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<document id="469CFC234F74A7DB214B1282D03D865E" ID-CLB-Dataset="20261" ID-DOI="10.11646/zootaxa.5115.2.7" ID-GBIF-Dataset="e75580eb-9ee1-41e2-af0b-5028b90576f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="6352464" ID-ZooBank="A33E7563-28AC-43E0-8B6D-2DE7E571AB07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647259553979" checkinUser="plazi" docAuthor="Pett, Brogan L. &amp; Rabemananjara, Paul Bienvenu" docDate="2022" docId="03E087D6E011FFDBC6FC282CFCF0AC16" docLanguage="en" docName="zootaxa.5115.2.7.pdf" docOrigin="Zootaxa 5115 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copa sakalava Pett &amp; Rabemananjara 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="287" masterDocId="FFD9FFAEE013FFDDC66B2B36FFDAAD0B" masterDocTitle="A new species of Copa (Araneae: Corinnidae: Castianeirinae) from dry forests in the north west of Madagascar" masterLastPageNumber="287" masterPageNumber="281" pageNumber="283" updateTime="1699318008879" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0E5ADB37C72291E18E303B3AE5C46036" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6F3082FE6C355DADB2335B367F8C9EA">
<mods:title id="0C96BD978CE7A4D08029606F406A70F3">A new species of Copa (Araneae: Corinnidae: Castianeirinae) from dry forests in the north west of Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="479E38D726602733CA2C1A8B313F46B5" type="personal">
<mods:role id="77E71753A30D443B0A615CB2928C47B3">
<mods:roleTerm id="13D31759EBD1F26F308BA1862426649D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81D50976533B808960E5D9BB348304F8">Pett, Brogan L.</mods:namePart>
<mods:affiliation id="8974F80E5ABE410890F3B55701A00A6E">SpiDivERse, Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium. &amp; Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Penryn Campus, Penryn, &amp; Operation Wallacea, Lincolnshire, UK.</mods:affiliation>
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<mods:name id="F714AC56CE7D2632B88080D6252F5318" type="personal">
<mods:role id="33FFC0FDAA0BA762AE8232251169783D">
<mods:roleTerm id="826F643270A1BBD15BB206BE2046EE5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03A5C3C7A3B05241870A414B7B00BA7F">Rabemananjara, Paul Bienvenu</mods:namePart>
<mods:nameIdentifier id="AC47DFA7CACE703422822A942C27A841" type="ORCID">0000-0001-6395-917X</mods:nameIdentifier>
<mods:affiliation id="D18127E310590EA5A8B576B8029B4AD3">Development and Biodiversity Conservation Action for Madagascar (DBCAM), Antananarivo, Madagascar. &amp; Faculté des Sciences, de Technologies et de l’environnement (FSTE), Université de Majunga, Mahajanga, Madagascar. https: // orcid. org / 0000 - 0001 - 6395 - 917 X</mods:affiliation>
</mods:name>
<mods:typeOfResource id="07C1DCFACE67DD53CE4094DD88391B11">text</mods:typeOfResource>
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<mods:titleInfo id="3E951334B132C5430A1EC3A524C28283">
<mods:title id="762076F55EF701BE9BA25C373E2D4A91">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EC7CDC1163BB2E1E11B670A40D4F6EB3">
<mods:date id="BD9602E3AD0F1109B16DAB591B1259E2">2022</mods:date>
<mods:detail id="7B487BC9B30681D93E6F1E876E944638" type="pubDate">
<mods:number id="B91848DCA0081BF4547D34A6ADDE4892">2022-03-14</mods:number>
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<mods:detail id="709F23A56E7E9714FCCBBA4020D9CC7D" type="volume">
<mods:number id="5F8340BD9A25A2EC6E585E89FF6A9B60">5115</mods:number>
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<mods:detail id="DE590C6C2B25C7D8EA957D75DFE82143" type="issue">
<mods:number id="95E5E498D1279973B20DE34ECF67E0AA">2</mods:number>
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<mods:start id="CBAA393C684853320D93A5820974DD21">281</mods:start>
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<mods:classification id="4BFDCF1B3DF9137B7E96ADF7E6338E30">journal article</mods:classification>
<mods:identifier id="08CC0EB3C93F0F95346EB7B4A91D8903" type="CLB-Dataset">20261</mods:identifier>
<mods:identifier id="D53C7876E668C443EBA0E050C164D718" type="DOI">10.11646/zootaxa.5115.2.7</mods:identifier>
<mods:identifier id="0311203217FFC50F49C84C02C59262F1" type="GBIF-Dataset">e75580eb-9ee1-41e2-af0b-5028b90576f0</mods:identifier>
<mods:identifier id="479A31E58BC19988E2B13325EE09EDC4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="798565D463E0AEB053880C5BB3D83CDD" type="Zenodo-Dep">6352464</mods:identifier>
<mods:identifier id="7BE87A37AD3DA6385E2C78B91186DC47" type="ZooBank">A33E7563-28AC-43E0-8B6D-2DE7E571AB07</mods:identifier>
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<treatment id="03E087D6E011FFDBC6FC282CFCF0AC16" ID-DOI="http://doi.org/10.5281/zenodo.6352479" ID-GBIF-Taxon="193983936" ID-Zenodo-Dep="6352479" LSID="urn:lsid:plazi:treatment:03E087D6E011FFDBC6FC282CFCF0AC16" httpUri="http://treatment.plazi.org/id/03E087D6E011FFDBC6FC282CFCF0AC16" lastPageId="6" lastPageNumber="287" pageId="2" pageNumber="283">
<subSubSection id="C353654BE011FFDFC6FC282CFC9FAE52" pageId="2" pageNumber="283" type="nomenclature">
<paragraph id="8BF636C0E011FFDFC6FC282CFE34AE3F" blockId="2.[151,837,793,892]" box="[151,494,793,820]" pageId="2" pageNumber="283">
<heading id="D0BE81ACE011FFDFC6FC282CFE34AE3F" bold="true" box="[151,494,793,820]" fontSize="11" level="1" pageId="2" pageNumber="283" reason="1">
<emphasis id="B93DEAD2E011FFDFC6FC282CFE34AE3F" bold="true" box="[151,494,793,820]" pageId="2" pageNumber="283">
<taxonomicName id="4C494D43E011FFDFC6FC282CFE59AE3F" ID-CoL="98ZHC" authority="Pett" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[151,387,793,820]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E011FFDFC6FC282CFE92AE3F" bold="true" box="[151,328,793,820]" italics="true" pageId="2" pageNumber="283">Copa sakalava</emphasis>
Pett
</taxonomicName>
,
<taxonomicNameLabel id="A20E57A9E011FFDFC7FB282CFE34AE3F" box="[400,494,794,820]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636C0E011FFDFC6FC2808FC9FAE52" blockId="2.[151,837,793,892]" box="[151,837,830,857]" pageId="2" pageNumber="283">http://zoobank.org/81d387d6-dc60-481a-a22a-0c47100f7985</paragraph>
</subSubSection>
<subSubSection id="C353654BE011FFDFC6FC2854FEDDAE77" box="[151,263,866,892]" pageId="2" pageNumber="283" type="description">
<paragraph id="8BF636C0E011FFDFC6FC2854FEDDAE77" blockId="2.[151,837,793,892]" box="[151,263,866,892]" pageId="2" pageNumber="283">
<figureCitation id="13722A45E011FFDFC6FC2854FEDDAE77" box="[151,263,866,892]" captionStart-0="FIGURES 2–4" captionStart-1="FIGURES 5–7" captionStart-2="FIGURES 8–11" captionStartId-0="3.[151,264,1490,1514]" captionStartId-1="4.[151,264,1492,1516]" captionStartId-2="5.[151,264,1834,1858]" captionTargetBox-0="[161,1432,191,1460]" captionTargetBox-1="[159,1436,420,1457]" captionTargetBox-2="[164,1422,195,1800]" captionTargetId-0="figure-19@3.[151,1436,181,1466]" captionTargetId-1="figure-166@4.[151,1436,417,1469]" captionTargetId-2="figure-19@5.[151,1436,181,1810]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 2–4. Copa sakalava sp. nov. Male holotype habitus. 2 dorsal, 3 line drawing of dorsal habitus detail, 4 ventral." captionText-1="FIGURES 5–7. Copa sakalava sp. nov. Female paratype habitus. 5 dorsal, 6 detail of dorsal habitus, 7 ventral." captionText-2="FIGURES 8–11. Copa sakalava sp. nov. genitalia illustrations. 8-9 male pedipalp, 10-11 female epigyne. 8, 10 ventral, 9 retrolateral, 11 dorsal. CO = copulatory openings, CD = copulatory duct, Em = embolus, FD = fertilisation duct, H = epigynal hood, LC = Lateral chamber of copulatory duct, PcS = paracymbial spine, SD = sperm duct, St = subtegulum, ST I &amp; ST II = spermathecae I and II. Scale bar for 10 &amp; 11 = 0.25mm." figureDoi-0="http://doi.org/10.5281/zenodo.6352466" figureDoi-1="http://doi.org/10.5281/zenodo.6352468" figureDoi-2="http://doi.org/10.5281/zenodo.6352470" httpUri-0="https://zenodo.org/record/6352466/files/figure.png" httpUri-1="https://zenodo.org/record/6352468/files/figure.png" httpUri-2="https://zenodo.org/record/6352470/files/figure.png" pageId="2" pageNumber="283">Figs 2–10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C353654BE011FFDFC6FC289FFCFEA93A" pageId="2" pageNumber="283" type="materials_examined">
<paragraph id="8BF636C0E011FFDFC6FC289FFD3CAEE2" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6FC289FFE50AEC8" bold="true" box="[151,394,937,963]" pageId="2" pageNumber="283">Material Examined.</emphasis>
<materialsCitation id="3B213C9DE011FFDFC7FE289CFD3CAEE2" ID-GBIF-Occurrence="3704589303" collectingDate="2018-06-19" collectionCode="RMCA" collectorName="Brogan L. Pett" country="Madagascar" latitude="-15.504529" location="MADAGASCAR" longLatPrecision="1" longitude="46.708603" pageId="2" pageNumber="283" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F28862E011FFDFC7FE289CFE27AECF" box="[405,509,938,964]" pageId="2" pageNumber="283" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4FFD49E011FFDFC46C289CFDC6AECF" box="[519,540,938,964]" pageId="2" pageNumber="283" type="male">♂</specimenCount>
.
<collectingCountry id="F35E7650E011FFDFC445289CFD20AECF" box="[558,762,938,964]" name="Madagascar" pageId="2" pageNumber="283">MADAGASCAR</collectingCountry>
:
<geoCoordinate id="EE7D5007E011FFDFC56C289CFC51AECE" box="[775,907,938,965]" degrees="15.504529" direction="south" orientation="latitude" pageId="2" pageNumber="283" precision="1" value="-15.504529">-15.504529</geoCoordinate>
,
<geoCoordinate id="EE7D5007E011FFDFC5F2289CFBC9AECF" box="[921,1043,938,964]" degrees="46.708601" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="1" value="46.708603">46.708601</geoCoordinate>
, Mariarano,
<date id="FFF71000E011FFDFC2C3289CFA9DAECF" box="[1192,1351,938,964]" pageId="2" pageNumber="283" value="2018-06-19">
<collectingDate id="EFB3E9E8E011FFDFC2C3289CFA9DAECF" box="[1192,1351,938,964]" pageId="2" pageNumber="283" value="2018-06-19">19 June 2018</collectingDate>
</date>
, forest, 21:07h,
<collectorName id="26BC5316E011FFDFC69828F8FE42AEE3" box="[243,408,974,1000]" pageId="2" pageNumber="283">Brogan L. Pett</collectorName>
leg (
<collectionCode id="ED58AE05E011FFDFC7A428F8FDF8AEE3" box="[463,546,974,1000]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="283" type="Museum">RMCA</collectionCode>
_ARA_ 247355).
</materialsCitation>
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC28C4FCFEA93A" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<materialsCitation id="3B213C9DE011FFDFC6AC28C4FCC5A93A" ID-GBIF-Occurrence="3704589302" collectingDate="2018-06-17" collectionCode="RMCA" collectorName="Brogan L. Pett" country="Madagascar" latitude="-15.478513" location="MADAGASCAR" longLatPrecision="1" longitude="46.683487" pageId="2" pageNumber="283" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="54F28862E011FFDFC6AC28C4FEE2A907" box="[199,312,1010,1036]" pageId="2" pageNumber="283" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4FFD49E011FFDFC72F28C4FEAAA907" box="[324,368,1010,1036]" pageId="2" pageNumber="283" type="female">2 ♀</specimenCount>
,
<collectingCountry id="F35E7650E011FFDFC71428C4FD91A907" box="[383,587,1010,1036]" name="Madagascar" pageId="2" pageNumber="283">MADAGASCAR</collectingCountry>
:
<geoCoordinate id="EE7D5007E011FFDFC43C28C4FD00A906" box="[599,730,1010,1037]" degrees="15.478513" direction="south" orientation="latitude" pageId="2" pageNumber="283" precision="1" value="-15.478513">-15.478513</geoCoordinate>
,
<geoCoordinate id="EE7D5007E011FFDFC48D28C4FCBAA907" box="[742,864,1010,1036]" degrees="46.683486" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="1" value="46.683487">46.683486</geoCoordinate>
, Mariarano,
<date id="FFF71000E011FFDFC59A28C4FB56A907" box="[1009,1164,1010,1036]" pageId="2" pageNumber="283" value="2018-06-17">
<collectingDate id="EFB3E9E8E011FFDFC59A28C4FB56A907" box="[1009,1164,1010,1036]" pageId="2" pageNumber="283" value="2018-06-17">17 June 2018</collectingDate>
</date>
, forest patch near riverbank, 20:00h,
<collectorName id="26BC5316E011FFDFC75C2F20FE06A93B" box="[311,476,1046,1072]" pageId="2" pageNumber="283">Brogan L. Pett</collectorName>
leg (
<collectionCode id="ED58AE05E011FFDFC4782F20FDB2A93A" box="[531,616,1046,1073]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="283" type="Museum">RMCA</collectionCode>
_ARA_247356)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C353654BE011FFDFC6AC2F0CFDCFA973" pageId="2" pageNumber="283" type="etymology">
<paragraph id="8BF636C0E011FFDFC6AC2F0CFDCFA973" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2F0CFE94A95F" bold="true" box="[199,334,1082,1108]" pageId="2" pageNumber="283">Etymology.</emphasis>
The species epithet honours the Sakalava ethnic group of
<collectingCountry id="F35E7650E011FFDFC5B42F0CFBBCA95F" box="[991,1126,1082,1108]" name="Madagascar" pageId="2" pageNumber="283">Madagascar</collectingCountry>
from the region of the
<typeStatus id="54F28862E011FFDFC3072F0CFA46A95F" box="[1388,1436,1082,1108]" pageId="2" pageNumber="283">type</typeStatus>
locality. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C353654BE011FFDFC6AC2FB7FE1CAB0F" pageId="2" pageNumber="283" type="diagnosis">
<paragraph id="8BF636C0E011FFDFC6AC2FB7FCAAA87F" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2FB7FE98A990" bold="true" box="[199,322,1153,1179]" pageId="2" pageNumber="283">Diagnosis.</emphasis>
<taxonomicName id="4C494D43E011FFDFC7212FB4FE2EA997" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[330,500,1154,1180]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E011FFDFC7212FB4FE2EA997" box="[330,500,1154,1180]" italics="true" pageId="2" pageNumber="283">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E011FFDFC7962FB4FD83A997" bold="true" box="[509,601,1154,1180]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A20E57A9E011FFDFC7962FB4FD83A997" box="[509,601,1154,1180]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily distinguished from all other
<taxonomicName id="4C494D43E011FFDFC2702FB4FB82A997" authorityName="Simon" authorityYear="1885" box="[1051,1112,1154,1180]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEAD2E011FFDFC2702FB4FB82A997" box="[1051,1112,1154,1180]" italics="true" pageId="2" pageNumber="283">Copa</emphasis>
</taxonomicName>
by: broad embolus with 1.5 turns, with the final half turn directed prolaterally, extending around 1/3
<superScript id="7C3C9B88E011FFDFC5AD2F93FC0EA9B8" attach="left" box="[966,980,1189,1203]" fontSize="6" pageId="2" pageNumber="283">rd</superScript>
to ½ of the length back across the basal embolus coil (rather than long, thin, and extending to apex of cymbium in
<emphasis id="B93DEAD2E011FFDFC5862FFCFB74A9EF" box="[1005,1198,1226,1252]" italics="true" pageId="2" pageNumber="283">
<taxonomicName id="4C494D43E011FFDFC5862FFCFB70A9EF" authorityName="Simon" authorityYear="1885" box="[1005,1194,1226,1252]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="flavoplumosa">C. flavoplumosa</taxonomicName>
,
</emphasis>
or relatively straight and medium width, extending to cymbial apex in
<taxonomicName id="4C494D43E011FFDFC4A92FD8FC76A803" authority="Haddad, 2013" authorityName="Haddad" authorityYear="2013" box="[706,940,1262,1288]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="kei">
<emphasis id="B93DEAD2E011FFDFC4A92FD8FCD9A803" box="[706,771,1262,1288]" italics="true" pageId="2" pageNumber="283">C. kei</emphasis>
<bibRefCitation id="EFD84B31E011FFDFC5612FD8FC76A803" author="Haddad, C. R." box="[778,940,1262,1288]" pageId="2" pageNumber="283" pagination="1 - 37" refId="ref3082" refString="Haddad, C. R. (2013 a) A revision of the continental species of Copa Simon, 1885 (Araneae, Corinnidae) in the Afrotropical region. ZooKeys, 276, 1 - 37. https: // doi. org / 10.3897 / zookeys. 276.4233" type="journal article" year="2013">Haddad, 2013</bibRefCitation>
</taxonomicName>
). Additionally,
<taxonomicName id="4C494D43E011FFDFC23D2FD8FB27A803" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[1110,1277,1262,1288]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E011FFDFC23D2FD8FB27A803" box="[1110,1277,1262,1288]" italics="true" pageId="2" pageNumber="283">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E011FFDFC36F2FD8FA87A803" bold="true" box="[1284,1373,1262,1288]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A20E57A9E011FFDFC36F2FD8FA87A803" box="[1284,1373,1262,1288]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a sharp but stubby paracymbial spine retrolaterally. Females are distinguished primarily by the lateral chamber of the CD that joins the median path of CD mid-way between the CO and connection between ST and CD, in ventral view the large semi-circular epigynal hood is also unique in the genus.
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2E4BFE1CAB0F" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2E4BFE42A893" bold="true" box="[199,408,1405,1432]" pageId="2" pageNumber="283">Taxonomic notes.</emphasis>
<taxonomicName id="4C494D43E011FFDFC7CB2E48FD93A893" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[416,585,1406,1432]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E011FFDFC7CB2E48FD93A893" box="[416,585,1406,1432]" italics="true" pageId="2" pageNumber="283">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E011FFDFC43A2E48FD76A893" bold="true" box="[593,684,1406,1432]" pageId="2" pageNumber="283">
<taxonomicNameLabel id="A20E57A9E011FFDFC43A2E48FD76A893" box="[593,684,1406,1432]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only species in the genus currently known to have a mostly uniform orangish brown habitus and, at present, it may be utilised as a diagnostic character. However, with a revision with more than 40 new species from
<collectingCountry id="F35E7650E011FFDFC4032EF0FD35A8EB" box="[616,751,1478,1504]" name="Madagascar" pageId="2" pageNumber="283">Madagascar</collectingCountry>
pending, we consider the apparent uniqueness of this character may quickly lose value.
</paragraph>
</subSubSection>
<subSubSection id="C353654BE011FFDBC6AC2D3BFCF0AC16" lastPageId="6" lastPageNumber="287" pageId="2" pageNumber="283" type="description">
<paragraph id="8BF636C0E011FFDFC6AC2D3BFE8BAB2C" blockId="2.[151,1437,937,2008]" box="[199,337,1549,1575]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2D3BFE8BAB2C" bold="true" box="[199,337,1549,1575]" pageId="2" pageNumber="283">Description</emphasis>
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2D04FD49AB46" blockId="2.[151,1437,937,2008]" box="[199,659,1586,1613]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2D04FEDEAB47" bold="true" box="[199,260,1586,1612]" pageId="2" pageNumber="283">Male</emphasis>
(
<materialsCitation id="3B213C9DE011FFDFC7782D04FD49AB46" ID-GBIF-Occurrence="3704589301" box="[275,659,1586,1613]" collectionCode="RMCA" pageId="2" pageNumber="283" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F28862E011FFDFC7782D04FEA1AB47" box="[275,379,1586,1612]" pageId="2" pageNumber="283" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED58AE05E011FFDFC7EE2D04FE0DAB47" box="[389,471,1586,1612]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="2" pageNumber="283" type="Museum">RMCA</collectionCode>
_ARA_ 247355)
</materialsCitation>
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2D60FAD8ABB3" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
Measurements. TL 5.30, CL 2.68, CW 2.07,
<collectingCountry id="F35E7650E011FFDFC4A82D60FD30AB7B" box="[707,746,1622,1648]" name="Switzerland" pageId="2" pageNumber="283">CH</collectingCountry>
1.12, SL 1.34, SW 1.12, AL 2.62, AW 1.72, chelicera length 0.77, chelicera width 0.43. Legs. I: 1.76, 0.71, 1.42, 1.36, 1.16. II: 1.69, 0.48, 1.20, 1.38, 1.10. III: 1.80, 0.79, 1.24, 1.67, 0.98. IV: 2.50, 1.02, 1.90, 2.58, 1.17. Eyes: AME—0.10, ALE—0.08, PME—0.09, PLE—0.08.
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2DF4FB03AA9B" blockId="2.[151,1437,937,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2DF4FE83ABD7" box="[199,345,1730,1756]" italics="true" pageId="2" pageNumber="283">Colouration:</emphasis>
Carapace orange to brownish-orange (
<figureCitation id="13722A45E011FFDFC57C2DF4FC5CABD7" box="[791,902,1730,1756]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[161,1432,191,1460]" captionTargetId="figure-19@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURES 2–4. Copa sakalava sp. nov. Male holotype habitus. 2 dorsal, 3 line drawing of dorsal habitus detail, 4 ventral." figureDoi="http://doi.org/10.5281/zenodo.6352466" httpUri="https://zenodo.org/record/6352466/files/figure.png" pageId="2" pageNumber="283">Figs. 2, 4</figureCitation>
), slightly darker at margins and lighter around fovea. Ocular region black. Broad, indistinct line of feathery black setae from PER to posterior slope of carapace. Black erect setae in line extending straight from just posterior of PME to just anterior to fovea. Sternum bright yellowish-orange (
<figureCitation id="13722A45E011FFDFC7012C18FE75AA43" box="[362,431,1838,1864]" captionStart="FIGURES 2–4" captionStartId="3.[151,264,1490,1514]" captionTargetBox="[161,1432,191,1460]" captionTargetId="figure-19@3.[151,1436,181,1466]" captionTargetPageId="3" captionText="FIGURES 2–4. Copa sakalava sp. nov. Male holotype habitus. 2 dorsal, 3 line drawing of dorsal habitus detail, 4 ventral." figureDoi="http://doi.org/10.5281/zenodo.6352466" httpUri="https://zenodo.org/record/6352466/files/figure.png" pageId="2" pageNumber="283">Fig. 3</figureCitation>
), coxae all paler. Dorsal sclerite deep reddish-orange, otherwise abdomen dorsally beige to greyish, venter beige to cream, epigastric region orangish. Legs I &amp; II generally yellowish-brown, legs III &amp; IV generally brownish-orange and clearly darker than first two pairs. Mt III &amp; IV deep orangish-red.
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2CACFA9BAABF" blockId="2.[151,1437,937,2008]" box="[199,1345,1945,1973]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2CACFEE5AABF" box="[199,319,1946,1972]" italics="true" pageId="2" pageNumber="283">Carapace:</emphasis>
Broad, width about 4/5
<superScript id="7C3C9B88E011FFDFC4232CAFFD8FAAAC" attach="left" box="[584,597,1945,1959]" fontSize="6" pageId="2" pageNumber="283">th</superScript>
length. Highest point at fovea, sloping abruptly posterior to fovea.
</paragraph>
<paragraph id="8BF636C0E011FFDFC6AC2C88FB47AAD3" blockId="2.[151,1437,937,2008]" box="[199,1181,1982,2008]" pageId="2" pageNumber="283">
<emphasis id="B93DEAD2E011FFDFC6AC2C88FEF4AAD3" box="[199,302,1982,2008]" italics="true" pageId="2" pageNumber="283">Sternum:</emphasis>
Broad, shield-shaped, anterior ridge straight. Widest between coxae II and III.
</paragraph>
<caption id="DF366648E010FFDEC6FC2EE4FAAAA8E1" ID-DOI="http://doi.org/10.5281/zenodo.6352466" ID-Zenodo-Dep="6352466" box="[151,1392,1490,1514]" httpUri="https://zenodo.org/record/6352466/files/figure.png" pageId="3" pageNumber="284" startId="3.[151,264,1490,1514]" targetBox="[161,1432,191,1460]" targetPageId="3">
<paragraph id="8BF636C0E010FFDEC6FC2EE4FAAAA8E1" blockId="3.[151,1392,1490,1514]" box="[151,1392,1490,1514]" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6FC2EE4FEE0A8E1" bold="true" box="[151,314,1490,1514]" pageId="3" pageNumber="284">FIGURES 2–4.</emphasis>
<taxonomicName id="4C494D43E010FFDEC72A2EE5FE02A8E1" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[321,472,1490,1514]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E010FFDEC72A2EE5FE02A8E1" box="[321,472,1490,1514]" italics="true" pageId="3" pageNumber="284">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E010FFDEC7B52EE4FDF5A8E1" bold="true" box="[478,559,1490,1514]" pageId="3" pageNumber="284">
<taxonomicNameLabel id="A20E57A9E010FFDEC7B52EE4FDF5A8E1" box="[478,559,1490,1514]" pageId="3" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male holotype habitus. 2 dorsal, 3 line drawing of dorsal habitus detail, 4 ventral.
</paragraph>
</caption>
<paragraph id="8BF636C0E010FFDEC6AC2D2FFB35AB5C" blockId="3.[151,1437,1560,2019]" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6AC2D2FFEDFAB39" box="[199,261,1561,1586]" italics="true" pageId="3" pageNumber="284">Eyes:</emphasis>
AER procurved with AMEs largest, close to touching ALE. PER strongly procurved, with PME slightly larger larger than PLE. Strong, short, ocular setae in horizontal rows of six between PLE and PME.
</paragraph>
<paragraph id="8BF636C0E010FFDEC6AC2D57FD89ABC9" blockId="3.[151,1437,1560,2019]" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6AC2D57FEDCAB71" box="[199,262,1633,1658]" italics="true" pageId="3" pageNumber="284">Legs:</emphasis>
Dorsal, prolateral and retrolateral spines absent from tibiae I, II. All femora with sparse erect ventral setae, patellae with and fine, long setae dorsally. Femur IV distinctly broader than others. Ti, Mt and Ta with dorsal, prolateral and retrolateral trichobothria.
</paragraph>
<paragraph id="8BF636C0E010FFDEC6AC2DFAFD4AAA01" blockId="3.[151,1437,1560,2019]" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6AC2DFAFE91ABED" box="[199,331,1740,1766]" italics="true" pageId="3" pageNumber="284">Chelicerae:</emphasis>
Covered with relatively long, erect setae on anterior face. Two teeth on retromargin, well-spaced, about equal-sized, distal tooth slightly larger.
</paragraph>
<paragraph id="8BF636C0E010FFDEC6AC2C22FB14AA91" blockId="3.[151,1437,1560,2019]" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6AC2C22FEE0AA25" box="[199,314,1812,1838]" italics="true" pageId="3" pageNumber="284">Abdomen:</emphasis>
Six strong straight setae on anterior margin of dorsal sclerite. Dorsal sclerite around half length of abdomen and posterior margin bifid. Dorsum covered by short straight black setae and feathery black setae, denser at lateral margins. Venter with sparser black setae. Inframamillary sclerite small, circular, with dense short setae. Epigastric region with moderate sclerotisation and straight black setae medially, absent laterally.
</paragraph>
<paragraph id="8BF636C0E010FFD9C6AC2C92FC04ADF2" blockId="3.[151,1437,1560,2019]" lastBlockId="4.[151,1436,151,393]" lastPageId="4" lastPageNumber="285" pageId="3" pageNumber="284">
<emphasis id="B93DEAD2E010FFDEC6AC2C92FEDFAAB5" box="[199,261,1956,1982]" italics="true" pageId="3" pageNumber="284">Palp:</emphasis>
(
<figureCitation id="13722A45E010FFDEC77F2C92FEA6AAB5" box="[276,380,1956,1982]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1834,1858]" captionTargetBox="[164,1422,195,1800]" captionTargetId="figure-19@5.[151,1436,181,1810]" captionTargetPageId="5" captionText="FIGURES 8–11. Copa sakalava sp. nov. genitalia illustrations. 8-9 male pedipalp, 10-11 female epigyne. 8, 10 ventral, 9 retrolateral, 11 dorsal. CO = copulatory openings, CD = copulatory duct, Em = embolus, FD = fertilisation duct, H = epigynal hood, LC = Lateral chamber of copulatory duct, PcS = paracymbial spine, SD = sperm duct, St = subtegulum, ST I &amp; ST II = spermathecae I and II. Scale bar for 10 &amp; 11 = 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.6352470" httpUri="https://zenodo.org/record/6352470/files/figure.png" pageId="3" pageNumber="284">Figs. 8, 9</figureCitation>
) Cymbium orange to brown. One large prolateral spine on tibia, one on prolateral edge of cymbium. Retrolateral paracymbial spine sharp but stubby. Tegulum pear-shaped with sperm duct deep purple to black. Embolus with relatively broad basal ridge around 2/3 width of cymbium, distally to ridge-embolic turn broader, around ¾ width of cymbium, turning 1½ times with embolus tip directed prolaterally and extending between 1/3 and ½ length of distal coil. Several thicker short setae at apex of cymbium.
</paragraph>
<paragraph id="8BF636C0E017FFD9C6AC2A35FE28AC82" blockId="4.[151,1436,151,393]" pageId="4" pageNumber="285">
<emphasis id="B93DEAD2E017FFD9C6AC2A35FEBAAC17" box="[199,352,259,284]" italics="true" pageId="4" pageNumber="285">Leg spination</emphasis>
: I: F = pl1 do3 rl1, P = d1, Ti = plv2 rlv2 (one additional much smaller spine plv), Mt = plv2 rlv2. II: F = pl1 do3 rl1, P = d1, Ti = plv2 rlv2 (one additional much smaller spine plv), Mt = plv2 rlv2. III: F = pl2 do3 rl2, P = do1, Ti = pl2 d1 rl3 plv3 rlv3, Mt = pl1 do4 rl1 plv2 rlv2. IV: F = pl2 d3 rl2, P = d1, Ti = pl2 d1 rl3 plv2 rlv2, Mt = pl2 d4 rl2 plv2 rlv2.
</paragraph>
<caption id="DF366648E017FFD9C6FC2EE2FB25A8E7" ID-DOI="http://doi.org/10.5281/zenodo.6352468" ID-Zenodo-Dep="6352468" box="[151,1279,1492,1516]" httpUri="https://zenodo.org/record/6352468/files/figure.png" pageId="4" pageNumber="285" startId="4.[151,264,1492,1516]" targetBox="[159,1436,420,1457]" targetPageId="4">
<paragraph id="8BF636C0E017FFD9C6FC2EE2FB25A8E7" blockId="4.[151,1279,1492,1516]" box="[151,1279,1492,1516]" pageId="4" pageNumber="285">
<emphasis id="B93DEAD2E017FFD9C6FC2EE2FEE0A8E7" bold="true" box="[151,314,1492,1516]" pageId="4" pageNumber="285">FIGURES 5–7.</emphasis>
<taxonomicName id="4C494D43E017FFD9C72A2EE2FE02A8E7" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[321,472,1492,1516]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E017FFD9C72A2EE2FE02A8E7" box="[321,472,1492,1516]" italics="true" pageId="4" pageNumber="285">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E017FFD9C7B52EE2FDF5A8E7" bold="true" box="[478,559,1492,1516]" pageId="4" pageNumber="285">
<taxonomicNameLabel id="A20E57A9E017FFD9C7B52EE2FDF5A8E7" box="[478,559,1492,1516]" pageId="4" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female paratype habitus. 5 dorsal, 6 detail of dorsal habitus, 7 ventral.
</paragraph>
</caption>
<paragraph id="8BF636C0E017FFD9C6AC2D2CFE43AB3F" blockId="4.[151,1436,1562,1985]" box="[199,409,1562,1588]" pageId="4" pageNumber="285">
<emphasis id="B93DEAD2E017FFD9C6AC2D2CFEC4AB3F" bold="true" box="[199,286,1562,1588]" pageId="4" pageNumber="285">Female</emphasis>
(
<typeStatus id="54F28862E017FFD9C7462D2CFE4BAB3F" box="[301,401,1562,1588]" pageId="4" pageNumber="285" type="paratype">paratype</typeStatus>
)
</paragraph>
<paragraph id="8BF636C0E017FFD9C6AC2D09FC82ABCF" blockId="4.[151,1436,1562,1985]" pageId="4" pageNumber="285">
Measurements. TL 6.36, CL 2.82, CW 2.12,
<collectingCountry id="F35E7650E017FFD9C4A82D08FD30AB53" box="[707,746,1598,1624]" name="Switzerland" pageId="4" pageNumber="285">CH</collectingCountry>
1.12, SL 1.42, SW 1.24, AL 3.54, AW 2.68, chelicera length 1.00, chelicera width 0.52. Legs. I: 1.82, 0.78, 1.48, 1.28, 1.00. II: 1.78, 0.70, 1.36, 1.30, 0.86. III: 1.73, 0.70, 1.22, 1.60, 0.88. IV: 2.04, 0.80, 1.44, 2.38, 1.10. Eyes: AME—0.10, ALE—0.08, PME—0.10, PLE 0.08. AME—ALE 0.02, PLE—PME—0.05, AME-PME—0.14, ALE-PLE—0.04.
</paragraph>
<paragraph id="8BF636C0E017FFD9C6AC2DF8FCDFAA06" blockId="4.[151,1436,1562,1985]" pageId="4" pageNumber="285">
Shape, colouration, details of eyes, legs and chelicerae all as in male, except: dorsal sclerite very small, covering only around 1/8
<superScript id="7C3C9B88E017FFD9C7182DC4FE5AAA0B" attach="left" box="[371,384,1778,1792]" fontSize="6" pageId="4" pageNumber="285">th</superScript>
of dorsum, light brownish orange.
</paragraph>
<paragraph id="8BF636C0E017FFD9C6AC2C21FB6BAACA" blockId="4.[151,1436,1562,1985]" pageId="4" pageNumber="285">
<emphasis id="B93DEAD2E017FFD9C6AC2C21FEF6AA3B" box="[199,300,1815,1840]" italics="true" pageId="4" pageNumber="285">Epigyne:</emphasis>
(
<figureCitation id="13722A45E017FFD9C7572C20FE18AA3B" box="[316,450,1814,1840]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1834,1858]" captionTargetBox="[164,1422,195,1800]" captionTargetId="figure-19@5.[151,1436,181,1810]" captionTargetPageId="5" captionText="FIGURES 8–11. Copa sakalava sp. nov. genitalia illustrations. 8-9 male pedipalp, 10-11 female epigyne. 8, 10 ventral, 9 retrolateral, 11 dorsal. CO = copulatory openings, CD = copulatory duct, Em = embolus, FD = fertilisation duct, H = epigynal hood, LC = Lateral chamber of copulatory duct, PcS = paracymbial spine, SD = sperm duct, St = subtegulum, ST I &amp; ST II = spermathecae I and II. Scale bar for 10 &amp; 11 = 0.25mm." figureDoi="http://doi.org/10.5281/zenodo.6352470" httpUri="https://zenodo.org/record/6352470/files/figure.png" pageId="4" pageNumber="285">Figs. 10, 11</figureCitation>
). Roughly square due to lateral chamber of CD, large semi-circular external ridges with lightly translucent hood around midpoint of epigyne, hood margin anteriorly at lateral margin of ST and touching at their mid-point for 1/3 their length posteriorly; CO just posterior to anterior apex of hood, CD highly distinctive, directed dorsally and slightly obliquely toward both ventral portion of CD and lateral chamber of CD (in anterior view) situated between CO and distinctive looped connection of CD to posterior end of ST II.
</paragraph>
<caption id="DF366648E016FFD8C6FC2C1CFD14AAA5" ID-DOI="http://doi.org/10.5281/zenodo.6352470" ID-Zenodo-Dep="6352470" httpUri="https://zenodo.org/record/6352470/files/figure.png" pageId="5" pageNumber="286" startId="5.[151,264,1834,1858]" targetBox="[164,1422,195,1800]" targetPageId="5">
<paragraph id="8BF636C0E016FFD8C6FC2C1CFD14AAA5" blockId="5.[151,1437,1834,1966]" pageId="5" pageNumber="286">
<emphasis id="B93DEAD2E016FFD8C6FC2C1CFE93AA49" bold="true" box="[151,329,1834,1858]" pageId="5" pageNumber="286">FIGURES 8–11.</emphasis>
<taxonomicName id="4C494D43E016FFD8C7392C1CFE37AA49" authority="Pett &amp; Rabemananjara, 2022" authorityName="Pett &amp; Rabemananjara" authorityYear="2022" box="[338,493,1834,1858]" class="Arachnida" family="Corinnidae" genus="Copa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="286" phylum="Arthropoda" rank="species" species="sakalava" status="sp. nov.">
<emphasis id="B93DEAD2E016FFD8C7392C1CFE37AA49" box="[338,493,1834,1858]" italics="true" pageId="5" pageNumber="286">Copa sakalava</emphasis>
</taxonomicName>
<emphasis id="B93DEAD2E016FFD8C79D2C1CFD90AA49" bold="true" box="[502,586,1834,1858]" pageId="5" pageNumber="286">
<taxonomicNameLabel id="A20E57A9E016FFD8C79D2C1CFD90AA49" box="[502,586,1834,1858]" pageId="5" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
genitalia illustrations. 8-9 male pedipalp, 10-11 female epigyne. 8, 10 ventral, 9 retrolateral, 11 dorsal. CO = copulatory openings, CD = copulatory duct, Em = embolus, FD = fertilisation duct, H = epigynal hood, LC = Lateral chamber of copulatory duct, PcS = paracymbial spine, SD = sperm duct, St = subtegulum, ST I &amp; ST II = spermathecae I and II. Scale bar for 10 &amp; 11 = 0.25mm.
</paragraph>
</caption>
<paragraph id="8BF636C0E015FFDBC6AC2BA1FCF0AC16" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="287">
<emphasis id="B93DEAD2E015FFDBC6AC2BA1FEB3ADBB" box="[199,361,151,176]" italics="true" pageId="6" pageNumber="287">Leg spination:</emphasis>
I: Ti = plv2 rlv2 (one additional much smaller spine at plv apex), Mt = plv2 rlv2. II: F = pl1 do3 rl1, P = d1, Ti = plv1 rlv2 (one additional much smaller spine at plv apex), Mt = plv2 rlv2. III: F = pl2 do3 rl2, P = d1, Ti = pl2 d1 rl2 plv3 rlv2, Mt = pl3 d4 rl1 plv2 rlv2 (4 distal spines around apex of segment). IV: F = pl2 do3 rl2, P = d1, Ti = pl2 d1 rl3 plv3 rlv2, Mt = pl2 d4 rl2 plv2 rlv2.
</paragraph>
</subSubSection>
</treatment>
</document>
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