei Monti Berici (Vicenza, Italia settentrionale). Lavori Societa Veneziana di Scienze Naturali 41: 121 - 128." type="journal article" year="2016">De Angeli & Alberti, 2016</bibRefCitation>
|
||
, all those genera clearly differ from
|
||
<emphasis id="B93DEAFFFFF05976FAD4A4DAFCD3FB14" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
|
||
n. gen., in having the anterolateral margins armed from four, to up to nine nodes or teeth, instead of only two small nodes in the new genus and different ratios (see
|
||
<bibRefCitation id="EFD84B1CFFF05976FAB1A4B9FC3AFAEB" author="FELDMANN R. M. & SCHWEITZER C. E. & BENNETT O. & FRANtESCU O. & RESAR N. & TRUDEAU A." pageId="5" pageNumber="50" pagination="323 - 353" refId="ref7468" refString="FELDMANN R. M., SCHWEITZER C. E., BENNETT O., FRANtESCU O., RESAR N. & TRUDEAU A. 2011. - Decapod crustaceans from the Eocene of Egypt. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlung 262: 323 - 353. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B93DEAFFFFF05976FCDEA55AFCFBFAF4" box="[813,866,1301,1327]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
: t. 3), and furthermore, in the case of
|
||
<emphasis id="B93DEAFFFFF05976FAA7A559FA31FAF4" box="[1364,1448,1301,1327]" italics="true" pageId="5" pageNumber="50">Ocalina</emphasis>
|
||
, in having a tuberculated dorsal surface (see
|
||
<bibRefCitation id="EFD84B1CFFF05976FAF4A579FCFCFAB4" author="FELDMANN R. M. & SCHWEITZER C. E. & BENNETT O. & FRANtESCU O. & RESAR N. & TRUDEAU A." pageId="5" pageNumber="50" pagination="323 - 353" refId="ref7468" refString="FELDMANN R. M., SCHWEITZER C. E., BENNETT O., FRANtESCU O., RESAR N. & TRUDEAU A. 2011. - Decapod crustaceans from the Eocene of Egypt. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlung 262: 323 - 353. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B93DEAFFFFF05976FA88A57AFA29FA94" box="[1403,1456,1333,1359]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
; 345-346, fig. 13;
|
||
<bibRefCitation id="EFD84B1CFFF05976FBCBA519FB41FAB4" author="RATHBUN M. J." box="[1080,1240,1365,1391]" pageId="5" pageNumber="50" pagination="1 - 4" refId="ref8324" refString="RATHBUN M. J. 1929. - A new crab from the Eocene of Florida. Proceedings of the U. S. National Museum 75: 1 - 4. https: // doi. org / 10.5479 / si. 00963801.75 - 2786.1" type="journal article" year="1929">Rathbun 1929</bibRefCitation>
|
||
; A. Milne-Edwards 1862: 51-53;
|
||
<bibRefCitation id="EFD84B1CFFF05976FC35A539FAD0FA54" author="BESCHIN C. & DE ANGELI A." box="[966,1353,1397,1424]" pageId="5" pageNumber="50" pagination="11 - 23" refId="ref6449" refString="BESCHIN C. & DE ANGELI A. 2006. - Il genere Palaeocarpilius A. Milne-Edwards, 1862 (Decapoda, Brachyura, Carpiliidae) nel Terziario del Vicentino (Italia Settentrionale). Studi e Ricerche 13: 11 - 23." type="journal article" year="2006">Beschin & De Angeli 2006: 11-23</bibRefCitation>
|
||
, figs 2-5, pls 1-4;
|
||
<bibRefCitation id="EFD84B1CFFF05976FC79A5D9FB2CFA74" author="BESCHIN C. & BUSULINI A. & DE ANGELI A. & TESSIER G." box="[906,1205,1429,1456]" pageId="5" pageNumber="50" refId="ref6347" refString="BESCHIN C., BUSULINI A., DE ANGELI A. & TESSIER G. 2007. - I decapodi dell'Eocene inferiore di Contrada Gecchelina (Vicenza, Italia settentrionale) (Anomura e Brachiura). Museo di Archeologia e Scienze Naturali " G. Zannato, " Montecchio Maggiore, Vicenza, Italy, 76 p." type="book" year="2007">
|
||
Beschin
|
||
<emphasis id="B93DEAFFFFF05976FC1BA5DAFB86FA74" box="[1000,1055,1429,1455]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2007: 42-43
|
||
</bibRefCitation>
|
||
, pl. 6, figs 1b, b;
|
||
<bibRefCitation id="EFD84B1CFFF05976FA8DA5D9FBD9FA14" author="COLLINS J. S. H. & MORRIS S. F." pageId="5" pageNumber="50" pagination="957 - 981" refId="ref6712" refString="COLLINS J. S. H. & MORRIS S. F. 1978. - New Lower Tertiary crabs from Pakistan. Palaeontology 21: 957 - 981. http: // www. biodiversitylibrary. org / page / 49759017" type="journal article" year="1978">Collins & Morris 1978: 972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B1CFFF05976FBBFA5F9FA32FA14" author="SCHWEITZER C. E. & FELDMANN R. M. & GINGERICH P. D." box="[1100,1451,1461,1487]" pageId="5" pageNumber="50" pagination="89 - 118" refId="ref8444" refString="SCHWEITZER C. E., FELDMANN R. M. & GINGERICH P. D. 2004. - New Decapoda (Crustacea) from the Middle and Late Eocene of Pakistan and a revision of Lobonotus A. Milne Edwards, 1864. Contributions from the Museum of Paleontology, University of Michigan 31 (4): 89 - 118." type="journal article" year="2004">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF05976FB34A5FAFB65FA14" box="[1223,1276,1461,1487]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2004: 100-102
|
||
</bibRefCitation>
|
||
, fig. 6;
|
||
<bibRefCitation id="EFD84B1CFFF05976FC85A599FBF0FA34" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[886,1129,1493,1519]" pageId="5" pageNumber="50" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF05976FC00A599FBB1FA35" box="[1011,1064,1493,1519]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B1CFFF05976FB8AA599FA06FA34" author="DE ANGELI A. & ALBERTI R." box="[1145,1439,1493,1519]" pageId="5" pageNumber="50" pagination="121 - 128" refId="ref7063" refString="DE ANGELI A. & ALBERTI R. 2016. - Tethyscarpilius bericus n. gen., n. sp. (Decapoda, Brachyura, Carpiliidae) dell'Eocene superiore dei Monti Berici (Vicenza, Italia settentrionale). Lavori Societa Veneziana di Scienze Naturali 41: 121 - 128." type="journal article" year="2016">De Angeli & Alberti 2016</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C494D6EFFF05976FCDEA5B9FBD4F9D4" authority="Desmarest, 1823" authorityName="Desmarest" authorityYear="1823" box="[813,1101,1525,1551]" class="Malacostraca" family="Carpiliidae" genus="Carpilius" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93DEAFFFFF05976FCDEA5B9FC14F9D4" box="[813,909,1525,1551]" italics="true" pageId="5" pageNumber="50">Carpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FC60A5B9FBD4F9D4" author="DESMAREST A. G." box="[915,1101,1525,1551]" pageId="5" pageNumber="50" pagination="211 - 285" refId="ref7244" refString="DESMAREST A. G. 1823. - Malacostraces, Malacostraca (Crust.), in CUVIER F. ed., Dictionnaire des Sciences naturelles. Vol. 28. F. G. Levrault & Le Normant, Strasbourg, Paris: 211 - 285. https: // biodiversitylibrary. org / page / 25300024" type="book chapter" year="1823">Desmarest, 1823</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the early Eocene to recent, differs from
|
||
<emphasis id="B93DEAFFFFF05976FC41A658FBC8F9F5" box="[946,1105,1556,1582]" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
|
||
n. gen. in having only one anterolateral tooth instead of two, and a short but well-developed oblique carina that extends from that tooth (epibranchial) to the dorsal surface (see
|
||
<bibRefCitation id="EFD84B1CFFF05976FBEBA639FAABF955" author="DESMAREST A. G." box="[1048,1330,1652,1679]" pageId="5" pageNumber="50" pagination="211 - 285" refId="ref7244" refString="DESMAREST A. G. 1823. - Malacostraces, Malacostraca (Crust.), in CUVIER F. ed., Dictionnaire des Sciences naturelles. Vol. 28. F. G. Levrault & Le Normant, Strasbourg, Paris: 211 - 285. https: // biodiversitylibrary. org / page / 25300024" type="book chapter" year="1823">Desmarest 1823: 227-228</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B1CFFF05976FACFA638FC2DF975" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." pageId="5" pageNumber="50" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF05976FCDEA6D9FCC7F975" box="[813,862,1684,1710]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2018: 3
|
||
</bibRefCitation>
|
||
, fig. 2a-b).
|
||
<emphasis id="B93DEAFFFFF05976FBD9A6D8FB5EF975" box="[1066,1223,1684,1710]" italics="true" pageId="5" pageNumber="50">Corallicarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FB3EA6D9FCF1F914" author="DE ANGELI A. & CECCON L." pageId="5" pageNumber="50" pagination="119 - 138" refId="ref7116" refString="DE ANGELI A. & CECCON L. 2015. - Nuovi Crostacei brachiuri dell'Eocene di Monte Magre (Vicenza, Italia settentrionale). Lavori Societa Veneziana di Scienze Naturali 40: 119 - 138." type="journal article" year="2015">De Angeli & Ceccon, 2015</bibRefCitation>
|
||
, from the early Eocene of northern
|
||
<collectingCountry id="F35E767DFFF05976FB15A6F8FA8EF915" box="[1254,1303,1716,1742]" name="Italy" pageId="5" pageNumber="50">Italy</collectingCountry>
|
||
, differs clearly from
|
||
<emphasis id="B93DEAFFFFF05976FC99A698FB90F935" box="[874,1033,1748,1774]" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
|
||
n. gen. in having a broader and more ovate carapace, without distinction between the posterolateral and posterior margin, and in having the dorsal surface ornated with irregular tubercles, instead of smooth in the new genus (see
|
||
<bibRefCitation id="EFD84B1CFFF05976FC27A718FAFCF8B5" author="DE ANGELI A. & CECCON L." box="[980,1381,1875,1902]" pageId="5" pageNumber="50" pagination="119 - 138" refId="ref7116" refString="DE ANGELI A. & CECCON L. 2015. - Nuovi Crostacei brachiuri dell'Eocene di Monte Magre (Vicenza, Italia settentrionale). Lavori Societa Veneziana di Scienze Naturali 40: 119 - 138." type="journal article" year="2015">De Angeli & Ceccon 2015: 125-126</bibRefCitation>
|
||
, fig. 4;
|
||
<bibRefCitation id="EFD84B1CFFF05976FCDEA73FFBA7F855" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[813,1086,1907,1934]" pageId="5" pageNumber="50" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF05976FC5AA738FC47F856" box="[937,990,1907,1933]" italics="true" pageId="5" pageNumber="50">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, fig. 3, 3a-b).
|
||
<emphasis id="B93DEAFFFFF05976FB2EA73FFAC9F856" box="[1245,1360,1907,1933]" italics="true" pageId="5" pageNumber="50">Eocarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF05976FAABA73FFC4EF876" author="BLOW W. C. & MANNING R. C." pageId="5" pageNumber="50" pagination="1 - 26" refId="ref6555" refString="BLOW W. C. & MANNING R. C. 1996. - Preliminary descriptions of 25 new decapod crustaceans from the Middle Eocene of the Carolinas, U. S. A. Tulane Studies in Geology and Paleontology 29 (1): 1 - 26." type="journal article" year="1996">Blow & Manning, 1996</bibRefCitation>
|
||
from the Eocene of eastern North America and westernmost Tethys, differs from
|
||
<emphasis id="B93DEAFFFFF05976FB33A7FFFAC6F816" box="[1216,1375,1971,1997]" italics="true" pageId="5" pageNumber="50">Maurocarpilius</emphasis>
|
||
n. gen. in having a longer ovoid carapace, anterolateral margins entire with only a small protuberance at the corner with antero- and posterolateral margin, posterolateral margins straight, and by its punctulate surface (see
|
||
<bibRefCitation id="EFD84B1CFFF35975FE5BA5B9FC9BF9D4" author="BLOW W. C. & MANNING R. C." box="[424,770,1525,1551]" pageId="6" pageNumber="51" pagination="1 - 26" refId="ref6555" refString="BLOW W. C. & MANNING R. C. 1996. - Preliminary descriptions of 25 new decapod crustaceans from the Middle Eocene of the Carolinas, U. S. A. Tulane Studies in Geology and Paleontology 29 (1): 1 - 26." type="journal article" year="1996">Blow & Manning 1996: 20-22</bibRefCitation>
|
||
, pl. 5, figs 2a, 2b;
|
||
<bibRefCitation id="EFD84B1CFFF35975FEBBA658FD1CF9F4" author="FELDMANN R. M. & BICE K. L. & SCHWEITZER C. E. & SALVA E. W. & PICKFORD K." box="[328,645,1556,1583]" pageId="6" pageNumber="51" pagination="1 - 28" refId="ref7403" refString="FELDMANN R. M., BICE K. L., SCHWEITZER C. E., SALVA E. W. & PICKFORD K. 1998. - Decapod crustaceans from the Eocene Castle Hayne Formation, North Carolina: paleoceanographic implications. Journal of Paleontology 48: 1 - 28. https: // doi. org / 10.1017 / S 0022336000059916" type="journal article" year="1998">
|
||
Feldmann
|
||
<emphasis id="B93DEAFFFFF35975FE4EA659FE6BF9F5" box="[445,498,1556,1582]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
1998: 11-12
|
||
</bibRefCitation>
|
||
, figs 13-14;
|
||
<bibRefCitation id="EFD84B1CFFF35975FF77A678FE0BF994" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[132,402,1588,1615]" pageId="6" pageNumber="51" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF35975FF0CA679FEAAF995" box="[255,307,1588,1614]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, fig. 3, 3a-b).
|
||
<emphasis id="B93DEAFFFFF35975FDDDA678FD27F995" box="[558,702,1588,1614]" italics="true" pageId="6" pageNumber="51">Holcocarcinus</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF35975FD36A678FF72F9B5" author="WITHERS T. H." pageId="6" pageNumber="51" pagination="94 - 97" refId="ref8774" refString="WITHERS T. H. 1924. - Eocene brachyurous decapod Crustacea from Nigeria. Annals and Magazine of Natural History (9) 13: 94 - 97. https: // doi. org / 10.1080 / 00222932408633010" type="journal article" year="1924">Withers, 1924</bibRefCitation>
|
||
, from the Eocene of
|
||
<collectingCountry id="F35E767DFFF35975FE3FA618FD86F9B5" box="[460,543,1620,1646]" name="Nigeria" pageId="6" pageNumber="51">Nigeria</collectingCountry>
|
||
, differs clearly for the new genus in having the carapace with two marked transverse dorsal ridges (see
|
||
<bibRefCitation id="EFD84B1CFFF35975FEC9A6D8FE6FF975" author="WITHERS T. H." box="[314,502,1684,1711]" pageId="6" pageNumber="51" pagination="94 - 97" refId="ref8774" refString="WITHERS T. H. 1924. - Eocene brachyurous decapod Crustacea from Nigeria. Annals and Magazine of Natural History (9) 13: 94 - 97. https: // doi. org / 10.1080 / 00222932408633010" type="journal article" year="1924">Withers 1924: 94</bibRefCitation>
|
||
, pl. 5, figs 1-2;
|
||
<bibRefCitation id="EFD84B1CFFF35975FD67A6D8FE8FF914" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." pageId="6" pageNumber="51" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF35975FF77A6F9FF20F915" box="[132,185,1716,1742]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, fig. 4.1).
|
||
<emphasis id="B93DEAFFFFF35975FE7BA6F8FD92F915" box="[392,523,1716,1742]" italics="true" pageId="6" pageNumber="51">Laticarpilius</emphasis>
|
||
Feldmann, Schweitzer, Bennet, Frantescu, Resar & Trudeau, 2011, from the Eocene of
|
||
<collectingCountry id="F35E767DFFF35975FF57A6B8FF70F8D5" box="[164,233,1780,1806]" name="Egypt" pageId="6" pageNumber="51">Egypt</collectingCountry>
|
||
, differs from
|
||
<emphasis id="B93DEAFFFFF35975FE70A6B8FDBEF8D5" box="[387,551,1780,1806]" italics="true" pageId="6" pageNumber="51">Maurocarpilius</emphasis>
|
||
n. gen. in having a broader carapace, and a less downturned and completely triangular front (see
|
||
<bibRefCitation id="EFD84B1CFFF35975FEACA77FFD28F895" author="FELDMANN R. M. & SCHWEITZER C. E. & BENNETT O. & FRANtESCU O. & RESAR N. & TRUDEAU A." box="[351,689,1843,1870]" pageId="6" pageNumber="51" pagination="323 - 353" refId="ref7468" refString="FELDMANN R. M., SCHWEITZER C. E., BENNETT O., FRANtESCU O., RESAR N. & TRUDEAU A. 2011. - Decapod crustaceans from the Eocene of Egypt. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlung 262: 323 - 353. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B93DEAFFFFF35975FE20A778FD9FF896" box="[467,518,1843,1869]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2011: 342-344
|
||
</bibRefCitation>
|
||
, fig. 12;
|
||
<bibRefCitation id="EFD84B1CFFF35975FF77A71FFE13F8B5" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[132,394,1875,1902]" pageId="6" pageNumber="51" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF35975FF0EA718FEB6F8B6" box="[253,303,1875,1901]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, figs 4, 2a, 2b).
|
||
<emphasis id="B93DEAFFFFF35975FDC2A71FFD10F8B6" box="[561,649,1875,1901]" italics="true" pageId="6" pageNumber="51">Liopsalis</emphasis>
|
||
von
|
||
<bibRefCitation id="EFD84B1CFFF35975FD4EA718FF25F856" author="MEYER H. VON" pageId="6" pageNumber="51" pagination="147 - 178" refId="ref8073" refString="MEYER H. VON 1862. - Tertiaere Decapoden aus den Alpen, von Oeningen und dem Taunus. Palaeontographica 10: 147 - 178. https: // biodiversitylibrary. org / page / 35372059" type="journal article" year="1862">Meyer, 1862</bibRefCitation>
|
||
from the Eocene of western Tethys and
|
||
<collectingCountry id="F35E767DFFF35975FD8FA73FFD2EF856" box="[636,695,1907,1933]" name="India" pageId="6" pageNumber="51">India</collectingCountry>
|
||
, differs from the new genus in having a completely entire margins, and a triangular front not lobed (see von
|
||
<bibRefCitation id="EFD84B1CFFF35975FDB1A7F8FF29F836" author="MEYER H. VON" pageId="6" pageNumber="51" pagination="147 - 178" refId="ref8073" refString="MEYER H. VON 1862. - Tertiaere Decapoden aus den Alpen, von Oeningen und dem Taunus. Palaeontographica 10: 147 - 178. https: // biodiversitylibrary. org / page / 35372059" type="journal article" year="1862">Meyer 1862: 161- 163</bibRefCitation>
|
||
, pl. 17, figs 9-12;
|
||
<bibRefCitation id="EFD84B1CFFF35975FE85A79FFD8CF836" author="STOLICZKA F." box="[374,533,2003,2029]" pageId="6" pageNumber="51" pagination="1 - 16" refId="ref8587" refString="STOLICZKA F. 1871. - Observations of fossil crabs from Tertiary deposits in Sind and Kutch. Memoirs of the Geological Survey of India, Palaeontologica Indica 7: 1 - 16." type="journal article" year="1871">Stoliczka 1871</bibRefCitation>
|
||
pl. 5, figs 6, 6a-b;
|
||
<bibRefCitation id="EFD84B1CFFF35975FD11A79FFC48FA14" author="VIA L." box="[738,977,1461,2029]" pageId="6" pageNumber="51" pagination="1 - 479" refId="ref8753" refString="VIA L. 1969. - Crustaceos Decapodos del Eoceno espanol. Pirineos 91 - 94: 1 - 479." type="journal article" year="1969">Vía 1969: 231-237</bibRefCitation>
|
||
, fig. 28, t. 23, figs 3-4;
|
||
<bibRefCitation id="EFD84B1CFFF35975FB35A5F9FC15FA2B" author="FELDMANN R. M. & SCHWEITZER C. E. & BENNETT O. & FRANtESCU O. & RESAR N. & TRUDEAU A." pageId="6" pageNumber="51" pagination="323 - 353" refId="ref7468" refString="FELDMANN R. M., SCHWEITZER C. E., BENNETT O., FRANtESCU O., RESAR N. & TRUDEAU A. 2011. - Decapod crustaceans from the Eocene of Egypt. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlung 262: 323 - 353. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0202" type="journal article" year="2011">
|
||
Feldmann
|
||
<emphasis id="B93DEAFFFFF35975FACAA5FAFAF2FA14" box="[1337,1387,1461,1487]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2011: 344-345
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B1CFFF35975FC64A599FB04FA2B" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." box="[919,1181,1493,1520]" pageId="6" pageNumber="51" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF35975FBE3A599FBDAFA34" box="[1040,1091,1493,1519]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, fig. 4, 3).
|
||
<emphasis id="B93DEAFFFFF35975FAE3A599FA36FA34" box="[1296,1455,1493,1519]" italics="true" pageId="6" pageNumber="51">Lovaracarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF35975FCDEA5B9FAD7F9D4" author="BESCHIN C. & DE ANGELI A. & CHECCHI A. & ZARANTONELLO G." box="[813,1358,1525,1551]" pageId="6" pageNumber="51" refId="ref6493" refString="BESCHIN C., DE ANGELI A., CHECCHI A. & ZARANTONELLO G. 2016. - Crostacei Decapodi del " Tufo a Lophoranina " (Luteziano inferiore) della Valle del Chiampo (Vicenza-Italia Nordorientale). Museo di Archeologia e Scienze Naturali " G. Zannato. " Montecchio Maggiore, Vicenza, Italy, 92 p." type="book" year="2016">Beschin, De Angeli, Checchi & Zarantonello, 2016</bibRefCitation>
|
||
from the Eocene of northern
|
||
<collectingCountry id="F35E767DFFF35975FBF0A658FBADF9F5" box="[1027,1076,1556,1582]" name="Italy" pageId="6" pageNumber="51">Italy</collectingCountry>
|
||
, differs from
|
||
<emphasis id="B93DEAFFFFF35975FB32A658FAF9F9F5" box="[1217,1376,1556,1582]" italics="true" pageId="6" pageNumber="51">Maurocarpilius</emphasis>
|
||
n. gen. in having shorter anterolateral margins, convex posterolateral margins, strongly punctulate surface and a triangular front (see
|
||
<bibRefCitation id="EFD84B1CFFF35975FCDEA638FBD9F954" author="BESCHIN C. & DE ANGELI A. & CHECCHI A. & ZARANTONELLO G." box="[813,1088,1652,1679]" pageId="6" pageNumber="51" refId="ref6493" refString="BESCHIN C., DE ANGELI A., CHECCHI A. & ZARANTONELLO G. 2016. - Crostacei Decapodi del " Tufo a Lophoranina " (Luteziano inferiore) della Valle del Chiampo (Vicenza-Italia Nordorientale). Museo di Archeologia e Scienze Naturali " G. Zannato. " Montecchio Maggiore, Vicenza, Italy, 92 p." type="book" year="2016">
|
||
Beschin
|
||
<emphasis id="B93DEAFFFFF35975FC76A639FC2FF955" box="[901,950,1652,1678]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2016: 55-56
|
||
</bibRefCitation>
|
||
, fig. 47, pl. 9, fig. 4A-B;
|
||
<bibRefCitation id="EFD84B1CFFF35975FACEA638FC22F974" author="SCHWEITZER C. E. & FELDMANN R. M. & KARASAWA H." pageId="6" pageNumber="51" pagination="1 - 22" refId="ref8508" refString="SCHWEITZER C. E., FELDMANN R. M. & KARASAWA H. 2018. - Part R, Revised, Volume 1, Chapter 8 T 2: Systematic descriptions: Superfamily Carpilioidea. Treatise Online 112: 1 - 22." type="journal article" year="2018">
|
||
Schweitzer
|
||
<emphasis id="B93DEAFFFFF35975FCDEA6D9FCF9F975" box="[813,864,1684,1710]" italics="true" pageId="6" pageNumber="51">et al.</emphasis>
|
||
2018: 5
|
||
</bibRefCitation>
|
||
, fig. 4, 4a-b).
|
||
<emphasis id="B93DEAFFFFF35975FBA1A6D8FB42F975" box="[1106,1243,1684,1710]" italics="true" pageId="6" pageNumber="51">Oscacarpilius</emphasis>
|
||
<bibRefCitation id="EFD84B1CFFF35975FB11A6D8FCFCF915" author="ARTAL P. & VAN BAKEL B." pageId="6" pageNumber="51" pagination="20 - 36" refId="ref6151" refString="ARTAL P. & VAN BAKEL B. 2018. - Carpiliids (Crustacea, Decapoda) from the Ypresian (Eocene) of the Northeast of Spain. Scripta Musei Geologici Seminarii Barcinonensis [Ser. palaeontologica] 12: 20 - 36." type="journal article" year="2018">Artal & Van Bakel, 2018</bibRefCitation>
|
||
from the northern of Iberian Peninsula, differs from the new genus in having a completely entire anterolateral margin, rounded lateral corners and a densely pitted surface (
|
||
<bibRefCitation id="EFD84B1CFFF35975FAAEA6B8FBB3F8F5" author="ARTAL P. & VAN BAKEL B." pageId="6" pageNumber="51" pagination="20 - 36" refId="ref6151" refString="ARTAL P. & VAN BAKEL B. 2018. - Carpiliids (Crustacea, Decapoda) from the Ypresian (Eocene) of the Northeast of Spain. Scripta Musei Geologici Seminarii Barcinonensis [Ser. palaeontologica] 12: 20 - 36." type="journal article" year="2018">Artal & Van Bakel 2018: 23-29</bibRefCitation>
|
||
, figs 1-2).
|
||
</paragraph>
|
||
<caption id="DF366665FFF35975FF77A570FA33FAA0" ID-DOI="http://doi.org/10.5281/zenodo.3696954" ID-Zenodo-Dep="3696954" httpUri="https://zenodo.org/record/3696954/files/figure.png" pageId="6" pageNumber="51" startId="6.[132,143,1340,1358]" targetBox="[132,1455,215,1295]" targetPageId="6">
|
||
<paragraph id="8BF636EDFFF35975FF77A570FA33FAA0" blockId="6.[132,1455,1339,1403]" pageId="6" pageNumber="51">
|
||
FIG. 3. —
|
||
<taxonomicName id="4C494D6EFFF35975FF22A570FE38FA95" authority="Ossó & Gagnaison & Bailleul, 2020" authorityName="Ossó & Gagnaison & Bailleul" authorityYear="2020" box="[209,417,1340,1358]" class="Malacostraca" family="Carpiliidae" genus="Maurocarpilius" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="binodosus" status="n. gen.,n.sp.">
|
||
<emphasis id="B93DEAFFFFF35975FF22A570FE38FA95" box="[209,417,1340,1358]" italics="true" pageId="6" pageNumber="51">Maurocarpilius binodosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20E5784FFF35975FE56A570FD8DFA95" box="[421,532,1340,1358]" pageId="6" pageNumber="51" rank="species">n. gen.,n.sp.</taxonomicNameLabel>
|
||
,from the Ypresian (early Eocene) of Kelâa M’Gouna (Morocco):
|
||
<emphasis id="B93DEAFFFFF35975FBFCA570FBA8FA95" bold="true" box="[1039,1073,1340,1358]" pageId="6" pageNumber="51">A -D</emphasis>
|
||
, holotype ULB-IV-A(1a);
|
||
<emphasis id="B93DEAFFFFF35975FB0BA570FA9CFA95" bold="true" box="[1272,1285,1340,1358]" pageId="6" pageNumber="51">A</emphasis>
|
||
, dorsal view (digitally cut out of the matrix);
|
||
<emphasis id="B93DEAFFFFF35975FECBA51FFEDCFABE" bold="true" box="[312,325,1363,1381]" pageId="6" pageNumber="51">B</emphasis>
|
||
, frontal view;
|
||
<emphasis id="B93DEAFFFFF35975FE44A51EFE5CFABF" bold="true" box="[439,453,1362,1380]" pageId="6" pageNumber="51">C</emphasis>
|
||
, right lateral view;
|
||
<emphasis id="B93DEAFFFFF35975FDADA51FFDF5FABE" bold="true" box="[606,620,1363,1381]" pageId="6" pageNumber="51">D</emphasis>
|
||
, posterior view;
|
||
<emphasis id="B93DEAFFFFF35975FD00A51FFC8CFABE" bold="true" box="[755,789,1362,1381]" pageId="6" pageNumber="51">E -H</emphasis>
|
||
, paratype ULB-IV-A(1b);
|
||
<emphasis id="B93DEAFFFFF35975FC10A51FFC76FABE" bold="true" box="[995,1007,1363,1381]" pageId="6" pageNumber="51">E</emphasis>
|
||
, dorsal view;
|
||
<emphasis id="B93DEAFFFFF35975FB93A51FFBF2FABE" bold="true" box="[1120,1131,1363,1381]" pageId="6" pageNumber="51">F</emphasis>
|
||
, frontal view;
|
||
<emphasis id="B93DEAFFFFF35975FB2EA51EFB72FABF" bold="true" box="[1245,1259,1362,1380]" pageId="6" pageNumber="51">G</emphasis>
|
||
, posterior view;
|
||
<emphasis id="B93DEAFFFFF35975FA81A51FFA19FABE" bold="true" box="[1394,1408,1363,1381]" pageId="6" pageNumber="51">H</emphasis>
|
||
, right lateral view;
|
||
<emphasis id="B93DEAFFFFF35975FF1FA525FF6BFAA0" bold="true" box="[236,242,1385,1403]" pageId="6" pageNumber="51">I</emphasis>
|
||
,
|
||
<emphasis id="B93DEAFFFFF35975FF0FA525FE9FFAA0" bold="true" box="[252,262,1385,1403]" pageId="6" pageNumber="51">J</emphasis>
|
||
, paratype ULB-IV-A(1c);
|
||
<emphasis id="B93DEAFFFFF35975FE25A525FE45FAA0" bold="true" box="[470,476,1385,1403]" pageId="6" pageNumber="51">I</emphasis>
|
||
, dorsal view;
|
||
<emphasis id="B93DEAFFFFF35975FDBCA525FDC0FAA0" bold="true" box="[591,601,1385,1403]" pageId="6" pageNumber="51">J</emphasis>
|
||
, frontal view.
|
||
<emphasis id="B93DEAFFFFF35975FD3FA525FC95FAA0" bold="true" box="[716,780,1385,1403]" pageId="6" pageNumber="51">Arrows</emphasis>
|
||
indicate the two anterolateral teeth distinctive of the species. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF636EDFFF35975FCB0A778FBE6F836" blockId="6.[813,1456,1461,2030]" pageId="6" pageNumber="51">
|
||
As mentioned above, the Moroccan specimens match perfectly with the general diagnosis of
|
||
<taxonomicName id="4C494D6EFFF35975FB5BA71FFABAF8B6" authorityName="Ortmann" authorityYear="1893" box="[1192,1315,1875,1901]" class="Malacostraca" family="Carpiliidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="51" phylum="Arthropoda" rank="family">Carpiliidae</taxonomicName>
|
||
. However, as can be seen in the comparisons, they do not fit completely with the characters or ratios of any of the aforementioned genera. Therefore, we propose the new genus,
|
||
<emphasis id="B93DEAFFFFF35975FAE3A7FFFA36F816" box="[1296,1455,1971,1997]" italics="true" pageId="6" pageNumber="51">Maurocarpilius</emphasis>
|
||
n. gen., to accommodate them.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |