865 lines
178 KiB
XML
865 lines
178 KiB
XML
<document id="13E13B5C3AF045756B94DFF49C71374B" ID-DOI="10.11646/zootaxa.3368.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5252292" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629789001705" checkinUser="felipe" docAuthor="Sørensen, Martin V., Rho, Hyun Soo, Min, Won-Gi, Kim, Dongsung & Chang, Cheon Young" docDate="2012" docId="03E08792E204FFA1FF36FDB2FEA1F9D8" docLanguage="en" docName="zt03368p196.pdf" docOrigin="Zootaxa 3368 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3368.1.8" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Echinoderes cernunnos Sørensen & Rho & Min & Kim & Chang 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="175" masterDocId="FFD9FFEAE20CFFAFFFA1FF8DFFF0FF88" masterDocTitle="An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934 *" masterLastPageNumber="196" masterPageNumber="161" pageNumber="169" updateTime="1699112222437" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F5CF658567244C21B4C7F46FA71BC255">
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<mods:title id="308F71E268209106702510A8601E51D1">An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934 *</mods:title>
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<mods:name id="07ECBC5C888BEA9A666920D48FE90F94" type="personal">
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<mods:namePart id="CCECA5473769D1DB9F462DE0652A3E23">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="D8B8540F365580FA25303A9EBBFE54C1">Zoological Museum, The Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="E18DA137665E34FA7A0E54CFF372AA1B" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="DE37CE1356A933E186541CD728214828">Rho, Hyun Soo</mods:namePart>
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<mods:affiliation id="08AF0E814884616BB69CAD1B5A966B8F">Dokdo Research Center, Korea Ocean Research and Development Institute, Uljin 767 - 813, Korea</mods:affiliation>
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</mods:name>
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<mods:name id="473E18D418DA8F4FB6D594CDFE38A2B9" type="personal">
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<mods:roleTerm id="CB7484F1F75FC1F3D94825EAED62B8DB">Author</mods:roleTerm>
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<mods:namePart id="687BD19D8F0724022797B56E3D00367B">Min, Won-Gi</mods:namePart>
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<mods:affiliation id="C6C4B0DD9B8588E18F85C174123B9C57">Dokdo Research Center, Korea Ocean Research and Development Institute, Uljin 767 - 813, Korea</mods:affiliation>
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<mods:namePart id="51194FF4973A18A8BFBAC6880E27E9BB">Kim, Dongsung</mods:namePart>
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<mods:affiliation id="68B55A1CF78B9C0C6885A4F9B8710795">Marine Living Resources Research Department, Korea Ocean Research and Development Institute, Ansan 425 - 600, Korea Department of Biological Sciences, College of Natural Sciences, Daegu University, Gyeongsan 712 - 714, Korea</mods:affiliation>
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<mods:namePart id="75AF012813E5CB6603A7CA5D5DD32DB1">Chang, Cheon Young</mods:namePart>
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<mods:title id="0BFC149EB37062012759EB6A5EC6CD6E">Zootaxa</mods:title>
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<mods:date id="77655CD876AB2EE99D9AB70175B59045">2012</mods:date>
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<mods:number id="F32EFF91D6A27F8A80C2F83E91D525C0">2012-07-04</mods:number>
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<mods:url id="51D062CDDAB3D1237E26CF03066253FE">https://biotaxa.org/Zootaxa/article/view/zootaxa.3368.1.8</mods:url>
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<mods:classification id="4A96BBC3134E7FF3FEA397D0E7E46765">journal article</mods:classification>
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<mods:identifier id="4568439E71E0D80AEFA5E4ED8A1819AB" type="DOI">10.11646/zootaxa.3368.1.8</mods:identifier>
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<mods:identifier id="822319FC9C4CE78CC14F6146B9716ABD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E08792E204FFA1FF36FDB2FEA1F9D8" LSID="urn:lsid:plazi:treatment:03E08792E204FFA1FF36FDB2FEA1F9D8" httpUri="http://treatment.plazi.org/id/03E08792E204FFA1FF36FDB2FEA1F9D8" lastPageId="14" lastPageNumber="175" pageId="8" pageNumber="169">
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<subSubSection id="C353650FE204FFA7FF36FDB2FDEBFDD1" box="[151,539,575,601]" pageId="8" pageNumber="169" type="nomenclature">
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<paragraph id="8BF63684E204FFA7FF36FDB2FDEBFDD1" blockId="8.[151,539,575,635]" box="[151,539,575,601]" pageId="8" pageNumber="169">
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<heading id="D0BE81E8E204FFA7FF36FDB2FDEBFDD1" bold="true" box="[151,539,575,601]" fontSize="11" level="1" pageId="8" pageNumber="169" reason="1">
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<emphasis id="B93DEA96E204FFA7FF36FDB2FDEBFDD1" bold="true" box="[151,539,575,601]" pageId="8" pageNumber="169">
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<taxonomicName id="4C494D07E204FFA7FF36FDB2FE45FDD1" ID-CoL="6DVK5" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[151,437,575,601]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
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<emphasis id="B93DEA96E204FFA7FF36FDB2FE45FDD1" bold="true" box="[151,437,575,601]" italics="true" pageId="8" pageNumber="169">Echinoderes cernunnos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57EDE204FFA7FE1CFDB2FDEBFDD1" box="[445,539,575,601]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353650FE204FFA7FF36FDEEFE46FDF3" box="[151,438,611,635]" pageId="8" pageNumber="169" type="description">
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<paragraph id="8BF63684E204FFA7FF36FDEEFE46FDF3" blockId="8.[151,539,575,635]" box="[151,438,611,635]" pageId="8" pageNumber="169">
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(
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<figureCitation id="13722A01E204FFA7FF3FFDEEFED8FDF3" box="[158,296,611,635]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="10.[151,244,1865,1886]" captionStartId-1="12.[151,244,1752,1773]" captionStartId-2="13.[151,244,1696,1717]" captionTargetBox-0="[217,1376,206,1837]" captionTargetBox-1="[220,1376,179,1732]" captionTargetBox-2="[186,1398,199,1673]" captionTargetId-0="figure-100@10.[207,1379,193,1844]" captionTargetId-1="figure-193@12.[212,1385,169,1739]" captionTargetId-2="figure-183@13.[185,1401,193,1675]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." captionText-1="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." captionText-2="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi-0="http://doi.org/10.5281/zenodo.5252308" figureDoi-1="http://doi.org/10.5281/zenodo.5252314" figureDoi-2="http://doi.org/10.5281/zenodo.5252318" httpUri-0="https://zenodo.org/record/5252308/files/figure.png" httpUri-1="https://zenodo.org/record/5252314/files/figure.png" httpUri-2="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Figures 4–6</figureCitation>
|
||
,
|
||
<tableCitation id="C6CB033FE204FFA7FE92FDEEFE5EFDF3" box="[307,430,611,635]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="9.[151,239,151,175]" captionStartId-1="9.[151,239,1339,1363]" captionText-0="TABLE 3. Measurements of female holotype and two female paratypes of Echinoderes cernunnos sp. nov. from the Korea Strait (MAP-27) and East China Sea (MAP-08). Abbreviations: (ac): acicular spine; (f): putative female condition of sexual dimorphic character; LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal; MSW-6: Maximum sternal width, measured on segment 6 in this species; S: segment lengths; SW-10: standard width, always measured on segment 10; TL: trunk length. Dash –: indicates that the structure could not be measured." captionText-1="TABLE 4. Summary of nature and location of sensory spots, glandular cell outlets and spines arranged by series in Echinoderes cernunnos sp. nov. from the Korean South Sea. Abbreviations: LA: Lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; (f): female condition of sexual dimorphic character; gco1/2: glandular cell outlet type 1/2; ltas: lateral terminal accessory spine; lts: lateral terminal spine; si: sieve plate; ss: sensory spot; tu: tubule." pageId="8" pageNumber="169">Tables 3–4</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353650FE204FFA7FF36FD24FCD9FC1C" pageId="8" pageNumber="169" type="diagnosis">
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<paragraph id="8BF63684E204FFA7FF36FD24FCD9FC1C" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
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<emphasis id="B93DEA96E204FFA7FF36FD24FEE1FD4A" bold="true" box="[151,273,681,706]" inLineHeading="true" pageId="8" pageNumber="169" reason="1">Diagnosis.</emphasis>
|
||
Segments 1 and 2 consisting of closed rings; segments 3 to 10 of one tergal and two sternal plates, and segment 11 consisting of two tergal and two sternal plates. Specimens with middorsal spines on segment 4–8, gradually increasing in length towards the more posterior segments; lateroventral tubule on segment 5; lateroventral spines on segments 6–9; very short, laterodorsal, distally fringed elements (tubules?) on segment 10. Glandular cell outlets
|
||
<typeStatus id="54F28826E204FFA7FE2EFCBAFE4FFCC7" box="[399,447,823,847]" pageId="8" pageNumber="169">type</typeStatus>
|
||
2 present in subdorsal, laterodorsal, sublateral and ventrolateral positions on segment 2, midlateral positions on segments 5 and 7, and in sublateral position of segment 8. Tergal extension of segment 11 extremely elongated, forming strong, horn-like extensions.
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C353650FE204FFA7FF66FC2DFE25FB51" pageId="8" pageNumber="169" type="materials_examined">
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<paragraph id="8BF63684E204FFA7FF66FC2DFF38FB1B" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
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<emphasis id="B93DEA96E204FFA7FF66FC2DFE86FC30" bold="true" box="[199,374,927,952]" inLineHeading="true" pageId="8" pageNumber="169" reason="1">Type material.</emphasis>
|
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<materialsCitation id="3B213CD9E204FFA7FEDEFC12FE83FBAD" collectingDate="2008-10-06" collectionCode="NIBR" collectorName="Fluoromount G & No" country="South Korea" county="Korean" elevation="96" latitude="34.2735" location="Strait" longLatPrecision="11" longitude="128.67334" municipality="Tsushima Island" pageId="8" pageNumber="169" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54F28826E204FFA7FEDEFC12FE1CFC3F" box="[383,492,927,951]" pageId="8" pageNumber="169" type="holotype">Holotype</typeStatus>
|
||
: adult female, collected on
|
||
<date id="FFF71044E204FFA7FC87FC12FC2BFC3F" box="[806,987,927,951]" pageId="8" pageNumber="169" value="2008-10-06">
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<collectingDate id="EFB3E9ACE204FFA7FC87FC12FC2BFC3F" box="[806,987,927,951]" pageId="8" pageNumber="169" value="2008-10-06">6 October 2008</collectingDate>
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</date>
|
||
, at station MAP-27, in the
|
||
<collectingCountry id="F35E7614E204FFA7FAB2FC2DFAA7FC3F" box="[1299,1367,928,951]" name="South Korea" pageId="8" pageNumber="169">Korea</collectingCountry>
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<location id="8E96605FE204FFA7FAC1FC12FA6CFC3F" LSID="urn:lsid:plazi:treatment:03E08792E204FFA1FF36FDB2FEA1F9D8:8E96605FE204FFA7FAC1FC12FA6CFC3F" box="[1376,1436,927,951]" country="South Korea" county="Korean" latitude="34.2735" longLatPrecision="11" longitude="128.67334" municipality="Tsushima Island" name="Strait" pageId="8" pageNumber="169">Strait</location>
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between
|
||
<collectingMunicipality id="6B92ACFEE204FFA7FF5DFC4AFE43FC57" box="[252,435,967,991]" pageId="8" pageNumber="169">Tsushima Island</collectingMunicipality>
|
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and the
|
||
<collectingCounty id="62974E08E204FFA7FDB4FC45FD97FC57" box="[533,615,968,991]" pageId="8" pageNumber="169">Korean</collectingCounty>
|
||
mainland (
|
||
<figureCitation id="13722A01E204FFA7FD49FC4AFCCEFC57" box="[744,830,967,991]" captionStart="FIGURE 1" captionStartId="1.[151,244,1947,1968]" captionTargetBox="[179,1407,1068,1890]" captionTargetId="figure-431@1.[179,1407,1068,1927]" captionTargetPageId="1" captionText="FIGURE 1. Maps showing collecting localities for: A, Echinoderes aspinosus sp. nov. and Echinoderes obtuspinosus sp. nov.; B, Echinoderes cernunnos sp. nov.; C, Echinoderes microaperturus sp. nov.; D, Echinoderes tchefouensis Lou, 1934. Type localities are marked with a ring around the dot." figureDoi="http://doi.org/10.5281/zenodo.5252296" httpUri="https://zenodo.org/record/5252296/files/figure.png" pageId="8" pageNumber="169">Fig. 1B</figureCitation>
|
||
),
|
||
<geoCoordinate id="EE7D5043E204FFA7FCF7FC4AFC25FC57" box="[854,981,962,991]" degrees="34" direction="north" minutes="16.41" orientation="latitude" pageId="8" pageNumber="169" precision="9" value="34.2735">
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||
34
|
||
<superScript id="7C3C9BCCE204FFA7FCD0FC4FFC89FC58" attach="none" box="[881,889,962,976]" fontSize="6" pageId="8" pageNumber="169">o</superScript>
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||
16.41’N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE7D5043E204FFA7FC7CFC4AFB9BFC57" box="[989,1131,962,991]" degrees="128" direction="east" minutes="40.40" orientation="longitude" pageId="8" pageNumber="169" precision="9" value="128.67334">
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128
|
||
<superScript id="7C3C9BCCE204FFA7FBA6FC4FFBFFFC58" attach="none" box="[1031,1039,962,976]" fontSize="6" pageId="8" pageNumber="169">o</superScript>
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40.40’E
|
||
</geoCoordinate>
|
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, from mud with tiny shells at
|
||
<elevation id="0064D1B7E204FFA7FF15FC67FEC7FB8A" box="[180,311,1002,1026]" metricMagnitude="1" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="169" unit="m" value="96.0">
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<quantity id="4CB19B61E204FFA7FF15FC67FF1EFB8A" box="[180,238,1002,1026]" metricMagnitude="1" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="169" unit="m" value="96.0">96 m</quantity>
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||
depth
|
||
</elevation>
|
||
, mounted in
|
||
<collectorName id="26BC5352E204FFA7FE6CFC67FD8BFB8A" box="[461,635,1002,1026]" pageId="8" pageNumber="169">Fluoromount G</collectorName>
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, deposited at
|
||
<collectionCode id="ED58AE41E204FFA7FCB9FC66FCABFB8A" box="[792,859,1003,1026]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="8" pageNumber="169">NIBR</collectionCode>
|
||
under accession number INBRIV0000245082.
|
||
<collectorName id="26BC5352E204FFA7FADBFC66FA6CFB8A" box="[1402,1436,1003,1026]" pageId="8" pageNumber="169">No</collectorName>
|
||
<typeStatus id="54F28826E204FFA7FF36FB80FF01FBAD" box="[151,241,1037,1061]" pageId="8" pageNumber="169" type="allotype">allotype</typeStatus>
|
||
designated
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B213CD9E204FFA7FEDFFB83FCBDFBF8" collectingDate="2008-06-06" collectionCode="NHMD" collectorName="Fluoromount G" country="China" county="Jeju Island" elevation="113" latitude="32.359833" location="East" longLatPrecision="12" longitude="126.772" municipality="Sea" pageId="8" pageNumber="169" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<typeStatus id="54F28826E204FFA7FEDFFB83FE1FFBAE" box="[382,495,1038,1062]" pageId="8" pageNumber="169" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4FFD0DE204FFA7FE58FB83FD74FBAD" box="[505,644,1037,1061]" pageId="8" pageNumber="169" type="female">two females</specimenCount>
|
||
, collected on
|
||
<date id="FFF71044E204FFA7FCBEFB80FC56FBAD" box="[799,934,1037,1061]" pageId="8" pageNumber="169" value="2008-06-06">
|
||
<collectingDate id="EFB3E9ACE204FFA7FCBEFB80FC56FBAD" box="[799,934,1037,1061]" pageId="8" pageNumber="169" value="2008-06-06">6 June 2008</collectingDate>
|
||
</date>
|
||
from station MAP-08 in the
|
||
<location id="8E96605FE204FFA7FB4AFB83FAEBFBAD" LSID="urn:lsid:plazi:treatment:03E08792E204FFA1FF36FDB2FEA1F9D8:8E96605FE204FFA7FB4AFB83FAEBFBAD" box="[1259,1307,1038,1061]" country="China" county="Jeju Island" latitude="32.359833" longLatPrecision="12" longitude="126.772" municipality="Sea" name="East" pageId="8" pageNumber="169">East</location>
|
||
<collectingCountry id="F35E7614E204FFA7FA83FB80FA95FBAD" box="[1314,1381,1037,1061]" name="China" pageId="8" pageNumber="169">China</collectingCountry>
|
||
<collectingMunicipality id="6B92ACFEE204FFA7FACDFB80FA67FBAD" box="[1388,1431,1037,1061]" pageId="8" pageNumber="169">Sea</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4CB19B61E204FFA7FF62FBB8FEECFBC5" box="[195,284,1077,1101]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="169" unit="km" value="100.0">100 km</quantity>
|
||
south of
|
||
<collectingCounty id="62974E08E204FFA7FE34FBB8FDE5FBC5" box="[405,533,1077,1101]" pageId="8" pageNumber="169">Jeju Island</collectingCounty>
|
||
, (
|
||
<figureCitation id="13722A01E204FFA7FD8FFBB8FD79FBC5" box="[558,649,1077,1101]" captionStart="FIGURE 1" captionStartId="1.[151,244,1947,1968]" captionTargetBox="[179,1407,1068,1890]" captionTargetId="figure-431@1.[179,1407,1068,1927]" captionTargetPageId="1" captionText="FIGURE 1. Maps showing collecting localities for: A, Echinoderes aspinosus sp. nov. and Echinoderes obtuspinosus sp. nov.; B, Echinoderes cernunnos sp. nov.; C, Echinoderes microaperturus sp. nov.; D, Echinoderes tchefouensis Lou, 1934. Type localities are marked with a ring around the dot." figureDoi="http://doi.org/10.5281/zenodo.5252296" httpUri="https://zenodo.org/record/5252296/files/figure.png" pageId="8" pageNumber="169">Fig. 1B</figureCitation>
|
||
),
|
||
<geoCoordinate id="EE7D5043E204FFA7FD04FBB8FCD4FBC5" box="[677,804,1072,1101]" degrees="32" direction="north" minutes="21.59" orientation="latitude" pageId="8" pageNumber="169" precision="9" value="32.359833">
|
||
32
|
||
<superScript id="7C3C9BCCE204FFA7FD61FBBDFD38FBB6" attach="none" box="[704,712,1072,1086]" fontSize="6" pageId="8" pageNumber="169">o</superScript>
|
||
21.59’N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE7D5043E204FFA7FC90FBB8FC4EFBC5" box="[817,958,1072,1101]" degrees="126" direction="east" minutes="46.32" orientation="longitude" pageId="8" pageNumber="169" precision="9" value="126.772">
|
||
126
|
||
<superScript id="7C3C9BCCE204FFA7FCFAFBBDFC93FBB6" attach="none" box="[859,867,1072,1086]" fontSize="6" pageId="8" pageNumber="169">o</superScript>
|
||
46.32’E
|
||
</geoCoordinate>
|
||
, from mud at
|
||
<elevation id="0064D1B7E204FFA7FBD1FBB8FAF8FBC5" box="[1136,1288,1077,1101]" metricMagnitude="2" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="169" unit="m" value="113.0">
|
||
<quantity id="4CB19B61E204FFA7FBD1FBB8FB4BFBC4" box="[1136,1211,1077,1101]" metricMagnitude="2" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="169" unit="m" value="113.0">113 m</quantity>
|
||
depth
|
||
</elevation>
|
||
, mounted in
|
||
<collectorName id="26BC5352E204FFA7FF36FBD5FEB4FBF8" box="[151,324,1112,1136]" pageId="8" pageNumber="169">Fluoromount G</collectorName>
|
||
(one in lateral position), deposited at
|
||
<collectionCode id="ED58AE41E204FFA7FD59FBD4FCBDFBF8" box="[760,845,1113,1136]" pageId="8" pageNumber="169">NHMD</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B213CD9E204FFA7FCF4FBD5FF34FB1B" collectionCode="ZMUC" country="China" pageId="8" pageNumber="169" specimenCode="KIN-536, KIN- 537" specimenCount="1" typeStatus="paratype">
|
||
under accession number
|
||
<collectionCode id="ED58AE41E204FFA7FBCDFBD5FB4DFBF8" box="[1132,1213,1112,1136]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="169">ZMUC</collectionCode>
|
||
<specimenCode id="DBEF9EFFE204FFA7FB65FBD5FAD9FBF8" box="[1220,1321,1112,1136]" collectionCode="KIN-" pageId="8" pageNumber="169">KIN-536</specimenCode>
|
||
and
|
||
<specimenCode id="DBEF9EFFE204FFA7FAC0FBD4FF34FB1B" collectionCode="KIN-" pageId="8" pageNumber="169">KIN- 537</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E204FFA7FF66FB13FE25FB51" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
<materialsCitation id="3B213CD9E204FFA7FF66FB13FE20FB51" collectingDate="2008-10-06" collectionCode="SEM" collectorName="Fluoromount G & No" country="South Korea" county="Korean" elevation="96" latitude="34.2735" location="Strait" longLatPrecision="11" longitude="128.67334" municipality="Tsushima Island" pageId="8" pageNumber="169" specimenCount="1">
|
||
<emphasis id="B93DEA96E204FFA7FF66FB13FE4AFB3F" bold="true" box="[199,442,1182,1207]" inLineHeading="true" pageId="8" pageNumber="169" reason="1">Additional material.</emphasis>
|
||
One female from the same locality as the holotype, mounted for
|
||
<collectionCode id="ED58AE41E204FFA7FB3EFB13FB28FB3E" box="[1183,1240,1182,1206]" pageId="8" pageNumber="169">SEM</collectionCode>
|
||
and stored in the personal collection of MVS
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353650FE204FFA7FF66FB68FCFDFAA8" pageId="8" pageNumber="169" type="etymology">
|
||
<paragraph id="8BF63684E204FFA7FF66FB68FCFDFAA8" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
<emphasis id="B93DEA96E204FFA7FF66FB68FEBDFB75" bold="true" box="[199,333,1253,1277]" inLineHeading="true" pageId="8" pageNumber="169" reason="1">Etymology.</emphasis>
|
||
The species is named after Cernunnos—The Horned God—from Celtic mythology, inspired by the species’ diagnostic long, horn-like tergal extensions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353650FE204FFA1FF66FAA6FEA1F9D8" lastPageId="14" lastPageNumber="175" pageId="8" pageNumber="169" type="description">
|
||
<paragraph id="8BF63684E204FFA7FF66FAA6FBB3FA01" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
<emphasis id="B93DEA96E204FFA7FF66FAA6FEA8FACC" bold="true" box="[199,344,1323,1348]" inLineHeading="true" pageId="8" pageNumber="169" reason="1">Description.</emphasis>
|
||
Adult specimens consist of a head, a neck and eleven trunk segments (
|
||
<figureCitation id="13722A01E204FFA7FB2EFAA6FAFFFACB" box="[1167,1295,1323,1347]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E204FFA7FABCFAA6FAB3FACB" box="[1309,1347,1323,1347]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">6A</figureCitation>
|
||
). Trunk cuticle appears relatively thin and flexible. Measurements and dimensions are given in
|
||
<tableCitation id="C6CB033FE204FFA7FB2CFAC3FAF8FAEE" box="[1165,1288,1358,1382]" captionStart="TABLE 3" captionStartId="9.[151,239,151,175]" captionText="TABLE 3. Measurements of female holotype and two female paratypes of Echinoderes cernunnos sp. nov. from the Korea Strait (MAP-27) and East China Sea (MAP-08). Abbreviations: (ac): acicular spine; (f): putative female condition of sexual dimorphic character; LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal; MSW-6: Maximum sternal width, measured on segment 6 in this species; S: segment lengths; SW-10: standard width, always measured on segment 10; TL: trunk length. Dash –: indicates that the structure could not be measured." pageId="8" pageNumber="169">Table 3. A</tableCitation>
|
||
summary of sensory spot, spine, tubule and glandular cell outlet positions is provided in
|
||
<tableCitation id="C6CB033FE204FFA7FC4BFAFCFBCFFA01" box="[1002,1087,1393,1417]" captionStart="TABLE 4" captionStartId="9.[151,239,1339,1363]" captionText="TABLE 4. Summary of nature and location of sensory spots, glandular cell outlets and spines arranged by series in Echinoderes cernunnos sp. nov. from the Korean South Sea. Abbreviations: LA: Lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; (f): female condition of sexual dimorphic character; gco1/2: glandular cell outlet type 1/2; ltas: lateral terminal accessory spine; lts: lateral terminal spine; si: sieve plate; ss: sensory spot; tu: tubule." pageId="8" pageNumber="169">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E204FFA7FF66FA18FEA7F92B" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
The head consists of a retractable mouth cone and an introvert. Inner oral styles are clearly present, but their exact arrangement could not be examined. Outer armature with nine outer oral styles composed of two subunits. Bases of outer oral styles with a single fringe, flanked by a pair of off-set spikes (
|
||
<figureCitation id="13722A01E204FFA7FBEBFA56FB54FA7B" box="[1098,1188,1499,1523]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Fig. 6B</figureCitation>
|
||
). The arrangement of scalids on the introvert (
|
||
<figureCitation id="13722A01E204FFA7FE12FA73FDFCF99E" box="[435,524,1534,1558]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Fig. 6C</figureCitation>
|
||
) is identical with the one found in
|
||
<taxonomicName id="4C494D07E204FFA7FC0CFA72FB8CF99E" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[941,1148,1535,1558]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="microaperturus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E204FFA7FC0CFA72FB8CF99E" box="[941,1148,1535,1558]" italics="true" pageId="8" pageNumber="169">E. microaperturus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E204FFA7FB27FA72FB13F99F" bold="true" box="[1158,1251,1535,1559]" pageId="8" pageNumber="169">
|
||
<taxonomicNameLabel id="A20E57EDE204FFA7FB27FA72FB13F99F" box="[1158,1251,1535,1559]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see
|
||
<figureCitation id="13722A01E204FFA7FA84FA73FA99F99E" box="[1317,1385,1534,1558]" captionStart="FIGURE 8" captionStartId="18.[151,244,1912,1933]" captionTargetBox="[174,1417,181,1884]" captionTargetId="figure-65@18.[173,1417,181,1884]" captionTargetPageId="18" captionText="FIGURE 8. Diagram of mouth cone (grey shaded area), introvert and placids in Echinoderes microaperturus sp. nov. with indication of inner and outer oral styles, scalid and placid distribution. Placids and trichoscalid plates are symbolized by the black-shaded bent bars and circles around the introvert diagram." figureDoi="http://doi.org/10.5281/zenodo.5252328" httpUri="https://zenodo.org/record/5252328/files/figure.png" pageId="8" pageNumber="169">Fig. 8</figureCitation>
|
||
and description below) and several other species described herein. Leaf-like scalids (see definition under description of the following species) are present as single ones in sections 1, 5, 6 and 7, and pairs in sections 3 and 9. A distinct band of longitudinal ridges that extend into fringe tips stretches around the introvert at the level between Rings 05 and 06 (
|
||
<figureCitation id="13722A01E204FFA7FF53F906FEBAF92B" box="[242,330,1675,1699]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Fig. 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E204FFA7FF66F923FB93F8B8" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
The neck consists of 16 placids, all measuring 12 µm in length and 7 µm in width at bases (
|
||
<figureCitation id="13722A01E204FFA7FB61F923FACEF94E" box="[1216,1342,1710,1734]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E204FFA7FAE8F923FA7FF94E" box="[1353,1423,1710,1734]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">5A–B</figureCitation>
|
||
), except midventral placid that measures 11 µm in width (
|
||
<figureCitation id="13722A01E204FFA7FCE5F95CFC56F961" box="[836,934,1745,1769]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E204FFA7FC19F95CFC2CF961" box="[952,988,1745,1769]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">5B</figureCitation>
|
||
). Placids number 2 and 16 (counting clockwise from midventral placid) with broad trichoscalid plate and attached trichoscalid (
|
||
<figureCitation id="13722A01E204FFA7FB6CF978FADBF885" box="[1229,1323,1781,1805]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">Fig. 5B</figureCitation>
|
||
). Smaller trichoscalid plates with trichoscalids on placids number 6, 8, 10, and 12 (
|
||
<figureCitation id="13722A01E204FFA7FC69F895FBD5F8B8" box="[968,1061,1816,1840]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E204FFA7FB90F895FBA7F8B8" box="[1073,1111,1816,1840]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E204FFA7FF66F8B6FCCEF868" blockId="8.[151,1437,681,2016]" pageId="8" pageNumber="169">
|
||
Segment 1 consists of one complete cuticular ring (
|
||
<figureCitation id="13722A01E204FFA7FCE2F8B6FC39F8DB" box="[835,969,1851,1875]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E204FFA7FC7DF8B6FBD3F8DB" box="[988,1059,1851,1875]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">5A–B</figureCitation>
|
||
). Pairs of subdorsal (
|
||
<figureCitation id="13722A01E204FFA7FA8CF8B6FA7EF8DB" box="[1325,1422,1851,1875]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="8" pageNumber="169">Fig. 5A</figureCitation>
|
||
), laterodorsal (
|
||
<figureCitation id="13722A01E204FFA7FE8BF8D3FE71F8FE" box="[298,385,1886,1910]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Fig. 6D</figureCitation>
|
||
) and ventromedial (
|
||
<figureCitation id="13722A01E204FFA7FDC2F8D3FD49F8FE" box="[611,697,1886,1910]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="8" pageNumber="169">Fig. 4B</figureCitation>
|
||
) sensory spots present. Sensory spots are small and rounded, with numerous papillae. Only few cuticular hairs present; hairs emerge through rounded perforation sites, and are scattered around the segment (
|
||
<figureCitation id="13722A01E204FFA7FE4BF828FDB4F835" box="[490,580,1957,1981]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="8" pageNumber="169">Fig. 6D</figureCitation>
|
||
). Posterior margin with pectinate fringe; fringe appears serrated on its dorsal side, whereas the ventral fringe tips extend into filiform tips.
|
||
</paragraph>
|
||
<caption id="DF36660CE205FFA6FF36FF1AFAFFFEBA" pageId="9" pageNumber="170" startId="9.[151,239,151,175]">
|
||
<paragraph id="8BF63684E205FFA6FF36FF1AFAFFFEBA" blockId="9.[151,1436,151,306]" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FF36FF1AFEFAFF27" bold="true" box="[151,266,151,175]" pageId="9" pageNumber="170">TABLE 3.</emphasis>
|
||
Measurements of female holotype and two female paratypes of
|
||
<taxonomicName id="4C494D07E205FFA6FC09FF15FB6DFF27" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[936,1181,152,175]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="9" pageNumber="170" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E205FFA6FC09FF15FB6DFF27" box="[936,1181,152,175]" italics="true" pageId="9" pageNumber="170">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E205FFA6FB02FF1AFB07FF27" bold="true" box="[1187,1271,151,175]" pageId="9" pageNumber="170">
|
||
<taxonomicNameLabel id="A20E57EDE205FFA6FB02FF1AFB07FF27" box="[1187,1271,151,175]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the Korea Strait (MAP-27) and East China Sea (MAP-08). Abbreviations: (ac): acicular spine; (f): putative female condition of sexual dimorphic character; LTAS: lateral terminal accessory spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal; MSW-6: Maximum sternal width, measured on segment 6 in this species; S: segment lengths; SW-10: standard width, always measured on segment 10; TL: trunk length. Dash –: indicates that the structure could not be measured.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E205FFA6FF3EFEC0FB6BFB98" pageId="9" pageNumber="170">
|
||
<table id="F949C424E2050050FF3EFEC0FAF6FB73" box="[159,1286,333,1275]" gridcols="4" gridrows="29" pageId="9" pageNumber="170">
|
||
<tr id="357934C6E2050050FF3EFEC0FAF6FEEA" box="[159,1286,333,354]" gridrow="0" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFEC0FEFEFEEA" box="[159,270,333,354]" gridcol="0" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FF3EFEC0FEF8FEEA" bold="true" box="[159,264,333,354]" pageId="9" pageNumber="170">Character</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FE6CFEC0FD8AFEEA" box="[461,634,333,354]" gridcol="1" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FE6CFEC0FD8AFEEA" bold="true" box="[461,634,333,354]" pageId="9" pageNumber="170">Length Holotype</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FCB4FEC0FC4EFEEA" box="[789,958,333,354]" gridcol="2" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FCB4FEC0FC4EFEEA" bold="true" box="[789,958,333,354]" pageId="9" pageNumber="170">Length KIN-536</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FBFCFEC0FAF6FEEA" box="[1117,1286,333,354]" gridcol="3" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FBFCFEC0FAF6FEEA" bold="true" box="[1117,1286,333,354]" pageId="9" pageNumber="170">Length KIN-537</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFE1EFAF6FE20" box="[159,1286,403,424]" gridrow="1" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFE1EFEFEFE20" box="[159,270,403,424]" gridcol="0" gridrow="1" pageId="9" pageNumber="170">TL</th>
|
||
<td id="76A85DBAE2050050FE6CFE1EFD8AFE20" box="[461,634,403,424]" gridcol="1" gridrow="1" pageId="9" pageNumber="170">279 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FE1EFC4EFE20" box="[789,958,403,424]" gridcol="2" gridrow="1" pageId="9" pageNumber="170">295 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFE1EFAF6FE20" box="[1117,1286,403,424]" gridcol="3" gridrow="1" pageId="9" pageNumber="170">299 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFE3DFAF6FE4D" box="[159,1286,432,453]" gridrow="2" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFE3DFEFEFE4D" box="[159,270,432,453]" gridcol="0" gridrow="2" pageId="9" pageNumber="170">MSW-6</th>
|
||
<td id="76A85DBAE2050050FE6CFE3DFD8AFE4D" box="[461,634,432,453]" gridcol="1" gridrow="2" pageId="9" pageNumber="170">49 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FE3DFC4EFE4D" box="[789,958,432,453]" gridcol="2" gridrow="2" pageId="9" pageNumber="170">50 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFE3DFAF6FE4D" box="[1117,1286,432,453]" gridcol="3" gridrow="2" pageId="9" pageNumber="170">–</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFE43FAF6FE6B" box="[159,1286,462,483]" gridrow="3" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFE43FEFEFE6B" box="[159,270,462,483]" gridcol="0" gridrow="3" pageId="9" pageNumber="170">MSW-6/TL</th>
|
||
<td id="76A85DBAE2050050FE6CFE43FD8AFE6B" box="[461,634,462,483]" gridcol="1" gridrow="3" pageId="9" pageNumber="170">17.6%</td>
|
||
<td id="76A85DBAE2050050FCB4FE43FC4EFE6B" box="[789,958,462,483]" gridcol="2" gridrow="3" pageId="9" pageNumber="170">16.9%</td>
|
||
<td id="76A85DBAE2050050FBFCFE43FAF6FE6B" box="[1117,1286,462,483]" gridcol="3" gridrow="3" pageId="9" pageNumber="170">–</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFE66FAF6FD88" box="[159,1286,491,512]" gridrow="4" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFE66FEFEFD88" box="[159,270,491,512]" gridcol="0" gridrow="4" pageId="9" pageNumber="170">SW-10</th>
|
||
<td id="76A85DBAE2050050FE6CFE66FD8AFD88" box="[461,634,491,512]" gridcol="1" gridrow="4" pageId="9" pageNumber="170">39 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FE66FC4EFD88" box="[789,958,491,512]" gridcol="2" gridrow="4" pageId="9" pageNumber="170">40 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFE66FAF6FD88" box="[1117,1286,491,512]" gridcol="3" gridrow="4" pageId="9" pageNumber="170">–</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFD85FAF6FD95" box="[159,1286,520,541]" gridrow="5" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFD85FEFEFD95" box="[159,270,520,541]" gridcol="0" gridrow="5" pageId="9" pageNumber="170">SW-10/TL</th>
|
||
<td id="76A85DBAE2050050FE6CFD85FD8AFD95" box="[461,634,520,541]" gridcol="1" gridrow="5" pageId="9" pageNumber="170">14.0%</td>
|
||
<td id="76A85DBAE2050050FCB4FD85FC4EFD95" box="[789,958,520,541]" gridcol="2" gridrow="5" pageId="9" pageNumber="170">13.6%</td>
|
||
<td id="76A85DBAE2050050FBFCFD85FAF6FD95" box="[1117,1286,520,541]" gridcol="3" gridrow="5" pageId="9" pageNumber="170">–</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFDABFAF6FDB3" box="[159,1286,550,571]" gridrow="6" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFDABFEFEFDB3" box="[159,270,550,571]" gridcol="0" gridrow="6" pageId="9" pageNumber="170">S1</th>
|
||
<td id="76A85DBAE2050050FE6CFDABFD8AFDB3" box="[461,634,550,571]" gridcol="1" gridrow="6" pageId="9" pageNumber="170">28 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FDABFC4EFDB3" box="[789,958,550,571]" gridcol="2" gridrow="6" pageId="9" pageNumber="170">28 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFDABFAF6FDB3" box="[1117,1286,550,571]" gridcol="3" gridrow="6" pageId="9" pageNumber="170">29 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFDCEFAF6FDD0" box="[159,1286,579,600]" gridrow="7" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFDCEFEFEFDD0" box="[159,270,579,600]" gridcol="0" gridrow="7" pageId="9" pageNumber="170">S2</th>
|
||
<td id="76A85DBAE2050050FE6CFDCEFD8AFDD0" box="[461,634,579,600]" gridcol="1" gridrow="7" pageId="9" pageNumber="170">28 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FDCEFC4EFDD0" box="[789,958,579,600]" gridcol="2" gridrow="7" pageId="9" pageNumber="170">28 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFDCEFAF6FDD0" box="[1117,1286,579,600]" gridcol="3" gridrow="7" pageId="9" pageNumber="170">28 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFDEDFAF6FDFD" box="[159,1286,608,629]" gridrow="8" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFDEDFEFEFDFD" box="[159,270,608,629]" gridcol="0" gridrow="8" pageId="9" pageNumber="170">S3</th>
|
||
<td id="76A85DBAE2050050FE6CFDEDFD8AFDFD" box="[461,634,608,629]" gridcol="1" gridrow="8" pageId="9" pageNumber="170">33 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FDEDFC4EFDFD" box="[789,958,608,629]" gridcol="2" gridrow="8" pageId="9" pageNumber="170">29 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFDEDFAF6FDFD" box="[1117,1286,608,629]" gridcol="3" gridrow="8" pageId="9" pageNumber="170">33 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFDF3FAF6FD1B" box="[159,1286,638,659]" gridrow="9" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFDF3FEFEFD1B" box="[159,270,638,659]" gridcol="0" gridrow="9" pageId="9" pageNumber="170">S4</th>
|
||
<td id="76A85DBAE2050050FE6CFDF3FD8AFD1B" box="[461,634,638,659]" gridcol="1" gridrow="9" pageId="9" pageNumber="170">32 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FDF3FC4EFD1B" box="[789,958,638,659]" gridcol="2" gridrow="9" pageId="9" pageNumber="170">30 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFDF3FAF6FD1B" box="[1117,1286,638,659]" gridcol="3" gridrow="9" pageId="9" pageNumber="170">40 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFD16FAF6FD38" box="[159,1286,667,688]" gridrow="10" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFD16FEFEFD38" box="[159,270,667,688]" gridcol="0" gridrow="10" pageId="9" pageNumber="170">S5</th>
|
||
<td id="76A85DBAE2050050FE6CFD16FD8AFD38" box="[461,634,667,688]" gridcol="1" gridrow="10" pageId="9" pageNumber="170">33 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FD16FC4EFD38" box="[789,958,667,688]" gridcol="2" gridrow="10" pageId="9" pageNumber="170">34 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFD16FAF6FD38" box="[1117,1286,667,688]" gridcol="3" gridrow="10" pageId="9" pageNumber="170">41 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFD35FAF6FD45" box="[159,1286,696,717]" gridrow="11" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFD35FEFEFD45" box="[159,270,696,717]" gridcol="0" gridrow="11" pageId="9" pageNumber="170">S6</th>
|
||
<td id="76A85DBAE2050050FE6CFD35FD8AFD45" box="[461,634,696,717]" gridcol="1" gridrow="11" pageId="9" pageNumber="170">34 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FD35FC4EFD45" box="[789,958,696,717]" gridcol="2" gridrow="11" pageId="9" pageNumber="170">34 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFD35FAF6FD45" box="[1117,1286,696,717]" gridcol="3" gridrow="11" pageId="9" pageNumber="170">44 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFD5BFAF6FD63" box="[159,1286,726,747]" gridrow="12" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFD5BFEFEFD63" box="[159,270,726,747]" gridcol="0" gridrow="12" pageId="9" pageNumber="170">S7</th>
|
||
<td id="76A85DBAE2050050FE6CFD5BFD8AFD63" box="[461,634,726,747]" gridcol="1" gridrow="12" pageId="9" pageNumber="170">38 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FD5BFC4EFD63" box="[789,958,726,747]" gridcol="2" gridrow="12" pageId="9" pageNumber="170">37 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFD5BFAF6FD63" box="[1117,1286,726,747]" gridcol="3" gridrow="12" pageId="9" pageNumber="170">45 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFD7EFAF6FC80" box="[159,1286,755,776]" gridrow="13" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFD7EFEFEFC80" box="[159,270,755,776]" gridcol="0" gridrow="13" pageId="9" pageNumber="170">S8</th>
|
||
<td id="76A85DBAE2050050FE6CFD7EFD8AFC80" box="[461,634,755,776]" gridcol="1" gridrow="13" pageId="9" pageNumber="170">43 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FD7EFC4EFC80" box="[789,958,755,776]" gridcol="2" gridrow="13" pageId="9" pageNumber="170">39 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFD7EFAF6FC80" box="[1117,1286,755,776]" gridcol="3" gridrow="13" pageId="9" pageNumber="170">44 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC9DFAF6FCAD" box="[159,1286,784,805]" gridrow="14" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC9DFEFEFCAD" box="[159,270,784,805]" gridcol="0" gridrow="14" pageId="9" pageNumber="170">S9</th>
|
||
<td id="76A85DBAE2050050FE6CFC9DFD8AFCAD" box="[461,634,784,805]" gridcol="1" gridrow="14" pageId="9" pageNumber="170">40 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FC9DFC4EFCAD" box="[789,958,784,805]" gridcol="2" gridrow="14" pageId="9" pageNumber="170">37 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC9DFAF6FCAD" box="[1117,1286,784,805]" gridcol="3" gridrow="14" pageId="9" pageNumber="170">42 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFCA3FAF6FCCB" box="[159,1286,814,835]" gridrow="15" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFCA3FEFEFCCB" box="[159,270,814,835]" gridcol="0" gridrow="15" pageId="9" pageNumber="170">S10</th>
|
||
<td id="76A85DBAE2050050FE6CFCA3FD8AFCCB" box="[461,634,814,835]" gridcol="1" gridrow="15" pageId="9" pageNumber="170">38 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FCA3FC4EFCCB" box="[789,958,814,835]" gridcol="2" gridrow="15" pageId="9" pageNumber="170">37 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFCA3FAF6FCCB" box="[1117,1286,814,835]" gridcol="3" gridrow="15" pageId="9" pageNumber="170">37 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFCC6FAF6FCE8" box="[159,1286,843,864]" gridrow="16" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFCC6FEFEFCE8" box="[159,270,843,864]" gridcol="0" gridrow="16" pageId="9" pageNumber="170">S11</th>
|
||
<td id="76A85DBAE2050050FE6CFCC6FD8AFCE8" box="[461,634,843,864]" gridcol="1" gridrow="16" pageId="9" pageNumber="170">52 µm</td>
|
||
<td id="76A85DBAE2050050FCB4FCC6FC4EFCE8" box="[789,958,843,864]" gridcol="2" gridrow="16" pageId="9" pageNumber="170">48 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFCC6FAF6FCE8" box="[1117,1286,843,864]" gridcol="3" gridrow="16" pageId="9" pageNumber="170">48 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC0BFAF6FC13" box="[159,1286,902,923]" gridrow="17" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC0BFEFEFC13" box="[159,270,902,923]" gridcol="0" gridrow="17" pageId="9" pageNumber="170">MD4 (ac)</th>
|
||
<td id="76A85DBAE2050050FE6CFC0BFD8AFC13" box="[461,634,902,923]" gridcol="1" gridrow="17" pageId="9" pageNumber="170">broken</td>
|
||
<td id="76A85DBAE2050050FCB4FC0BFC4EFC13" box="[789,958,902,923]" gridcol="2" gridrow="17" pageId="9" pageNumber="170">27 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC0BFAF6FC13" box="[1117,1286,902,923]" gridcol="3" gridrow="17" pageId="9" pageNumber="170">30 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC2EFAF6FC30" box="[159,1286,931,952]" gridrow="18" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC2EFEFEFC30" box="[159,270,931,952]" gridcol="0" gridrow="18" pageId="9" pageNumber="170">MD5 (ac)</th>
|
||
<td id="76A85DBAE2050050FE6CFC2EFD8AFC30" box="[461,634,931,952]" gridcol="1" gridrow="18" pageId="9" pageNumber="170">broken</td>
|
||
<td id="76A85DBAE2050050FCB4FC2EFC4EFC30" box="[789,958,931,952]" gridcol="2" gridrow="18" pageId="9" pageNumber="170">35 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC2EFAF6FC30" box="[1117,1286,931,952]" gridcol="3" gridrow="18" pageId="9" pageNumber="170">42 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC4DFAF6FC5D" box="[159,1286,960,981]" gridrow="19" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC4DFEFEFC5D" box="[159,270,960,981]" gridcol="0" gridrow="19" pageId="9" pageNumber="170">MD6 (ac)</th>
|
||
<td id="76A85DBAE2050050FE6CFC4DFD8AFC5D" box="[461,634,960,981]" gridcol="1" gridrow="19" pageId="9" pageNumber="170">broken</td>
|
||
<td id="76A85DBAE2050050FCB4FC4DFC4EFC5D" box="[789,958,960,981]" gridcol="2" gridrow="19" pageId="9" pageNumber="170">46 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC4DFAF6FC5D" box="[1117,1286,960,981]" gridcol="3" gridrow="19" pageId="9" pageNumber="170">52 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC53FAF6FC7B" box="[159,1286,990,1011]" gridrow="20" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC53FEFEFC7B" box="[159,270,990,1011]" gridcol="0" gridrow="20" pageId="9" pageNumber="170">MD7 (ac)</th>
|
||
<td id="76A85DBAE2050050FE6CFC53FD8AFC7B" box="[461,634,990,1011]" gridcol="1" gridrow="20" pageId="9" pageNumber="170">broken</td>
|
||
<td id="76A85DBAE2050050FCB4FC53FC4EFC7B" box="[789,958,990,1011]" gridcol="2" gridrow="20" pageId="9" pageNumber="170">57 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC53FAF6FC7B" box="[1117,1286,990,1011]" gridcol="3" gridrow="20" pageId="9" pageNumber="170">61 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3EFC76FAF6FB98" box="[159,1286,1019,1040]" gridrow="21" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3EFC76FEFEFB98" box="[159,270,1019,1040]" gridcol="0" gridrow="21" pageId="9" pageNumber="170">MD8 (ac)</th>
|
||
<td id="76A85DBAE2050050FE6CFC76FD8AFB98" box="[461,634,1019,1040]" gridcol="1" gridrow="21" pageId="9" pageNumber="170">broken</td>
|
||
<td id="76A85DBAE2050050FCB4FC76FC4EFB98" box="[789,958,1019,1040]" gridcol="2" gridrow="21" pageId="9" pageNumber="170">68 µm</td>
|
||
<td id="76A85DBAE2050050FBFCFC76FAF6FB98" box="[1117,1286,1019,1040]" gridcol="3" gridrow="21" pageId="9" pageNumber="170">74 µm</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF36660CE205FFA6FF36FAB6FA67FA69" pageId="9" pageNumber="170" startId="9.[151,239,1339,1363]">
|
||
<paragraph id="8BF63684E205FFA6FF36FAB6FA67FA69" blockId="9.[151,1437,1339,1505]" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FF36FAB6FEFFFADB" bold="true" box="[151,271,1339,1363]" pageId="9" pageNumber="170">TABLE 4.</emphasis>
|
||
Summary of nature and location of sensory spots, glandular cell outlets and spines arranged by series in
|
||
<taxonomicName id="4C494D07E205FFA6FF36FAD3FE7BFAFD" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[151,395,1374,1397]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="9" pageNumber="170" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E205FFA6FF36FAD3FE7BFAFD" box="[151,395,1374,1397]" italics="true" pageId="9" pageNumber="170">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E205FFA6FE35FAD0FE19FAFD" bold="true" box="[404,489,1373,1397]" pageId="9" pageNumber="170">
|
||
<taxonomicNameLabel id="A20E57EDE205FFA6FE35FAD0FE19FAFD" box="[404,489,1373,1397]" pageId="9" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the Korean South Sea. Abbreviations: LA: Lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; (f): female condition of sexual dimorphic character; gco1/2: glandular cell outlet type 1/2; ltas: lateral terminal accessory spine; lts: lateral terminal spine; si: sieve plate; ss: sensory spot; tu: tubule.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E205FFA6FF3CFA79FBC4F849" pageId="9" pageNumber="170">
|
||
<table id="F949C424E2050050FF3CFA79FA94F849" box="[157,1380,1524,1985]" gridcols="12" gridrows="12" pageId="9" pageNumber="170">
|
||
<tr id="357934C6E2050050FF3CFA79FA94F9AE" box="[157,1380,1524,1574]" gridrow="0" pageId="9" pageNumber="170">
|
||
<th id="76A85DBAE2050050FF3CFA79FF05F9AE" box="[157,245,1524,1574]" gridcol="0" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FF3CFA79FF05F9AE" bold="true" box="[157,245,1524,1574]" pageId="9" pageNumber="170">Position Segment</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FEACFA79FE80F9AE" box="[269,368,1524,1574]" gridcol="1" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FEACFA79FEC5F981" bold="true" box="[269,309,1524,1545]" pageId="9" pageNumber="170">MD</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FE30FA79FE28F9AE" box="[401,472,1524,1574]" gridcol="2" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FE30FA79FE41F981" bold="true" box="[401,433,1524,1545]" pageId="9" pageNumber="170">PD</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FE4CFA79FDECF9AE" box="[493,540,1524,1574]" gridcol="3" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FE4CFA79FDFBF981" bold="true" box="[493,523,1524,1545]" pageId="9" pageNumber="170">SD</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FDD7FA79FD27F9AE" box="[630,727,1524,1574]" gridcol="4" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FDD7FA79FD67F981" bold="true" box="[630,663,1524,1545]" pageId="9" pageNumber="170">LD</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FCA5FA79FCC3F9AE" box="[772,819,1524,1574]" gridcol="5" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FCA5FA79FCDAF981" bold="true" box="[772,810,1524,1545]" pageId="9" pageNumber="170">ML</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FCC2FA79FC62F9AE" box="[867,914,1524,1574]" gridcol="6" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FCC2FA79FC70F981" bold="true" box="[867,896,1524,1545]" pageId="9" pageNumber="170">SL</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FC61FA79FBF4F9AE" box="[960,1028,1524,1574]" gridcol="7" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FC61FA79FC11F981" bold="true" box="[960,993,1524,1545]" pageId="9" pageNumber="170">LA</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FBBFFA79FBCDF9AE" box="[1054,1085,1524,1574]" gridcol="8" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FBBFFA79FBCDF981" bold="true" box="[1054,1085,1524,1545]" pageId="9" pageNumber="170">LV</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FBF7FA79FB75F9AE" box="[1110,1157,1524,1574]" gridcol="9" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FBF7FA79FB87F981" bold="true" box="[1110,1143,1524,1545]" pageId="9" pageNumber="170">VL</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FB67FA79FAFDF9AE" box="[1222,1293,1524,1574]" gridcol="10" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FB67FA79FB1EF981" bold="true" box="[1222,1262,1524,1545]" pageId="9" pageNumber="170">VM</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE2050050FA97FA79FA94F9AE" box="[1334,1380,1524,1574]" gridcol="11" gridrow="0" pageId="9" pageNumber="170">
|
||
<emphasis id="B93DEA96E205FFA6FA97FA79FAA6F981" bold="true" box="[1334,1366,1524,1545]" pageId="9" pageNumber="170">PV</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF9BAFA94F9C4" box="[157,1380,1591,1612]" gridrow="1" pageId="9" pageNumber="170" rowspan-1="1" rowspan-11="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF9BAFF05F9C4" box="[157,245,1591,1612]" gridcol="0" gridrow="1" pageId="9" pageNumber="170">1</th>
|
||
<td id="76A85DBAE2050050FE4CF9BAFDECF9C4" box="[493,540,1591,1612]" gridcol="3" gridrow="1" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FDD7F9BAFD27F9C4" box="[630,727,1591,1612]" gridcol="4" gridrow="1" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FB67F9BAFAFDF9C4" box="[1222,1293,1591,1612]" gridcol="10" gridrow="1" pageId="9" pageNumber="170">ss</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF9D1FA94F9F9" box="[157,1380,1628,1649]" gridrow="2" pageId="9" pageNumber="170" rowspan-11="1" rowspan-2="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="76A85DBAE2050050FF3CF9D1FF05F9F9" box="[157,245,1628,1649]" gridcol="0" gridrow="2" pageId="9" pageNumber="170">2</th>
|
||
<td id="76A85DBAE2050050FEACF9D1FE80F9F9" box="[269,368,1628,1649]" gridcol="1" gridrow="2" pageId="9" pageNumber="170">gco1,ss</td>
|
||
<td id="76A85DBAE2050050FE4CF9D1FDECF9F9" box="[493,540,1628,1649]" gridcol="3" gridrow="2" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FDD7F9D1FD27F9F9" box="[630,727,1628,1649]" gridcol="4" gridrow="2" pageId="9" pageNumber="170">ss,gco2,ss</td>
|
||
<td id="76A85DBAE2050050FCC2F9D1FC62F9F9" box="[867,914,1628,1649]" gridcol="6" gridrow="2" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FBF7F9D1FB75F9F9" box="[1110,1157,1628,1649]" gridcol="9" gridrow="2" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FB67F9D1FAFDF9F9" box="[1222,1293,1628,1649]" gridcol="10" gridrow="2" pageId="9" pageNumber="170">ss</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF90CFA94F91E" box="[157,1380,1665,1686]" gridrow="3" pageId="9" pageNumber="170" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF90CFF05F91E" box="[157,245,1665,1686]" gridcol="0" gridrow="3" pageId="9" pageNumber="170">3</th>
|
||
<td id="76A85DBAE2050050FEACF90CFE80F91E" box="[269,368,1665,1686]" gridcol="1" gridrow="3" pageId="9" pageNumber="170">gco1</td>
|
||
<td id="76A85DBAE2050050FE4CF90CFDECF91E" box="[493,540,1665,1686]" gridcol="3" gridrow="3" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FCA5F90CFCC3F91E" box="[772,819,1665,1686]" gridcol="5" gridrow="3" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FB67F90CFAFDF91E" box="[1222,1293,1665,1686]" gridcol="10" gridrow="3" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF92AFA94F934" box="[157,1380,1703,1724]" gridrow="4" pageId="9" pageNumber="170" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF92AFF05F934" box="[157,245,1703,1724]" gridcol="0" gridrow="4" pageId="9" pageNumber="170">4</th>
|
||
<td id="76A85DBAE2050050FEACF92AFE80F934" box="[269,368,1703,1724]" gridcol="1" gridrow="4" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FE4CF92AFDECF934" box="[493,540,1703,1724]" gridcol="3" gridrow="4" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FB67F92AFAFDF934" box="[1222,1293,1703,1724]" gridcol="10" gridrow="4" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF941FA94F969" box="[157,1380,1740,1761]" gridrow="5" pageId="9" pageNumber="170" rowspan-11="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF941FF05F969" box="[157,245,1740,1761]" gridcol="0" gridrow="5" pageId="9" pageNumber="170">5</th>
|
||
<td id="76A85DBAE2050050FEACF941FE80F969" box="[269,368,1740,1761]" gridcol="1" gridrow="5" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FE30F941FE28F969" box="[401,472,1740,1761]" gridcol="2" gridrow="5" pageId="9" pageNumber="170">gco1</td>
|
||
<td id="76A85DBAE2050050FE4CF941FDECF969" box="[493,540,1740,1761]" gridcol="3" gridrow="5" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FCA5F941FCC3F969" box="[772,819,1740,1761]" gridcol="5" gridrow="5" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FBBFF941FBCDF969" box="[1054,1085,1740,1761]" gridcol="8" gridrow="5" pageId="9" pageNumber="170">tu</td>
|
||
<td id="76A85DBAE2050050FB67F941FAFDF969" box="[1222,1293,1740,1761]" gridcol="10" gridrow="5" pageId="9" pageNumber="170">ss,gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF97CFA94F88E" box="[157,1380,1777,1798]" gridrow="6" pageId="9" pageNumber="170" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF97CFF05F88E" box="[157,245,1777,1798]" gridcol="0" gridrow="6" pageId="9" pageNumber="170">6</th>
|
||
<td id="76A85DBAE2050050FEACF97CFE80F88E" box="[269,368,1777,1798]" gridcol="1" gridrow="6" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FE30F97CFDECF88E" box="[401,540,1777,1798]" colspan="2" colspanRight="1" gridcol="2" gridrow="6" pageId="9" pageNumber="170">gco1.ss ss</td>
|
||
<td id="76A85DBAE2050050FCA5F97CFCC3F88E" box="[772,819,1777,1798]" gridcol="5" gridrow="6" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FBBFF97CFBCDF88E" box="[1054,1085,1777,1798]" gridcol="8" gridrow="6" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FB67F97CFAFDF88E" box="[1222,1293,1777,1798]" gridcol="10" gridrow="6" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FA97F97CFA94F88E" box="[1334,1380,1777,1798]" gridcol="11" gridrow="6" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF89AFA94F8A4" box="[157,1380,1815,1836]" gridrow="7" pageId="9" pageNumber="170" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF89AFF05F8A4" box="[157,245,1815,1836]" gridcol="0" gridrow="7" pageId="9" pageNumber="170">7</th>
|
||
<td id="76A85DBAE2050050FEACF89AFE80F8A4" box="[269,368,1815,1836]" gridcol="1" gridrow="7" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FE30F89AFE28F8A4" box="[401,472,1815,1836]" gridcol="2" gridrow="7" pageId="9" pageNumber="170">gco1,ss</td>
|
||
<td id="76A85DBAE2050050FE4CF89AFDECF8A4" box="[493,540,1815,1836]" gridcol="3" gridrow="7" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FCA5F89AFCC3F8A4" box="[772,819,1815,1836]" gridcol="5" gridrow="7" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FBBFF89AFBCDF8A4" box="[1054,1085,1815,1836]" gridcol="8" gridrow="7" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FB67F89AFAFDF8A4" box="[1222,1293,1815,1836]" gridcol="10" gridrow="7" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FA97F89AFA94F8A4" box="[1334,1380,1815,1836]" gridcol="11" gridrow="7" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF8B1FA94F8D9" box="[157,1380,1852,1873]" gridrow="8" pageId="9" pageNumber="170" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF8B1FF05F8D9" box="[157,245,1852,1873]" gridcol="0" gridrow="8" pageId="9" pageNumber="170">8</th>
|
||
<td id="76A85DBAE2050050FEACF8B1FE80F8D9" box="[269,368,1852,1873]" gridcol="1" gridrow="8" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FE30F8B1FE28F8D9" box="[401,472,1852,1873]" gridcol="2" gridrow="8" pageId="9" pageNumber="170">gco1,ss</td>
|
||
<td id="76A85DBAE2050050FE4CF8B1FDECF8D9" box="[493,540,1852,1873]" gridcol="3" gridrow="8" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FCC2F8B1FC62F8D9" box="[867,914,1852,1873]" gridcol="6" gridrow="8" pageId="9" pageNumber="170">gco2</td>
|
||
<td id="76A85DBAE2050050FBBFF8B1FBCDF8D9" box="[1054,1085,1852,1873]" gridcol="8" gridrow="8" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FA97F8B1FA94F8D9" box="[1334,1380,1852,1873]" gridcol="11" gridrow="8" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF8ECFA94F8FE" box="[157,1380,1889,1910]" gridrow="9" pageId="9" pageNumber="170" rowspan-1="1" rowspan-10="1" rowspan-4="1" rowspan-6="1">
|
||
<th id="76A85DBAE2050050FF3CF8ECFF05F8FE" box="[157,245,1889,1910]" gridcol="0" gridrow="9" pageId="9" pageNumber="170">9</th>
|
||
<td id="76A85DBAE2050050FE30F8ECFE28F8FE" box="[401,472,1889,1910]" gridcol="2" gridrow="9" pageId="9" pageNumber="170">gco1,ss</td>
|
||
<td id="76A85DBAE2050050FE4CF8ECFDECF8FE" box="[493,540,1889,1910]" gridcol="3" gridrow="9" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FCA5F8ECFCC3F8FE" box="[772,819,1889,1910]" gridcol="5" gridrow="9" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FC61F8ECFBF4F8FE" box="[960,1028,1889,1910]" gridcol="7" gridrow="9" pageId="9" pageNumber="170">si</td>
|
||
<td id="76A85DBAE2050050FBBFF8ECFBCDF8FE" box="[1054,1085,1889,1910]" gridcol="8" gridrow="9" pageId="9" pageNumber="170">ac</td>
|
||
<td id="76A85DBAE2050050FBF7F8ECFB75F8FE" box="[1110,1157,1889,1910]" gridcol="9" gridrow="9" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FA97F8ECFA94F8FE" box="[1334,1380,1889,1910]" gridcol="11" gridrow="9" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF80AFA94F814" box="[157,1380,1927,1948]" gridrow="10" pageId="9" pageNumber="170" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="76A85DBAE2050050FF3CF80AFF05F814" box="[157,245,1927,1948]" gridcol="0" gridrow="10" pageId="9" pageNumber="170">10</th>
|
||
<td id="76A85DBAE2050050FEACF80AFE80F814" box="[269,368,1927,1948]" gridcol="1" gridrow="10" pageId="9" pageNumber="170">gco1,gco1</td>
|
||
<td id="76A85DBAE2050050FE4CF80AFDECF814" box="[493,540,1927,1948]" gridcol="3" gridrow="10" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FDD7F80AFD27F814" box="[630,727,1927,1948]" gridcol="4" gridrow="10" pageId="9" pageNumber="170">tu?</td>
|
||
<td id="76A85DBAE2050050FBF7F80AFB75F814" box="[1110,1157,1927,1948]" gridcol="9" gridrow="10" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FA97F80AFA94F814" box="[1334,1380,1927,1948]" gridcol="11" gridrow="10" pageId="9" pageNumber="170">gco1</td>
|
||
</tr>
|
||
<tr id="357934C6E2050050FF3CF821FA94F849" box="[157,1380,1964,1985]" gridrow="11" pageId="9" pageNumber="170" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="76A85DBAE2050050FF3CF821FF05F849" box="[157,245,1964,1985]" gridcol="0" gridrow="11" pageId="9" pageNumber="170">11</th>
|
||
<td id="76A85DBAE2050050FE30F821FE28F849" box="[401,472,1964,1985]" gridcol="2" gridrow="11" pageId="9" pageNumber="170">ss</td>
|
||
<td id="76A85DBAE2050050FC61F821FBF4F849" box="[960,1028,1964,1985]" gridcol="7" gridrow="11" pageId="9" pageNumber="170">ltas(f?)</td>
|
||
<td id="76A85DBAE2050050FBBFF821FBCDF849" box="[1054,1085,1964,1985]" gridcol="8" gridrow="11" pageId="9" pageNumber="170">lts</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF36660CE206FFA5FF36F8C4FDDCF85C" ID-DOI="http://doi.org/10.5281/zenodo.5252308" ID-Zenodo-Dep="5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="10" pageNumber="171" startId="10.[151,244,1865,1886]" targetBox="[217,1376,206,1837]" targetPageId="10">
|
||
<paragraph id="8BF63684E206FFA5FF36F8C4FDDCF85C" blockId="10.[151,1437,1865,2004]" pageId="10" pageNumber="171">
|
||
<emphasis id="B93DEA96E206FFA5FF36F8C4FEFDF8D6" bold="true" box="[151,269,1865,1886]" pageId="10" pageNumber="171">FIGURE 4.</emphasis>
|
||
Line art illustrations showing general female habitus in
|
||
<taxonomicName id="4C494D07E206FFA5FC8FF8C4FBE0F8D6" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[814,1040,1865,1886]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="171" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E206FFA5FC8FF8C4FBE0F8D6" box="[814,1040,1865,1886]" italics="true" pageId="10" pageNumber="171">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20E57EDE206FFA5FBB7F8C4FB90F8D6" box="[1046,1120,1865,1886]" pageId="10" pageNumber="171" rank="species">sp. nov</taxonomicNameLabel>
|
||
: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E207FFA4FF66FF1AFEB8FD9F" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 2 consists of one complete cuticular ring. Pairs of conspicuous glandular cell outlets
|
||
<typeStatus id="54F28826E207FFA4FB42FF15FAE3FF38" box="[1251,1299,152,176]" pageId="11" pageNumber="172">type</typeStatus>
|
||
2 present in subdorsal, laterodorsal, sublateral and ventrolateral positions (
|
||
<figureCitation id="13722A01E207FFA4FCF0FF31FC20FF5C" box="[849,976,188,212]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FC7DFF31FBD1FF5C" box="[988,1057,188,212]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FB8FFF31FBA3FF5C" box="[1070,1107,188,212]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="11" pageNumber="172">6D</figureCitation>
|
||
). Sensory spots present as an unpaired one in middorsal position, two pairs in laterodorsal position on each side of the gco2 (
|
||
<figureCitation id="13722A01E207FFA4FB72FF52FADEFF7F" box="[1235,1326,223,247]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="11" pageNumber="172">Fig. 6D</figureCitation>
|
||
), and one pair in ventromedial positions; sensory spots on this and following eight segments are minute, consisting of a few papillae forming a circle around a central pore. An unpaired middorsal pore field (i.e., glandular cell outlet
|
||
<typeStatus id="54F28826E207FFA4FAF7FEA5FA76FEC8" box="[1366,1414,296,320]" pageId="11" pageNumber="172">type</typeStatus>
|
||
1 (gco1)) is present in middorsal position (
|
||
<figureCitation id="13722A01E207FFA4FDD9FEC1FD26FEEC" box="[632,726,332,356]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5A</figureCitation>
|
||
). Cuticular hairs on this and the following segments emerge through slit-like perforation sites; hairs on dorsal and lateral sides are scattered in a median belt that is limited posteriorly by the IJ-line; no hairs present on ventral side. A secondary pectinate fringe was not observed on this or any of the following segments. Posterior segment margin with regular pectinate fringe on dorsal and ventral sides, whereas the fringes on the lateral sides are composed of well-spaced long and narrow fringe tips, with much shorter tips in between.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66FDA9FF0CFC9C" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 3 and following seven segments, down to segment 10, consist of one tergal and two sternal plates (
|
||
<figureCitation id="13722A01E207FFA4FF3EFDCAFF0BFDD7" box="[159,251,583,607]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Fig. 4B</figureCitation>
|
||
). On all segments with this composition, pachycycli are well-developed along anterior margins of the sternal plates, and along anterior 1/3 of tergosternal and midsternal junctions. Segment with sensory spots in subdorsal and midlateral positions, and unpaired middorsal and paired ventromedial pore fields (gco1). Cuticular hairs scattered in a median belt around the tergal plate and onto the ventrolateral areas of the sternal plates; no cuticular hairs in ventromedial and paraventral areas. Posterior segment margin as lateral margin on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66FC92FADDFCD4" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 4 with middorsal spine (
|
||
<figureCitation id="13722A01E207FFA4FDE1FC92FD6BFCBF" box="[576,667,799,823]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Fig. 4A</figureCitation>
|
||
). One pair of sensory spots present in subdorsal position. Pore fields (gco1) in ventromedial positions (
|
||
<figureCitation id="13722A01E207FFA4FDB2FCC9FD9BFCD4" box="[531,619,836,860]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Fig. 4B</figureCitation>
|
||
). Cuticular hairs and pectinate fringe as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66FCEAFE81FBBC" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 5 with middorsal spine and lateroventral tubules (length = 13 µm, estimated with SEM) (
|
||
<figureCitation id="13722A01E207FFA4FAB1FCEAFA60FCF7" box="[1296,1424,871,895]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4A–B</figureCitation>
|
||
). Paired paradorsal pore fields (gco1) present, anterior to the attachment point of the middorsal spine (
|
||
<figureCitation id="13722A01E207FFA4FAA0FC01FAAFFC2C" box="[1281,1375,908,932]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FACAFC01FA7EFC2C" box="[1387,1422,908,932]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
|
||
); one additional pair present in ventromedial position. One pair of conspicuous glandular cell outlets
|
||
<typeStatus id="54F28826E207FFA4FB5CFC3DFADDFC40" box="[1277,1325,944,968]" pageId="11" pageNumber="172">type</typeStatus>
|
||
2 present in midlateral positions (
|
||
<figureCitation id="13722A01E207FFA4FE08FC59FDF7FC64" box="[425,519,980,1004]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FDB6FC59FDCAFC64" box="[535,570,980,1004]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
|
||
). Sensory spots present in subdorsal and ventromedial positions; subdorsal sensory spots with short, tube-shaped subcuticular structure (
|
||
<figureCitation id="13722A01E207FFA4FCF0FC7AFC5DFB87" box="[849,941,1015,1039]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5D</figureCitation>
|
||
). Cuticular hairs and pectinate fringe as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66FBB2FE81FAB8" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 6 with middorsal spine and lateroventral spines (
|
||
<figureCitation id="13722A01E207FFA4FCD7FBB2FC09FBDF" box="[886,1017,1087,1111]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FBA8FBB2FBDCFBDF" box="[1033,1068,1087,1111]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FB9CFBB2FB90FBDF" box="[1085,1120,1087,1111]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="11" pageNumber="172">6E</figureCitation>
|
||
). Paired pore fields (gco1) present slightly anterior to attachment point of the middorsal spine (
|
||
<figureCitation id="13722A01E207FFA4FC66FBE9FBD4FBF4" box="[967,1060,1124,1148]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5C</figureCitation>
|
||
); one additional pair present in paraventral position. Sensory spots present in paradorsal, subdorsal, midlateral (
|
||
<figureCitation id="13722A01E207FFA4FBFCFB05FB4AFB28" box="[1117,1210,1160,1184]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5C</figureCitation>
|
||
) and ventromedial positions; paradorsal ones are very close to the subdorsal area, and do not readily appear as perispinal sensory spots; subdorsal sensory spots with tube-shaped subcuticular structure (
|
||
<figureCitation id="13722A01E207FFA4FC4CFB5DFBBDFB60" box="[1005,1101,1232,1256]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5D</figureCitation>
|
||
); ventromedial ones located slightly closer to the midsternal line than those on the preceding segment. Cuticular hairs and pectinate fringe as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66FAB1FCB4F980" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 7 with middorsal spine and lateroventral spines. Paired pore fields (gco1) present slightly anterior to attachment point of the middorsal spine (
|
||
<figureCitation id="13722A01E207FFA4FDC0FAEDFD4EFAF0" box="[609,702,1376,1400]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FD68FAEDFD1DFAF0" box="[713,749,1376,1400]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
|
||
); one additional pair present in paraventral position. One pair of conspicuous glandular cell outlets
|
||
<typeStatus id="54F28826E207FFA4FD97FA09FD96FA14" box="[566,614,1412,1436]" pageId="11" pageNumber="172">type</typeStatus>
|
||
2 present in midlateral positions (
|
||
<figureCitation id="13722A01E207FFA4FC41FA09FBCAFA14" box="[992,1082,1412,1436]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FBE7FA09FB99FA14" box="[1094,1129,1412,1436]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E207FFA4FBD5FA09FB67FA14" box="[1140,1175,1412,1436]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="11" pageNumber="172">6E</figureCitation>
|
||
). Sensory spots present in paradorsal (as on preceding segment), subdorsal and ventromedial positions; subdorsal sensory spots with tubeshaped subcuticular structure; ventromedial ones located slightly more lateral than those on the preceding segment. Cuticular hairs and pectinate fringe as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66F999FBB4F934" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 8 with middorsal spine and lateroventral spines. Paired pore fields (gco1) present slightly anterior to attachment point of the middorsal spine; one additional pair present in paraventral position. One pair of conspicuous glandular cell outlets
|
||
<typeStatus id="54F28826E207FFA4FD92F9D1FD93F9FC" box="[563,611,1628,1652]" pageId="11" pageNumber="172">type</typeStatus>
|
||
2 present in sublateral positions (
|
||
<figureCitation id="13722A01E207FFA4FBA0F9D1FB90F9FC" box="[1025,1120,1628,1652]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="11" pageNumber="172">Fig. 4B</figureCitation>
|
||
). Sensory spots present in paradorsal (as on preceding segment) and subdorsal positions; subdorsal sensory spots with tube-shaped subcuticular structure. Cuticular hairs and pectinate fringe as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E207FFA4FF66F945FBA5F830" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="172">
|
||
Segment 9 without middorsal spine, but with lateroventral spines. Paired pore fields (gco1) present in paradorsal and paraventral positions; paraventral pore fields closer to midsternal line than those on preceding segments. Sensory spots present in paradorsal (as on preceding segment), subdorsal, midlateral and ventromedial positions; subcuticular structures associated with subdorsal sensory spots are anteriorly expanded, giving them a funnel-shaped rather than tube-shaped appearance (
|
||
<figureCitation id="13722A01E207FFA4FD5FF8D5FCACF8F8" box="[766,860,1880,1904]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="11" pageNumber="172">Fig. 5E</figureCitation>
|
||
). Small, rounded sieve plates present in lateral accessory positions. Cuticular hairs as on preceding segment, but almost lacking in mid- and paradorsal areas. Pectinate fringe with uniform, well-developed fringe tips around the segment margin.
|
||
</paragraph>
|
||
<caption id="DF36660CE200FFA3FF36F955FDE7F851" ID-DOI="http://doi.org/10.5281/zenodo.5252314" ID-Zenodo-Dep="5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="12" pageNumber="173" startId="12.[151,244,1752,1773]" targetBox="[220,1376,179,1732]" targetPageId="12">
|
||
<paragraph id="8BF63684E200FFA3FF36F955FDE7F851" blockId="12.[151,1436,1752,2009]" pageId="12" pageNumber="173">
|
||
<emphasis id="B93DEA96E200FFA3FF36F955FEFDF965" bold="true" box="[151,269,1752,1773]" pageId="12" pageNumber="173">FIGURE 5.</emphasis>
|
||
Light micrographs showing details in neck and trunk morphology of the female holotype of
|
||
<taxonomicName id="4C494D07E200FFA3FB37F954FA89F966" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[1174,1401,1753,1774]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="12" pageNumber="173" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E200FFA3FB37F954FA89F966" box="[1174,1401,1753,1774]" italics="true" pageId="12" pageNumber="173">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E200FFA3FADEF955FF30F883" bold="true" pageId="12" pageNumber="173">
|
||
<taxonomicNameLabel id="A20E57EDE200FFA3FADEF955FF30F883" pageId="12" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF36660CE201FFA2FF36F92DFF2FF828" ID-DOI="http://doi.org/10.5281/zenodo.5252318" ID-Zenodo-Dep="5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="13" pageNumber="174" startId="13.[151,244,1696,1717]" targetBox="[186,1398,199,1673]" targetPageId="13">
|
||
<paragraph id="8BF63684E201FFA2FF36F92DFF2FF828" blockId="13.[151,1437,1696,1952]" pageId="13" pageNumber="174">
|
||
<emphasis id="B93DEA96E201FFA2FF36F92DFEFFF93D" bold="true" box="[151,271,1696,1717]" pageId="13" pageNumber="174">FIGURE 6.</emphasis>
|
||
Scanning electron micrographs showing overviews and details in head and trunk morphology of female
|
||
<taxonomicName id="4C494D07E201FFA2FA84F92CFF0BF95B" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="13" pageNumber="174" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E201FFA2FA84F92CFF0BF95B" italics="true" pageId="13" pageNumber="174">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E201FFA2FEA2F933FEA1F95B" bold="true" box="[259,337,1726,1747]" pageId="13" pageNumber="174">
|
||
<taxonomicNameLabel id="A20E57EDE201FFA2FEA2F933FEA1F95B" box="[259,337,1726,1747]" pageId="13" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E202FFA1FF66FF1AFE2DFE47" blockId="14.[151,1436,151,1616]" pageId="14" pageNumber="175">
|
||
Segment 10 without spines. Two unpaired, middorsal pore fields (gco1) (
|
||
<figureCitation id="13722A01E202FFA1FBA1FF1AFBA5FF27" box="[1024,1109,151,175]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="14" pageNumber="175">Fig. 5E</figureCitation>
|
||
) and paired paraventral ones present; paraventral pore fields closer to midsternal line than those on preceding segments. Sensory spots with funnel-shaped subcuticular structure present in subdorsal positions; a regular pair of sensory spots present in ventrolateral position. A pair of fringed tufts extends slightly beyond the posterior segment margin in the laterodorsal position; the structures could be very short tubules with fringed tips, but this is uncertain (structure marked as “ldt?” on
|
||
<figureCitation id="13722A01E202FFA1FE03FEC1FDF7FEEC" box="[418,519,332,356]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FDBDFEC1FDCDFEEC" box="[540,573,332,356]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">6F</figureCitation>
|
||
). Cuticular hairs scattered in middorsal, laterodorsal and lateroventral/ ventrolateral clusters. Posterior segment margin with pectinate fringe; fringe tips short along dorsal margin, and along the lateral ones; paraventrally the sternal plates and the fringe tips are elongated, almost reaching the margin of segment 11 (
|
||
<figureCitation id="13722A01E202FFA1FEE2FE3AFE6EFE47" box="[323,414,439,463]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FE0AFE3AFE20FE47" box="[427,464,439,463]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">6H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E202FFA1FF66FE51FC38FD2F" blockId="14.[151,1436,151,1616]" pageId="14" pageNumber="175">
|
||
Segment 11 is composed of two tergal and two sternal plates (
|
||
<figureCitation id="13722A01E202FFA1FCDEFE51FC0DFE7C" box="[895,1021,476,500]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FBA8FE51FBDAFE7C" box="[1033,1066,476,500]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="14" pageNumber="175">5G</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FB97FE51FBA7FE7C" box="[1078,1111,476,500]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">6F</figureCitation>
|
||
). Lateral terminal spines and lateral terminal accessory spines present (
|
||
<figureCitation id="13722A01E202FFA1FDCCFE72FD1CFD9F" box="[621,748,511,535]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FD59FE72FCE9FD9F" box="[760,793,511,535]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="14" pageNumber="175">5F</figureCitation>
|
||
); the latter is probably a sexually dimorphic female trait, but male specimens were unavailable to confirm this. Sensory spots present in paradorsal position (
|
||
<figureCitation id="13722A01E202FFA1FA94FDA9FA7FFDB4" box="[1333,1423,548,572]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">Fig. 6F</figureCitation>
|
||
). Cuticular hairs not present. Tergal extensions extremely elongated and posteriorly projecting (
|
||
<figureCitation id="13722A01E202FFA1FB42FDCAFAB5FDD7" box="[1251,1349,583,607]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FAF4FDCAFA67FDD7" box="[1365,1431,583,607]" captionStart="FIGURE 5" captionStartId="12.[151,244,1752,1773]" captionTargetBox="[220,1376,179,1732]" captionTargetId="figure-193@12.[212,1385,169,1739]" captionTargetPageId="12" captionText="FIGURE 5. Light micrographs showing details in neck and trunk morphology of the female holotype of Echinoderes cernunnos sp. nov., NIBR Acc. No. INBRIV0000245082: A, neck and segments 1 and 2, dorsal view; B, neck and segments 1 and 2, ventral view; C, Segments 5 to 7, middorsal to midlateral regions on right half of tergal plates; D, segments 5 and 6, close-up of middorsal to subdorsal regions on right half of tergal plates; E, segments 9 and 10, close-up of middorsal and subdorsal areas; F, segment 11, focused on left side tergal extension and terminal spines; G, segment 11, focused on midtergal junction; H, segment 11, focused on right side tergal extension. Abbreviations: gco I/II, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; sd (tu/fu), subdorsal sensory spot (with tube-shaped/funnel-shaped subcuticular structure); te, tergal extension; trp, trichoscalid plate. Digits after the labels refer to the segment numbers." figureDoi="http://doi.org/10.5281/zenodo.5252314" httpUri="https://zenodo.org/record/5252314/files/figure.png" pageId="14" pageNumber="175">5F–H</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FF36FDE1FF10FD0C" box="[151,224,620,644]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">6G–H</figureCitation>
|
||
), measuring 38 µm and constituting 70% of the total segment length; sternal extensions short and rounded with fringed margins and a few longer filiform extensions (
|
||
<figureCitation id="13722A01E202FFA1FC8FFD02FC79FD2F" box="[814,905,655,679]" captionStart="FIGURE 4" captionStartId="10.[151,244,1865,1886]" captionTargetBox="[217,1376,206,1837]" captionTargetId="figure-100@10.[207,1379,193,1844]" captionTargetPageId="10" captionText="FIGURE 4. Line art illustrations showing general female habitus in Echinoderes cernunnos sp. nov: A, dorsal view; B, ventral view. Abbreviations: gco I/II, glandular cell outlet type 1/2; ldt?, laterodorsal tubule?; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mdp, middorsal placid; mds, middorsal spine; ml, midlateral sensory spot; mtj, midtergal junction; mvp, midventral placid; s, segment followed by segment number; sd, subdorsal sensory spot; te, tergal extension; vm, ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.5252308" httpUri="https://zenodo.org/record/5252308/files/figure.png" pageId="14" pageNumber="175">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E202FFA1FC34FD02FC4BFD2F" box="[917,955,655,679]" captionStart="FIGURE 6" captionStartId="13.[151,244,1696,1717]" captionTargetBox="[186,1398,199,1673]" captionTargetId="figure-183@13.[185,1401,193,1675]" captionTargetPageId="13" captionText="FIGURE 6. Scanning electron micrographs showing overviews and details in head and trunk morphology of female Echinoderes cernunnos sp. nov.: A, lateral overview of whole specimen; B, detail of head, showing partly extended mouth cone and outer oral styles; C, detail of head showing introvert section 8; D, segments 1 and 2, dorsolateral view; E, segments 6 and 7, midlateral view; F, segment 11, subdorsal view; G, segment 11, lateral view; H, segment 11, ventral view. Abbreviations: ff, free flap, fs, fringe of style basis; gco II, glandular cell outlet type 2; go, gonopore; ld, laterodorsal sensory spot; ldt?, laterodorsal tubule?; lvs, lateroventral spine; lvt, lateroventral tubule; ml, midlateral sensory spot; mtj, midtergal junction; oos, outer oral styles; pd, paradorsal sensory spot; sc, scalids; se, sternal extension; sp, spinoscalids; tb, transverse band of ridges; te, tergal extension; tr, trichoscalid; trp, trichoscalid plate. Digits after labels refer to the introvert ring numbers on C, otherwise to segment number. Lambda symbols Λ mark attachment point of scalids." figureDoi="http://doi.org/10.5281/zenodo.5252318" httpUri="https://zenodo.org/record/5252318/files/figure.png" pageId="14" pageNumber="175">6H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E202FFA1FF66FD39FF3EFB87" blockId="14.[151,1436,151,1616]" pageId="14" pageNumber="175">
|
||
<emphasis id="B93DEA96E202FFA1FF66FD39FD39FD44" bold="true" box="[199,713,691,716]" inLineHeading="true" pageId="14" pageNumber="175" reason="1">Notes on diagnostic features and affinities.</emphasis>
|
||
Even though
|
||
<taxonomicName id="4C494D07E202FFA1FCCAFD39FB83FD44" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[875,1139,692,716]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FCCAFD39FB83FD44" box="[875,1139,692,716]" italics="true" pageId="14" pageNumber="175">Echinoderes cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FBDDFD39FB28FD44" bold="true" box="[1148,1240,692,716]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FBDDFD39FB28FD44" box="[1148,1240,692,716]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
displays a rather classic spine pattern, with five middorsal spines and lateroventral spines/tubules on segments 5 to 9, the species possesses several unique traits that make it impossible to confuse with any other taxon. Also the lack of ventrolateral tubules is uncommon, and only shared with a minority of its congeners. The most prominent characteristic, however, is its extremely elongated tergal extensions. Their lengths clearly exceed tergal extensions in any other species, inclusive those in
|
||
<taxonomicName id="4C494D07E202FFA1FDF4FCE4FC1AFCF7" authority="Huys & Coomans, 1989" authorityName="Huys & Coomans" authorityYear="1989" box="[597,1002,871,896]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="higginsi">
|
||
<emphasis id="B93DEA96E202FFA1FDF4FCE4FD20FC08" box="[597,720,872,896]" italics="true" pageId="14" pageNumber="175">E. higginsi</emphasis>
|
||
<bibRefCitation id="EFD84B75E202FFA1FD79FCE5FC1AFCF7" author="Huys, R. & Coomans, A." box="[728,1002,871,896]" pageId="14" pageNumber="175" pagination="211 - 221" refId="ref19530" refString="Huys, R. & Coomans, A. (1989). Echinoderes higginsi sp. n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparede. Zoologisca Scripta, 18, 211 - 221." type="journal article" year="1989">Huys & Coomans, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D07E202FFA1FB82FCE4FEDFFC2C" authority="Sorensen, Heiner & Ziemer, 2005" authorityName="Sorensen, Heiner & Ziemer" authorityYear="2005" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="spinifurca">
|
||
<emphasis id="B93DEA96E202FFA1FB82FCE4FB44FC08" box="[1059,1204,872,896]" italics="true" pageId="14" pageNumber="175">E. spinifurca</emphasis>
|
||
<bibRefCitation id="EFD84B75E202FFA1FB1DFCEAFEDFFC2C" author="Sorensen, M. V. & Heiner, I. & Ziemer, O." pageId="14" pageNumber="175" pagination="499 - 508" refId="ref19911" refString="Sorensen, M. V., Heiner, I. & Ziemer, O. (2005) A new species of Echinoderes from Florida (Kinorhyncha: Cyclorhagida). Proceedings of the Biological Society of Washington, 118, 499 - 508." type="journal article" year="2005">Sørensen, Heiner & Ziemer, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
that otherwise possess the longest tergal extensions among the species known so far (see
|
||
<bibRefCitation id="EFD84B75E202FFA1FAE3FC01FEB4FC4F" author="Huys, R. & Coomans, A." pageId="14" pageNumber="175" pagination="211 - 221" refId="ref19530" refString="Huys, R. & Coomans, A. (1989). Echinoderes higginsi sp. n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparede. Zoologisca Scripta, 18, 211 - 221." type="journal article" year="1989">Huys & Coomans 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B75E202FFA1FEF3FC22FDB0FC4F" author="Sorensen, M. V. & Heiner, I. & Ziemer, O." box="[338,576,943,968]" pageId="14" pageNumber="175" pagination="499 - 508" refId="ref19911" refString="Sorensen, M. V., Heiner, I. & Ziemer, O. (2005) A new species of Echinoderes from Florida (Kinorhyncha: Cyclorhagida). Proceedings of the Biological Society of Washington, 118, 499 - 508." type="journal article" year="2005">
|
||
Sørensen
|
||
<emphasis id="B93DEA96E202FFA1FE60FC3CFE0BFC40" box="[449,507,944,968]" italics="true" pageId="14" pageNumber="175">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
). As stated above, the tergal extensions of
|
||
<taxonomicName id="4C494D07E202FFA1FB8DFC3CFB32FC40" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[1068,1218,945,968]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FB8DFC3CFB32FC40" box="[1068,1218,945,968]" italics="true" pageId="14" pageNumber="175">E. cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FB6AFC3DFAD6FC40" bold="true" box="[1227,1318,944,968]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FB6AFC3DFAD6FC40" box="[1227,1318,944,968]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
constitute 70% of the total segment length, whereas the corresponding ratios in
|
||
<taxonomicName id="4C494D07E202FFA1FC0DFC58FBD6FC64" authorityName="Huys & Coomans" authorityYear="1989" box="[940,1062,980,1004]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="higginsi">
|
||
<emphasis id="B93DEA96E202FFA1FC0DFC58FBD6FC64" box="[940,1062,980,1004]" italics="true" pageId="14" pageNumber="175">E. higginsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D07E202FFA1FBFEFC58FB00FC64" authorityName="Sorensen, Heiner & Ziemer" authorityYear="2005" box="[1119,1264,980,1004]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="spinifurca">
|
||
<emphasis id="B93DEA96E202FFA1FBFEFC58FB00FC64" box="[1119,1264,980,1004]" italics="true" pageId="14" pageNumber="175">E. spinifurca</emphasis>
|
||
</taxonomicName>
|
||
are about 50% only.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E202FFA1FF66FB91FD97FB28" blockId="14.[151,1436,151,1616]" pageId="14" pageNumber="175">
|
||
Also the distribution of glandular cell outlets
|
||
<typeStatus id="54F28826E202FFA1FD50FB91FCD1FBBC" box="[753,801,1052,1076]" pageId="14" pageNumber="175">type</typeStatus>
|
||
2 (gco2) appears to be unique for this species. Our information about the presence and distribution of gco2 is still quite limited, because previous descriptions have tended to neglect this character. However, to our knowledge, no other species displays the specific pattern of gco2 as the one found in
|
||
<taxonomicName id="4C494D07E202FFA1FED2FB04FDF7FB28" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[371,519,1161,1184]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FED2FB04FDF7FB28" box="[371,519,1161,1184]" italics="true" pageId="14" pageNumber="175">E. cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FDACFB05FD97FB28" bold="true" box="[525,615,1160,1184]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FDACFB05FD97FB28" box="[525,615,1160,1184]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF63684E202FFA1FF66FB21FEA1F9D8" blockId="14.[151,1436,151,1616]" pageId="14" pageNumber="175">
|
||
The third character that attracts special attention and makes
|
||
<taxonomicName id="4C494D07E202FFA1FCCDFB20FBF0FB4C" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[876,1024,1197,1220]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FCCDFB20FBF0FB4C" box="[876,1024,1197,1220]" italics="true" pageId="14" pageNumber="175">E. cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FBA6FB21FB91FB4C" bold="true" box="[1031,1121,1196,1220]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FBA6FB21FB91FB4C" box="[1031,1121,1196,1220]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
unique among congeners is the middorsal division of segment 11 that splits the tergal plate into two paired plates. No other species of the genus has this segment composition, and it inevitably makes the generic assignation of the species problematic. Among the five genera of
|
||
<taxonomicName id="4C494D07E202FFA1FEC8FA95FDFEFAB8" authorityName="Butschli" authorityYear="1876" box="[361,526,1304,1328]" class="Cyclorhagida" family="Echinoderidae" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="family">Echinoderidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D07E202FFA1FDBAFA95FC7DFAB8" authority="Adrianov, 1999" authorityName="Adrianov" authorityYear="1999" box="[539,909,1304,1328]" class="Cyclorhagida" family="Echinoderidae" genus="Cephalorhyncha" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B93DEA96E202FFA1FDBAFA95FD25FAB8" box="[539,725,1304,1328]" italics="true" pageId="14" pageNumber="175">Cephalorhyncha</emphasis>
|
||
<bibRefCitation id="EFD84B75E202FFA1FD7CFA95FC7DFAB8" author="Adrianov, A. V. & Malakhov, V. V." box="[733,909,1304,1328]" pageId="14" pageNumber="175" refId="ref18869" refString="Adrianov, A. V. & Malakhov, V. V. (1999) Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow, 328 pp." type="book" year="1999">
|
||
Adrianov,
|
||
<quantity id="4CB19B61E202FFA1FCF4FA95FC7DFAB8" box="[853,909,1304,1328]" metricMagnitude="1" metricUnit="m" metricValue="5.07746" pageId="14" pageNumber="175" unit="in" value="1999.0">1999</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFD84B75E202FFA1FC15FA95FB0FFAB8" author="Adrianov, A. V. & Malakhov, V. V." box="[948,1279,1304,1328]" pageId="14" pageNumber="175" refId="ref18869" refString="Adrianov, A. V. & Malakhov, V. V. (1999) Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow, 328 pp." type="book" year="1999">Adrianov & Malakhov, 1999</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C494D07E202FFA1FAADFA95FE38FADC" authority="Neuhaus & Blasche, 2006" authorityName="Neuhaus & Blasche" authorityYear="2006" class="Cyclorhagida" family="Echinoderidae" genus="Fissuroderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B93DEA96E202FFA1FAADFA95FA6CFAB8" box="[1292,1436,1304,1328]" italics="true" pageId="14" pageNumber="175">Fissuroderes</emphasis>
|
||
<bibRefCitation id="EFD84B75E202FFA1FF36FAB1FE38FADC" author="Neuhaus, B. & Blasche, T." box="[151,456,1340,1364]" pageId="14" pageNumber="175" pagination="19 - 52" refId="ref19700" refString="Neuhaus, B. & Blasche, T. (2006) Fissuroderes, a new genus of Kinorhyncha (Cyclorhagida) from the deep sea and continental shelf of New Zealand and from the continental shelf of Costa Rica. Zoologischer Anzeiger, 245, 19 - 52." type="journal article" year="2006">Neuhaus & Blasche, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C494D07E202FFA1FDA7FAB1FC60FADC" authority="Herranz et al., 2012" authorityName="Herranz" authorityYear="2012" box="[518,912,1340,1364]" class="Cyclorhagida" family="Echinoderidae" genus="Meristoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B93DEA96E202FFA1FDA7FAB1FD68FADC" box="[518,664,1340,1364]" italics="true" pageId="14" pageNumber="175">Meristoderes</emphasis>
|
||
<bibRefCitation id="EFD84B75E202FFA1FD05FAB1FC60FADC" author="Herranz, M. & Thormar, J. & Benito, J. & Sanchez, N. & Pardos, F." box="[676,912,1340,1364]" pageId="14" pageNumber="175" refId="ref19161" refString="Herranz, M., Thormar, J., Benito, J., Sanchez, N. & Pardos, F. (2012) Meristoderes gen. nov., a new kinorhynch genus, with the description of two new species and their implications for echinoderid phylogeny (Kinorhyncha: Cyclorhagida, Echinoderidae). Zoologischer Anzeiger, DOI: 10.1016 / j. jcz. 2011.08.004" type="book" year="2012">
|
||
Herranz
|
||
<emphasis id="B93DEA96E202FFA1FCABFAB0FCB6FADC" box="[778,838,1340,1364]" italics="true" pageId="14" pageNumber="175">et al.</emphasis>
|
||
, 2012
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
accommodate species with middorsally split tergal plate on segment 11 (
|
||
<bibRefCitation id="EFD84B75E202FFA1FE6EFAEDFD0BFAF0" author="Neuhaus, B. & Blasche, T." box="[463,763,1376,1400]" pageId="14" pageNumber="175" pagination="19 - 52" refId="ref19700" refString="Neuhaus, B. & Blasche, T. (2006) Fissuroderes, a new genus of Kinorhyncha (Cyclorhagida) from the deep sea and continental shelf of New Zealand and from the continental shelf of Costa Rica. Zoologischer Anzeiger, 245, 19 - 52." type="journal article" year="2006">Neuhaus & Blasche, 2006</bibRefCitation>
|
||
; for
|
||
<taxonomicName id="4C494D07E202FFA1FC91FAEDFBC1FAF0" authority="Sorensen" authorityName="Sorensen" box="[816,1073,1376,1400]" class="Cyclorhagida" family="Echinoderidae" genus="Meristoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B93DEA96E202FFA1FC91FAEDFC33FAF0" box="[816,963,1376,1400]" italics="true" pageId="14" pageNumber="175">Meristoderes</emphasis>
|
||
Sørensen
|
||
</taxonomicName>
|
||
pers. obs. from yet undescribed species). However, these genera are characterized by species with either fully or at least partly differentiated sternal plates on segment 2 (
|
||
<bibRefCitation id="EFD84B75E202FFA1FE23FA25FD34FA48" author="Adrianov, A. V. & Malakhov, V. V." box="[386,708,1448,1472]" pageId="14" pageNumber="175" refId="ref18869" refString="Adrianov, A. V. & Malakhov, V. V. (1999) Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow, 328 pp." type="book" year="1999">Adrianov & Malakhov 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B75E202FFA1FD71FA25FC1FFA48" author="Neuhaus, B. & Blasche, T." box="[720,1007,1448,1472]" pageId="14" pageNumber="175" pagination="19 - 52" refId="ref19700" refString="Neuhaus, B. & Blasche, T. (2006) Fissuroderes, a new genus of Kinorhyncha (Cyclorhagida) from the deep sea and continental shelf of New Zealand and from the continental shelf of Costa Rica. Zoologischer Anzeiger, 245, 19 - 52." type="journal article" year="2006">Neuhaus & Blasche 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B75E202FFA1FC5AFA25FB28FA48" author="Herranz, M. & Thormar, J. & Benito, J. & Sanchez, N. & Pardos, F." box="[1019,1240,1448,1472]" pageId="14" pageNumber="175" refId="ref19161" refString="Herranz, M., Thormar, J., Benito, J., Sanchez, N. & Pardos, F. (2012) Meristoderes gen. nov., a new kinorhynch genus, with the description of two new species and their implications for echinoderid phylogeny (Kinorhyncha: Cyclorhagida, Echinoderidae). Zoologischer Anzeiger, DOI: 10.1016 / j. jcz. 2011.08.004" type="book" year="2012">
|
||
Herranz
|
||
<emphasis id="B93DEA96E202FFA1FBFDFA24FB64FA48" box="[1116,1172,1448,1472]" italics="true" pageId="14" pageNumber="175">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). Segment
|
||
<quantity id="4CB19B61E202FFA1FAF2FA25FA8DFA48" box="[1363,1405,1448,1472]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="175" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C494D07E202FFA1FA25FA24FEFCFA6C" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FA25FA24FEFCFA6C" italics="true" pageId="14" pageNumber="175">E. cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FEB5FA41FE9FFA6C" bold="true" box="[276,367,1484,1508]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FEB5FA41FE9FFA6C" box="[276,367,1484,1508]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is composed of a closed cuticular ring, which characterizes species of
|
||
<taxonomicName id="4C494D07E202FFA1FB36FA41FAD2FA6C" authorityName="Claparede" authorityYear="1863" box="[1175,1314,1484,1508]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B93DEA96E202FFA1FB36FA41FAD2FA6C" box="[1175,1314,1484,1508]" italics="true" pageId="14" pageNumber="175">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, hence we tentatively assign the species to this genus. The conflicting characters could indicate that a future reassignment could be required, and in any case, this character combination clearly makes
|
||
<taxonomicName id="4C494D07E202FFA1FC5DF998FB60F9A4" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[1020,1168,1557,1580]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="175" phylum="Kinorhyncha" rank="species" species="cernunnos" status="sp. nov.">
|
||
<emphasis id="B93DEA96E202FFA1FC5DF998FB60F9A4" box="[1020,1168,1557,1580]" italics="true" pageId="14" pageNumber="175">E. cernunnos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E202FFA1FB39F999FB03F9A4" bold="true" box="[1176,1267,1556,1580]" pageId="14" pageNumber="175">
|
||
<taxonomicNameLabel id="A20E57EDE202FFA1FB39F999FB03F9A4" box="[1176,1267,1556,1580]" pageId="14" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
special among all echinoderids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |