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<document id="97CCD444FA2C8D86D1654BBBEAD11205" ID-CLB-Dataset="26904" ID-DOI="10.5852/ejt.2019.523" ID-GBIF-Dataset="7cefc0b8-0126-4022-a865-d2a8c0323c7c" ID-Zenodo-Dep="2673211" ID-ZooBank="0108D1A3-49B5-4EEE-8152-7714CDD17F3E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557293973202" checkinUser="plazi" docAuthor="Enghoff, Henrik &amp; Reboleira, Ana Sofia P. S." docDate="2019" docId="03E087EEFF9E2C7DFDAAFBBB712DFC8D" docLanguage="en" docName="ejt-523_enghoff_reboleira.pdf.imf" docOrigin="European Journal of Taxonomy 523" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Jeekelosoma abadi Enghoff &amp; Reboleira 2019, comb. nov." docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFD9FF96FF9C2C78FFABFFE9733AFFF0" masterDocTitle="The genus Jeekelosoma Mauriès, 1985 - Moroccan cave millipedes (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="3" updateTime="1698726463198" updateUser="ExternalLinkService">
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<mods:title id="6EDBDA8CCF3DDACBEEED0810DC635B29">The genus Jeekelosoma Mauriès, 1985 - Moroccan cave millipedes (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="05756D30D672F8888C63A890B4783E7E">Enghoff, Henrik</mods:namePart>
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<mods:namePart id="29D2F6DD00585101925704E0F0D8C067">Reboleira, Ana Sofia P. S.</mods:namePart>
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<subSubSection id="C3536573FF9E2C7AFDAAFBBB7709FB9D" box="[513,1075,1106,1134]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BF636F8FF9E2C7AFDAAFBBB7709FB9D" blockId="2.[513,1075,1106,1168]" box="[513,1075,1106,1134]" pageId="2" pageNumber="3">
<taxonomicName id="4C494D7BFF9E2C7AFDAAFBBB7090FB9E" authority="(Mauries, 1985)" authorityName="Enghoff &amp; Reboleira" authorityYear="2019" baseAuthorityName="Mauries" baseAuthorityYear="1985" box="[513,938,1106,1134]" class="Diplopoda" family="Paradoxosomatidae" genus="Jeekelosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="abadi" status="comb. nov.">
<emphasis id="B93DEAEAFF9E2C7AFDAAFBBB71DCFB9D" bold="true" box="[513,742,1106,1133]" italics="true" pageId="2" pageNumber="3">Jeekelosoma abadi</emphasis>
(
<bibRefCitation id="EFD84B09FF9E2C7AFD5EFBBA7098FB9E" author="Mauries J. - P." box="[757,930,1107,1134]" pageId="2" pageNumber="3" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès, 1985</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20E5791FF9E2C7AFC1AFBBA7709FB9D" box="[945,1075,1107,1133]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C3536573FF9E2C7AFD4CFB9F7076FB60" box="[743,844,1142,1168]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BF636F8FF9E2C7AFD4CFB9F7076FB60" blockId="2.[513,1075,1106,1168]" box="[743,844,1142,1168]" pageId="2" pageNumber="3">
<figureCitation id="13722A7DFF9E2C7AFD4CFB9F7076FB60" box="[743,844,1142,1168]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[189,232,1962,1988]" captionStartId-1="4.[189,232,1843,1869]" captionStartId-2="5.[189,232,1929,1955]" captionStartId-3="6.[189,232,1825,1851]" captionTargetBox-0="[189,1398,1287,1921]" captionTargetBox-1="[190,1399,378,1800]" captionTargetBox-2="[189,1398,961,1885]" captionTargetBox-3="[223,1365,265,1779]" captionTargetId-0="figure@2.[189,1398,1284,1924]" captionTargetId-1="figure@4.[190,1399,378,1800]" captionTargetId-2="figure@5.[189,1398,960,1885]" captionTargetId-3="figure@6.[223,1365,265,1779]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 1. Jeekelosoma abadi (Mauriès, 1985) comb. nov., living specimen from the type locality. Courtesy of Carles Hernando." captionText-1="Fig. 2. Jeekelosoma abadi (Mauriès, 1985) comb. nov., topotypes. AC. Female (NHMD 302022). DG. Male (NHMD 302021). A. Head and body rings 15, lateral view. B. Head and collum, frontal view. C. Body rings 810 and posterior half of ring 7, seen from the right. D. Rings 1011, seen from the left. E. Ring 11 and legs, posterior view. F. Rings 1011, dorsal view. G. Ozopore from ring 18, right side (anterior to the right). Scale bars: AF = 0.2 mm; G = 0.02 mm." captionText-2="Fig. 3. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021). A. Penultimate ring (podous), ultimate ring (apodous) and telson, oblique lateral view. B. Telson, posterior view. C. Midbody leg. D. Claw of midbody leg. Scale bars: AC = 0.1 mm; D = 0.01 mm." captionText-3="Fig. 4. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021), left gonopod in situ. A. Ventral view. B. Slightly meso-ventral view to show the keel a. C. Subapical view. D. Lateral view. E. Mesal view. Abbreviations: a = posterior-median keel of gonopod aperture; app = terminal appendage of solenophore; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore; spha = apical lamella of solenophore. Scale bars: AB, DE = 0.05 mm; C = 0.01 mm." httpUri-0="https://zenodo.org/record/2673213/files/figure.png" httpUri-1="https://zenodo.org/record/2673215/files/figure.png" httpUri-2="https://zenodo.org/record/2673217/files/figure.png" httpUri-3="https://zenodo.org/record/2673219/files/figure.png" pageId="2" pageNumber="3">Figs 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3536573FF9E2C7AFF16FB5E70A6FB21" box="[189,924,1207,1234]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BF636F8FF9E2C7AFF16FB5E70A6FB21" blockId="2.[189,924,1207,1234]" box="[189,924,1207,1234]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB5911D6FF9E2C7AFF16FB5E70A6FB21" box="[189,924,1207,1234]" pageId="2" pageNumber="3">
<taxonomicName id="4C494D7BFF9E2C7AFF16FB5E7026FB22" ID-CoL="3DHSC" authority="Mauries, 1985: 52" authorityName="Mauries" authorityPageNumber="52" authorityYear="1985" box="[189,796,1207,1234]" class="Diplopoda" family="Paradoxosomatidae" genus="Eviulisoma" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="abadi" subGenus="Jeekelosoma">
<emphasis id="B93DEAEAFF9E2C7AFF16FB5E7106FB21" box="[189,572,1207,1233]" italics="true" pageId="2" pageNumber="3">Eviulisoma (Jeekelosoma) abadi</emphasis>
<treatmentCitation id="0AE810E9FF9E2C7AFDE8FB5E7026FB22" author="Mauries J. - P." box="[579,796,1207,1234]" page="52" pageId="2" pageNumber="3" year="1985">
<bibRefCitation id="EFD84B09FF9E2C7AFDE8FB5E71CBFB22" author="Mauries J. - P." box="[579,753,1207,1234]" pageId="2" pageNumber="3" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès, 1985</bibRefCitation>
: 52
</treatmentCitation>
</taxonomicName>
, figs 113.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF366670FF9E2C7AFF16F8437291F817" httpUri="https://zenodo.org/record/2673213/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,232,1962,1988]" targetBox="[189,1398,1287,1921]" targetPageId="2">
<paragraph id="8BF636F8FF9E2C7AFF16F8437291F817" blockId="2.[189,1399,1962,2023]" pageId="2" pageNumber="3">
<emphasis id="B93DEAEAFF9E2C7AFF16F8437230F834" bold="true" box="[189,266,1962,1988]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
<taxonomicName id="4C494D7BFF9E2C7AFEBBF843718BF835" authority="(Mauries, 1985)" authorityName="Enghoff &amp; Reboleira" authorityYear="2019" baseAuthorityName="Mauries" baseAuthorityYear="1985" box="[272,689,1962,1989]" class="Diplopoda" family="Paradoxosomatidae" genus="Jeekelosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="abadi" status="comb. nov.">
<emphasis id="B93DEAEAFF9E2C7AFEBBF84372D5F834" box="[272,495,1962,1988]" italics="true" pageId="2" pageNumber="3">Jeekelosoma abadi</emphasis>
(Mauriès, 1985)
</taxonomicName>
<taxonomicNameLabel id="A20E5791FF9E2C7AFD13F8437001F835" box="[696,827,1962,1989]" pageId="2" pageNumber="3" rank="species">comb. nov.</taxonomicNameLabel>
, living specimen from the type locality. Courtesy of Carles Hernando.
</paragraph>
</caption>
<subSubSection id="C3536573FF9F2C7BFF16FEE2706DFE6A" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BF636F8FF9F2C7BFF16FEE2720CFED5" blockId="3.[189,310,267,293]" box="[189,310,267,293]" pageId="3" pageNumber="4">
<heading id="D0BE8194FF9F2C7BFF16FEE2720CFED5" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B93DEAEAFF9F2C7BFF16FEE2720CFED5" bold="true" box="[189,310,267,293]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FED3706DFE6A" blockId="3.[189,1399,313,410]" pageId="3" pageNumber="4">
A species of
<taxonomicName id="4C494D7BFF9F2C7BFEFFFED072D0FEA3" authorityName="Mauries" authorityYear="1985" box="[340,490,313,339]" class="Diplopoda" family="Paradoxosomatidae" genus="Jeekelosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEAEAFF9F2C7BFEFFFED072D0FEA3" box="[340,490,313,339]" italics="true" pageId="3" pageNumber="4">Jeekelosoma</emphasis>
</taxonomicName>
with 17 podous rings, with the ozopores borne on mushroomlike peritremata each provided with one macroseta, with a horizontal row of tubercles on podous rings 5-17, and with a terminal curved appendage on the gonopod solenophore.
</paragraph>
</subSubSection>
<subSubSection id="C3536573FF9F2C7BFF16FE287037FD84" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BF636F8FF9F2C7BFF16FE287290FE2B" blockId="3.[189,426,449,475]" box="[189,426,449,475]" pageId="3" pageNumber="4">
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<emphasis id="B93DEAEAFF9F2C7BFF16FE287290FE2B" bold="true" box="[189,426,449,475]" pageId="3" pageNumber="4">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FE197037FD84" blockId="3.[189,1399,496,628]" pageId="3" pageNumber="4">
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,
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,
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;
<location id="8E966023FF9F2C7BFCEDFE1970F7FDFA" LSID="urn:lsid:plazi:treatment:03E087EEFF9E2C7DFDAAFBBB712DFC8D:8E966023FF9F2C7BFCEDFE1970F7FDFA" box="[838,973,496,522]" country="Morocco" county="cave Kef Aziza" municipality="Boudenib" name="Tazouguert" pageId="3" pageNumber="4">Tazouguert</location>
,
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,
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;
<collectingDate id="EFB3E9D0FF9F2C7BFAB7FE1973C1FDDD" pageId="3" pageNumber="4" value="2001-05-31">
<date id="FFF71038FF9F2C7BFAB7FE1973C1FDDD" pageId="3" pageNumber="4" value="2001-05-31">31 May 2001</date>
</collectingDate>
;
<collectorName id="26BC532EFF9F2C7BFEA0FDFA72BAFDDD" box="[267,384,531,557]" pageId="3" pageNumber="4">O. Escola</collectorName>
leg.;
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<collectionCode id="ED58AE3DFF9F2C7BFE62FDFD7133FDDE" box="[457,521,532,558]" collectionName="MZB" pageId="3" pageNumber="4">MZB</collectionCode>
2003-1386
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,
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2019-0 0 14
</specimenCode>
,
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<collectionCode id="ED58AE3DFF9F2C7BFC2BFDFD70FAFDDE" box="[896,960,532,558]" collectionName="MZB" pageId="3" pageNumber="4">MZB</collectionCode>
2019-0 0 15
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,
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2019-0 0 16
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,
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2019-0 0 17
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2019-0 0 18
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,
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2019-0 0 19
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<materialsCitation id="3B213CA5FF9F2C7BFCA3FDDF7033FD84" ID-GBIF-Occurrence="2239085371" collectingDate="2001-05-31" collectionCode="NHMD" collectorName="O. Escola" country="Morocco" county="cave Kef Aziza" location="Tazouguert" municipality="Boudenib" pageId="3" pageNumber="4" specimenCode="NHMD 302019, NHMD 302020, NHMD 302021, NHMD 302022" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="9D4FFD71FF9F2C7BFCA3FDDF7074FDA1" box="[776,846,566,593]" pageId="3" pageNumber="4" type="male">2 ♂♂</specimenCount>
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; same collecting data;
<specimenCode id="DBEF9E83FF9F2C7BFB1EFDDE7648FDA0" box="[1205,1394,566,593]" collectionCode="NHMD" pageId="3" pageNumber="4">NHMD 302019</specimenCode>
,
<specimenCode id="DBEF9E83FF9F2C7BFF16FDB37243FD84" box="[189,377,602,628]" collectionCode="NHMD" pageId="3" pageNumber="4">NHMD 302020</specimenCode>
,
<specimenCode id="DBEF9E83FF9F2C7BFE2EFDB3717BFD84" box="[389,577,602,628]" collectionCode="NHMD" pageId="3" pageNumber="4">NHMD 302021</specimenCode>
,
<specimenCode id="DBEF9E83FF9F2C7BFDE7FDB37033FD84" box="[588,777,602,628]" collectionCode="NHMD" pageId="3" pageNumber="4">NHMD 302022</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536573FF9F2C7DFF16FD7271CEFDE7" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
<paragraph id="8BF636F8FF9F2C7BFF16FD7272A9FD45" blockId="3.[189,403,667,693]" box="[189,403,667,693]" pageId="3" pageNumber="4">
<heading id="D0BE8194FF9F2C7BFF16FD7272A9FD45" bold="true" box="[189,403,667,693]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B93DEAEAFF9F2C7BFF16FD7272A9FD45" bold="true" box="[189,403,667,693]" pageId="3" pageNumber="4">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FD2071A7FCF7" blockId="3.[189,1399,713,775]" pageId="3" pageNumber="4">
Based on studied specimens, to supplement original description (
<bibRefCitation id="EFD84B09FF9F2C7BFC42FD2077AFFD14" author="Mauries J. - P." box="[1001,1173,713,740]" pageId="3" pageNumber="4" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès 1985</bibRefCitation>
) and to introduce gonopod terminology of
<bibRefCitation id="EFD84B09FF9F2C7BFE4AFD0471A3FCF7" author="Enghoff H." box="[481,665,749,775]" pageId="3" pageNumber="4" pagination="1 - 90" refId="ref4451" refString="Enghoff H. 2018. A mountain of millipedes VII. The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae). European Journal of Taxonomy 445: 1 - 90. https: // doi. org / 10.5852 / ejt. 2018.445" type="journal article" year="2018">Enghoff (2018)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF41FCC7722FFC7F" blockId="3.[189,1399,814,911]" pageId="3" pageNumber="4">
<emphasis id="B93DEAEAFF9F2C7BFF41FCC77210FCB8" bold="true" box="[234,298,814,840]" pageId="3" pageNumber="4">Male</emphasis>
SIZE. Body length
<quantity id="4CB19B1DFF9F2C7BFE3CFCB8712DFC9C" box="[407,535,849,876]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="4" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
, max. width
<quantity id="4CB19B1DFF9F2C7BFD06FCB87023FC9C" box="[685,793,849,876]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="4" unit="mm" value="0.67">0.67 mm</quantity>
, 19 segments (17 podous rings + 1 apodous ring + telson).
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FC5F764AFAFD" blockId="3.[189,1399,948,1293]" pageId="3" pageNumber="4">
MIDBODY RINGS (
<figureCitation id="13722A7DFF9F2C7BFE27FC5F7136FC21" box="[396,524,950,977]" captionStart="Fig" captionStartId="4.[189,232,1843,1869]" captionTargetBox="[190,1399,378,1800]" captionTargetId="figure@4.[190,1399,378,1800]" captionTargetPageId="4" captionText="Fig. 2. Jeekelosoma abadi (Mauriès, 1985) comb. nov., topotypes. AC. Female (NHMD 302022). DG. Male (NHMD 302021). A. Head and body rings 15, lateral view. B. Head and collum, frontal view. C. Body rings 810 and posterior half of ring 7, seen from the right. D. Rings 1011, seen from the left. E. Ring 11 and legs, posterior view. F. Rings 1011, dorsal view. G. Ozopore from ring 18, right side (anterior to the right). Scale bars: AF = 0.2 mm; G = 0.02 mm." httpUri="https://zenodo.org/record/2673215/files/figure.png" pageId="3" pageNumber="4">Fig. 2D, F</figureCitation>
). Elongated, length of metazona equaling ca ¾ of maximum width. Body extremely narrow in anterior part, maximum width attained a little behind midbody. From there until and including ring 18 (the apodous ring in front of telson) body parallel-sided. Surface of metazona with very pronounced microcellulate structure, borders between cuticular scutes raised. Metazona with diameter increasing towards rear margin and with a clear constriction ca at the middle; posterior margin of rings finely crenulate but without a proper limbus. Each metazona with three rows of short setae borne on distinct tubercles; each row on midbody rings with ca 6 setae; intermediate row situated just behind constriction. A longitudinal row of tubercles on each side of metazona, consisting of a series of ca 5 tubercles in front of constriction plus, in ozopore-less rings, an additional series of ca 5 tubercles behind constriction. Posteriormost tubercles in each row with setae similar to those in transverse rows.
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FADD70EDFA65" blockId="3.[189,1399,1332,1429]" pageId="3" pageNumber="4">
OZOPORES (
<figureCitation id="13722A7DFF9F2C7BFEE3FADD7292FABE" box="[328,424,1332,1359]" captionStart="Fig" captionStartId="4.[189,232,1843,1869]" captionTargetBox="[190,1399,378,1800]" captionTargetId="figure@4.[190,1399,378,1800]" captionTargetPageId="4" captionText="Fig. 2. Jeekelosoma abadi (Mauriès, 1985) comb. nov., topotypes. AC. Female (NHMD 302022). DG. Male (NHMD 302021). A. Head and body rings 15, lateral view. B. Head and collum, frontal view. C. Body rings 810 and posterior half of ring 7, seen from the right. D. Rings 1011, seen from the left. E. Ring 11 and legs, posterior view. F. Rings 1011, dorsal view. G. Ozopore from ring 18, right side (anterior to the right). Scale bars: AF = 0.2 mm; G = 0.02 mm." httpUri="https://zenodo.org/record/2673215/files/figure.png" pageId="3" pageNumber="4">Fig. 2G</figureCitation>
). On peculiar mushroomlike outgrowths (modified peritremata); each mushroom with one long seta behind pore and several minute setae scattered over the surface; outline of mushrooms almost circular on anterior part of body, more oval towards the rear.
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FA557702FA26" blockId="3.[189,1080,1468,1495]" box="[189,1080,1468,1495]" pageId="3" pageNumber="4">STERNA. Unmodified, no process on sternum 5, no excavation on sternum 6.</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16FA1772FBF9CB" blockId="3.[189,1398,1533,1595]" pageId="3" pageNumber="4">
LEGS (
<figureCitation id="13722A7DFF9F2C7BFEA1FA177251F9E8" box="[266,363,1533,1560]" captionStart="Fig" captionStartId="4.[189,232,1843,1869]" captionTargetBox="[190,1399,378,1800]" captionTargetId="figure@4.[190,1399,378,1800]" captionTargetPageId="4" captionText="Fig. 2. Jeekelosoma abadi (Mauriès, 1985) comb. nov., topotypes. AC. Female (NHMD 302022). DG. Male (NHMD 302021). A. Head and body rings 15, lateral view. B. Head and collum, frontal view. C. Body rings 810 and posterior half of ring 7, seen from the right. D. Rings 1011, seen from the left. E. Ring 11 and legs, posterior view. F. Rings 1011, dorsal view. G. Ozopore from ring 18, right side (anterior to the right). Scale bars: AF = 0.2 mm; G = 0.02 mm." httpUri="https://zenodo.org/record/2673215/files/figure.png" pageId="3" pageNumber="4">Figs 2E</figureCitation>
;
<figureCitation id="13722A7DFF9F2C7BFED1FA1772FEF9E8" box="[378,452,1534,1560]" captionStart="Fig" captionStartId="5.[189,232,1929,1955]" captionTargetBox="[189,1398,961,1885]" captionTargetId="figure@5.[189,1398,960,1885]" captionTargetPageId="5" captionText="Fig. 3. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021). A. Penultimate ring (podous), ultimate ring (apodous) and telson, oblique lateral view. B. Telson, posterior view. C. Midbody leg. D. Claw of midbody leg. Scale bars: AC = 0.1 mm; D = 0.01 mm." httpUri="https://zenodo.org/record/2673217/files/figure.png" pageId="3" pageNumber="4">3CD</figureCitation>
). Slender, but femora considerably shorter than body diameter; femora distally swollen. No scopulae.
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16F98B7290F8F9" blockId="3.[189,1398,1634,1801]" pageId="3" pageNumber="4">
TELSON (
<figureCitation id="13722A7DFF9F2C7BFE8DF98B729FF98D" box="[294,421,1634,1661]" captionStart="Fig" captionStartId="5.[189,232,1929,1955]" captionTargetBox="[189,1398,961,1885]" captionTargetId="figure@5.[189,1398,960,1885]" captionTargetPageId="5" captionText="Fig. 3. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021). A. Penultimate ring (podous), ultimate ring (apodous) and telson, oblique lateral view. B. Telson, posterior view. C. Midbody leg. D. Claw of midbody leg. Scale bars: AC = 0.1 mm; D = 0.01 mm." httpUri="https://zenodo.org/record/2673217/files/figure.png" pageId="3" pageNumber="4">Fig. 3AB</figureCitation>
). Preanal ring triangular in dorsal view, lateral tubercles indicated by steps shortly before the subrectangular tip (epiproct); two transverse rows of setae plus several marginal setae. Anal valves (paraprocts) with raised mesal margin (lips), each with two setae, the ventral one situated slightly closer to the mesal margin than the dorsal one. Subanal scale (hypoproct) trapezoid, with two long marginal setae.
</paragraph>
<paragraph id="8BF636F8FF9F2C7BFF16F8D9704BF89E" blockId="3.[189,1398,1840,1902]" pageId="3" pageNumber="4">
GONOPODAL APERTURE (
<figureCitation id="13722A7DFF9F2C7BFE7CF8D9714CF8BB" box="[471,630,1840,1867]" captionStart="Fig" captionStartId="6.[189,232,1825,1851]" captionTargetBox="[223,1365,265,1779]" captionTargetId="figure@6.[223,1365,265,1779]" captionTargetPageId="6" captionText="Fig. 4. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021), left gonopod in situ. A. Ventral view. B. Slightly meso-ventral view to show the keel a. C. Subapical view. D. Lateral view. E. Mesal view. Abbreviations: a = posterior-median keel of gonopod aperture; app = terminal appendage of solenophore; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore; spha = apical lamella of solenophore. Scale bars: AB, DE = 0.05 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/2673219/files/figure.png" pageId="3" pageNumber="4">Fig. 4AB, E</figureCitation>
).
<figureCitation id="13722A7DFF9F2C7BFD22F8D971D1F8BA" box="[649,747,1840,1866]" captionStart="Fig" captionStartId="11.[189,232,1040,1066]" captionTargetBox="[526,1054,265,991]" captionTargetId="figure@11.[482,1106,265,999]" captionTargetPageId="11" captionText="Fig. 8. Jeekelosoma heptarachne sp. nov., holotype, male (MZB 73-2849), left gonopod. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2673227/files/figure.png" pageId="3" pageNumber="4">Figure 8</figureCitation>
-shaped, almost complete divided in midline by vertical, compressed lobe arising from posterior margin of aperture.
</paragraph>
<paragraph id="8BF636F8FF9F2C7DFF16F87C7286FE43" blockId="3.[189,1398,1941,2003]" lastBlockId="5.[189,1399,267,435]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
GONOPODS (
<figureCitation id="13722A7DFF9F2C7BFEF8F87C729BF85F" box="[339,417,1941,1968]" captionStart="Fig" captionStartId="6.[189,232,1825,1851]" captionTargetBox="[223,1365,265,1779]" captionTargetId="figure@6.[223,1365,265,1779]" captionTargetPageId="6" captionText="Fig. 4. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021), left gonopod in situ. A. Ventral view. B. Slightly meso-ventral view to show the keel a. C. Subapical view. D. Lateral view. E. Mesal view. Abbreviations: a = posterior-median keel of gonopod aperture; app = terminal appendage of solenophore; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore; spha = apical lamella of solenophore. Scale bars: AB, DE = 0.05 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/2673219/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). Coxa (not visible in
<figureCitation id="13722A7DFF9F2C7BFD14F87C7031F85F" box="[703,779,1941,1967]" captionStart="Fig" captionStartId="6.[189,232,1825,1851]" captionTargetBox="[223,1365,265,1779]" captionTargetId="figure@6.[223,1365,265,1779]" captionTargetPageId="6" captionText="Fig. 4. Jeekelosoma abadi (Mauriès, 1985) comb. nov., male topotype (NHMD 302021), left gonopod in situ. A. Ventral view. B. Slightly meso-ventral view to show the keel a. C. Subapical view. D. Lateral view. E. Mesal view. Abbreviations: a = posterior-median keel of gonopod aperture; app = terminal appendage of solenophore; map = mesal acropodital process; prf = prefemoral part; slm = solenomere; sph = solenophore; spha = apical lamella of solenophore. Scale bars: AB, DE = 0.05 mm; C = 0.01 mm." httpUri="https://zenodo.org/record/2673219/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
, but see
<bibRefCitation id="EFD84B09FF9F2C7BFC2BF87C7711F840" author="Mauries J. - P." box="[896,1067,1941,1968]" pageId="3" pageNumber="4" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès 1985</bibRefCitation>
: fig. 13) simple, without a distomedial lobe); prefemoral part (
<emphasis id="B93DEAEAFF9F2C7BFDC8F85171BFF822" box="[611,645,1976,2002]" italics="true" pageId="3" pageNumber="4">prf</emphasis>
) less than half as long as acropodite; mesal acropodital process (
<emphasis id="B93DEAEAFF992C7DFF6CFEE573C0FED5" box="[199,250,268,293]" italics="true" pageId="5" pageNumber="6">map</emphasis>
, “T” of
<bibRefCitation id="EFD84B09FF992C7DFEC9FEE27132FED6" author="Mauries J. - P." box="[354,520,267,294]" pageId="5" pageNumber="6" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès 1985</bibRefCitation>
) a simple broad, dorso-ventrally flattened lobe; solenophore (
<emphasis id="B93DEAEAFF992C7DFB5BFEE27623FED5" box="[1264,1305,267,293]" italics="true" pageId="5" pageNumber="6">sph</emphasis>
, “S” of
<bibRefCitation id="EFD84B09FF992C7DFF16FEC6725EFEB9" author="Mauries J. - P." box="[189,356,302,329]" pageId="5" pageNumber="6" pagination="51 - 62" refId="ref5057" refString="Mauries J. - P. 1985. Polydesmide et Craspedosomides nouveaux de France et du Maroc (Myriapoda - Diplopoda). International Journal of Speleology 14: 51 - 62." type="journal article" year="1985">Mauriès 1985</bibRefCitation>
) as long as
<emphasis id="B93DEAEAFF992C7DFE57FEC67115FEB8" box="[508,559,303,328]" italics="true" pageId="5" pageNumber="6">map</emphasis>
, slender, apically dividing into a subrectangular lamella (
<emphasis id="B93DEAEAFF992C7DFB46FEC67614FEB9" box="[1261,1326,303,329]" italics="true" pageId="5" pageNumber="6">spha</emphasis>
) with distal corners drawn out into long processes, and a terminal slender appendage (
<emphasis id="B93DEAEAFF992C7DFBDFFEBA779AFE9C" box="[1140,1184,339,364]" italics="true" pageId="5" pageNumber="6">app</emphasis>
) making a U-turn resulting in its rugose tip pointed towards gonopod base; solenomere (
<emphasis id="B93DEAEAFF992C7DFB8BFE9C7773FE7F" box="[1056,1097,373,399]" italics="true" pageId="5" pageNumber="6">slm</emphasis>
) slender, simple, largely concealed within
<emphasis id="B93DEAEAFF992C7DFE27FE71728FFE42" box="[396,437,408,434]" italics="true" pageId="5" pageNumber="6">sph</emphasis>
.
</paragraph>
<caption id="DF366670FF982C7CFF16F8DA70DAF82A" httpUri="https://zenodo.org/record/2673215/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1843,1869]" targetBox="[190,1399,378,1800]" targetPageId="4">
<paragraph id="8BF636F8FF982C7CFF16F8DA70DAF82A" blockId="4.[189,1399,1843,2010]" pageId="4" pageNumber="5">
<emphasis id="B93DEAEAFF982C7CFF16F8DA7234F8BD" bold="true" box="[189,270,1843,1869]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="4C494D7BFF982C7CFEB1F8DA71F3F8BE" authority="(Mauries, 1985)" authorityName="Enghoff &amp; Reboleira" authorityYear="2019" baseAuthorityName="Mauries" baseAuthorityYear="1985" box="[282,713,1843,1870]" class="Diplopoda" family="Paradoxosomatidae" genus="Jeekelosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="abadi" status="comb. nov.">
<emphasis id="B93DEAEAFF982C7CFEB1F8DA72C4F8BD" box="[282,510,1843,1869]" italics="true" pageId="4" pageNumber="5">Jeekelosoma abadi</emphasis>
(Mauriès, 1985)
</taxonomicName>
<taxonomicNameLabel id="A20E5791FF982C7CFD7EF8DA7061F8BD" box="[725,859,1843,1869]" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
, topotypes.
<emphasis id="B93DEAEAFF982C7CFC5AF8DA7710F8BD" bold="true" box="[1009,1066,1843,1869]" pageId="4" pageNumber="5">AC</emphasis>
. Female (NHMD 302022).
<emphasis id="B93DEAEAFF982C7CFF16F8BF73C2F880" bold="true" box="[189,248,1878,1904]" pageId="4" pageNumber="5">DG</emphasis>
. Male (NHMD 302021).
<emphasis id="B93DEAEAFF982C7CFD87F8BF717BF880" bold="true" box="[556,577,1878,1904]" pageId="4" pageNumber="5">A</emphasis>
. Head and body rings 15, lateral view.
<emphasis id="B93DEAEAFF982C7CFB83F8BF7706F880" bold="true" box="[1064,1084,1878,1904]" pageId="4" pageNumber="5">B</emphasis>
. Head and collum, frontal view.
<emphasis id="B93DEAEAFF982C7CFEAFF8907223F863" bold="true" box="[260,281,1913,1939]" pageId="4" pageNumber="5">C</emphasis>
. Body rings 810 and posterior half of ring 7, seen from the right.
<emphasis id="B93DEAEAFF982C7CFB9CF8907776F863" bold="true" box="[1079,1100,1913,1939]" pageId="4" pageNumber="5">D</emphasis>
. Rings 1011, seen from the left.
<emphasis id="B93DEAEAFF982C7CFEB0F8747215F847" bold="true" box="[283,303,1949,1975]" pageId="4" pageNumber="5">E</emphasis>
. Ring 11 and legs, posterior view.
<emphasis id="B93DEAEAFF982C7CFD15F87471EAF847" bold="true" box="[702,720,1949,1975]" pageId="4" pageNumber="5">F</emphasis>
. Rings 1011, dorsal view.
<emphasis id="B93DEAEAFF982C7CFBBBF875771DF846" bold="true" box="[1040,1063,1948,1974]" pageId="4" pageNumber="5">G</emphasis>
. Ozopore from ring 18, right side (anterior to the right). Scale bars: AF = 0.2 mm; G = 0.02 mm.
</paragraph>
</caption>
<paragraph id="8BF636F8FF992C7DFF41FE33727FFE04" blockId="5.[189,756,474,535]" box="[234,325,474,500]" pageId="5" pageNumber="6">
<emphasis id="B93DEAEAFF992C7DFF41FE33727FFE04" bold="true" box="[234,325,474,500]" pageId="5" pageNumber="6">Female</emphasis>
</paragraph>
<paragraph id="8BF636F8FF992C7DFF16FE1471CEFDE7" blockId="5.[189,756,474,535]" box="[189,756,509,535]" pageId="5" pageNumber="6">
SIZE. Body length
<quantity id="4CB19B1DFF992C7DFE33FE1472D7FDE7" box="[408,493,509,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="mm" value="12.0">12 mm</quantity>
, max. width
<quantity id="4CB19B1DFF992C7DFD2FFE1471CAFDE7" box="[644,752,509,535]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="5" pageNumber="6" unit="mm" value="0.83">0.83 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536573FF992C7DFF16FDD7712DFC8D" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BF636F8FF992C7DFF16FDD7764CFD77" blockId="5.[189,494,574,600]" lastBlockId="5.[189,1399,620,893]" pageId="5" pageNumber="6">
<emphasis id="B93DEAEAFF992C7DFF16FDD772D4FDA8" bold="true" box="[189,494,574,601]" pageId="5" pageNumber="6">Distribution and habitat</emphasis>
Known only from the
<typeStatus id="54F2885AFF992C7DFE65FD84713BFD77" box="[462,513,621,647]" pageId="5" pageNumber="6">type</typeStatus>
locality, the Kef Aziza cave, situated at
<geoCoordinate id="EE7D503FFF992C7DFC5AFD8277B4FD77" box="[1009,1166,619,647]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="15" value="-3.7880554">03°4717″ W</geoCoordinate>
,
<geoCoordinate id="EE7D503FFF992C7DFB37FD82760FFD77" box="[1180,1333,619,647]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="15" value="32.029446">32°0146″ N</geoCoordinate>
, near
</paragraph>
<paragraph id="8BF636F8FF992C7DFF16FD78712DFC8D" blockId="5.[189,1399,620,893]" pageId="5" pageNumber="6">
Tazzouguert (http://www.openspeleo.org/openspeleo/caves-view-294.html). This cave is also home to other cave-adapted species including the aquatic isopod
<taxonomicName id="4C494D7BFF992C7DFCEFFD5D77FDFD3D" authority="Magniez, 1977" authorityName="Magniez" authorityYear="1977" box="[836,1223,691,717]" class="Malacostraca" family="Stenasellidae" genus="Magniezia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gardei">
<emphasis id="B93DEAEAFF992C7DFCEFFD5D7728FD3D" box="[836,1042,691,717]" italics="true" pageId="5" pageNumber="6">Magniezia gardei</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFBBCFD5A77FDFD3D" author="Magniez G." box="[1047,1223,691,717]" pageId="5" pageNumber="6" pagination="321 - 329" refId="ref4948" refString="Magniez G. 1977. Magniezia gardei n. sp. (Crustacea Isopoda Asellote): un Stenasellide des eaux souterraines du Maroc sud-oriental. International Journal of Speleology 9: 321 - 329." type="journal article" year="1977">Magniez, 1977</bibRefCitation>
</taxonomicName>
, the palpigrade
<taxonomicName id="4C494D7BFF992C7DFF16FD3F7022FD01" authority="Barranco &amp; Mayoral, 2008" authorityName="Barranco &amp; Mayoral" authorityYear="2008" box="[189,792,726,753]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maroccana">
<emphasis id="B93DEAEAFF992C7DFF16FD3F72EBFD00" box="[189,465,726,752]" italics="true" pageId="5" pageNumber="6">Eukoenenia maroccana</emphasis>
Barranco &amp; Mayoral, 2008
</taxonomicName>
, the spider
<taxonomicName id="4C494D7BFF992C7DFC36FD3F77D6FD01" authority="Ribera, 1993" authorityName="Ribera" authorityYear="1993" box="[925,1260,726,753]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B93DEAEAFF992C7DFC36FD3F7771FD00" box="[925,1099,726,752]" italics="true" pageId="5" pageNumber="6">Dysdera caeca</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFBF8FD3F77D6FD01" author="Ribera C." box="[1107,1260,726,753]" pageId="5" pageNumber="6" pagination="1 - 7" refId="ref5207" refString="Ribera C. 1993. Dysdera caeca n. sp. y Harpactea stalitoides n. sp. (Araneae), dos especies cavernicolas de Marruecos y Portugal. Revue arachnologique 10 (1): 1 - 7." type="journal article" year="1993">Ribera, 1993</bibRefCitation>
</taxonomicName>
, the ground beetle
<taxonomicName id="4C494D7BFF992C7DFEA4FD13705EFCE4" authority="Lagar, 1978" authorityName="Lagar" authorityYear="1978" box="[271,868,762,788]" class="Insecta" family="Carabidae" genus="Platyderus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="presahariensis">
<emphasis id="B93DEAEAFF992C7DFEA4FD1371FFFCE4" box="[271,709,762,788]" italics="true" pageId="5" pageNumber="6">Platyderus insignitus presahariensis</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFD7AFD13705EFCE4" author="Lagar A." box="[721,868,762,788]" pageId="5" pageNumber="6" pagination="19 - 20" refId="ref4818" refString="Lagar A. 1978. Un nou Platyderus del Marroc (Coleoptera Pterostichidae). Excursionisme 41: 19 - 20." type="journal article" year="1978">Lagar, 1978</bibRefCitation>
</taxonomicName>
, the rove beetle
<taxonomicName id="4C494D7BFF992C7DFB93FD1373C0FCC7" authority="Espanol, 1969" authorityName="Espanol" authorityYear="1969" class="Insecta" family="Staphylinidae" genus="Apteranillus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ruei">
<emphasis id="B93DEAEAFF992C7DFB93FD13763EFCE3" box="[1080,1284,762,788]" italics="true" pageId="5" pageNumber="6">Apteranillus ruei</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFABAFD1373C0FCC7" author="Espanol F." pageId="5" pageNumber="6" pagination="171 - 176" refId="ref4652" refString="Espanol F. 1969. Un nuevo Apteranillus cavernicola de la region de Boudenib, Marruecos presahariano (Col. Staphylinidae). Annales de Speleologie 24 (1): 171 - 176." type="journal article" year="1969">Español, 1969</bibRefCitation>
</taxonomicName>
, the darkling beetle
<taxonomicName id="4C494D7BFF992C7DFE4EFCF470FAFCC8" authority="Antoine, 1953" authorityName="Antoine" authorityYear="1953" box="[485,960,797,824]" class="Insecta" family="Tenebrionidae" genus="Scaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="tingitanus" subSpecies="gimeli">
<emphasis id="B93DEAEAFF992C7DFE4EFCF47035FCC7" box="[485,783,797,823]" italics="true" pageId="5" pageNumber="6">Scaurus tingitanus gimeli</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFCBDFCF470FAFCC8" author="Antoine M." box="[790,960,797,824]" pageId="5" pageNumber="6" pagination="58 - 59" refId="ref4270" refString="Antoine M. 1953. Presentation des Coleopteres troglobies. Comptes rendus mensuelles des Seances de la Societe de Sciences Naturelles et Physiques du Maroc 19 (4): 58 - 59." type="journal article" year="1953">Antoine, 1953</bibRefCitation>
</taxonomicName>
, and the weevil
<taxonomicName id="4C494D7BFF992C7DFBD7FCF77266FCAA" authority="Stuben, 2009" authorityName="Stuben" authorityYear="2009" class="Insecta" family="Curculionidae" genus="Torneuma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="troglodytis">
<emphasis id="B93DEAEAFF992C7DFBD7FCF7764CFCC7" box="[1148,1398,797,823]" italics="true" pageId="5" pageNumber="6">Torneuma troglodytis</emphasis>
<bibRefCitation id="EFD84B09FF992C7DFF16FCA97266FCAA" author="Stuben P. E." box="[189,348,832,858]" pageId="5" pageNumber="6" pagination="94 - 106" refId="ref5248" refString="Stuben P. E. 2009. Neubeschreibungen westpalaarktischer Cryptorhynchinae III. (Torneuma, Echinodera, Acalles). Snudebiller 10: 94 - 106." type="journal article" year="2009">Stüben, 2009</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFD84B09FF992C7DFEC4FCA97123FCAB" author="Antoine M." box="[367,537,832,859]" pageId="5" pageNumber="6" pagination="58 - 59" refId="ref4270" refString="Antoine M. 1953. Presentation des Coleopteres troglobies. Comptes rendus mensuelles des Seances de la Societe de Sciences Naturelles et Physiques du Maroc 19 (4): 58 - 59." type="journal article" year="1953">Antoine 1953</bibRefCitation>
;
<bibRefCitation id="EFD84B09FF992C7DFD81FCA971E8FCAA" author="Espanol F." box="[554,722,832,858]" pageId="5" pageNumber="6" pagination="171 - 176" refId="ref4652" refString="Espanol F. 1969. Un nuevo Apteranillus cavernicola de la region de Boudenib, Marruecos presahariano (Col. Staphylinidae). Annales de Speleologie 24 (1): 171 - 176." type="journal article" year="1969">Español 1969</bibRefCitation>
;
<bibRefCitation id="EFD84B09FF992C7DFD49FCA970A8FCAA" author="Magniez G." box="[738,914,832,858]" pageId="5" pageNumber="6" pagination="321 - 329" refId="ref4948" refString="Magniez G. 1977. Magniezia gardei n. sp. (Crustacea Isopoda Asellote): un Stenasellide des eaux souterraines du Maroc sud-oriental. International Journal of Speleology 9: 321 - 329." type="journal article" year="1977">Magniez 1977</bibRefCitation>
;
<bibRefCitation id="EFD84B09FF992C7DFC09FCA8770AFCAA" author="Lagar A." box="[930,1072,832,859]" pageId="5" pageNumber="6" pagination="19 - 20" refId="ref4818" refString="Lagar A. 1978. Un nou Platyderus del Marroc (Coleoptera Pterostichidae). Excursionisme 41: 19 - 20." type="journal article" year="1978">Lagar 1978</bibRefCitation>
;
<bibRefCitation id="EFD84B09FF992C7DFBEAFCA977E2FCAA" author="Ribera C." box="[1089,1240,832,858]" pageId="5" pageNumber="6" pagination="1 - 7" refId="ref5207" refString="Ribera C. 1993. Dysdera caeca n. sp. y Harpactea stalitoides n. sp. (Araneae), dos especies cavernicolas de Marruecos y Portugal. Revue arachnologique 10 (1): 1 - 7." type="journal article" year="1993">Ribera 1993</bibRefCitation>
; Barranco &amp; Mayoral 2008;
<bibRefCitation id="EFD84B09FF992C7DFED9FC8A7131FC8D" author="Stuben P. E." box="[370,523,867,893]" pageId="5" pageNumber="6" pagination="94 - 106" refId="ref5248" refString="Stuben P. E. 2009. Neubeschreibungen westpalaarktischer Cryptorhynchinae III. (Torneuma, Echinodera, Acalles). Snudebiller 10: 94 - 106." type="journal article" year="2009">Stüben 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>