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<document id="6F01D627AE68F6722AB19DC65CAE6710" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD6411413CFDFD58FB46FEDB" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola northlandensis Park and Carlton, new species" docType="treatment" docVersion="22" lastPageNumber="14" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="14" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CBDB6D83C3AFB8CA8084F94A32C01BAC">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="528B7C8BB2D3AF92AE6C56A7AB731FED">Park, J. - S.</mods:namePart>
<mods:affiliation id="F07DA3EB6264CDF4931CF782F1FAB299">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="CDEDA3290579328DCAB62931466E932F">Carlton, C. E.</mods:namePart>
<mods:affiliation id="76F3A29D788BEE9856DFD29DFE6DBDE0">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="598C450A2FCF0732767C9CBD54DF1FC4">The Coleopterists Bulletin</mods:title>
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<mods:date id="0D1479C3418000565DBC1AD211F69A70">2014</mods:date>
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<mods:title id="9BCEBE187AE002D3DB60A244BDD6DEB9">mo 13</mods:title>
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<treatment id="03D82A3EBD6411413CFDFD58FB46FEDB" ID-DOI="http://doi.org/10.5281/zenodo.5765235" ID-GBIF-Taxon="191245116" ID-Zenodo-Dep="5765235" LSID="urn:lsid:zoobank.org:act:A5C02EC2-107C-4F45-BCF5-14E08697343D" httpUri="http://treatment.plazi.org/id/03D82A3EBD6411413CFDFD58FB46FEDB" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="C36BC8A3BD6411413CFDFD58FE13FD40" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BCE9B28BD6411413CFDFD58FE13FD40" blockId="13.[184,605,683,757]" pageId="13" pageNumber="14">
<heading id="D0862C44BD6411413CFDFD58FE13FD40" centered="true" fontSize="10" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B905473ABD6411413CFDFD58FE13FD40" bold="true" pageId="13" pageNumber="14">
<taxonomicName id="4C71E0ABBD6411413CFDFD58FD8EFD5A" ID-CoL="4TYZT" authority="Park and Carlton" authorityName="Park and Carlton" box="[184,600,683,704]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="northlandensis" status="sp. nov.">
<emphasis id="B905473ABD6411413CFDFD58FE40FD5A" bold="true" box="[184,406,683,704]" italics="true" pageId="13" pageNumber="14">Sagola northlandensis</emphasis>
Park and Carlton
</taxonomicName>
,
<taxonomicNameLabel id="A236FA41BD6411413D0AFD36FE13FD40" box="[335,453,709,730]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36BC8A3BD6411413D79FD13FE0FFD6F" box="[316,473,736,757]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BCE9B28BD6411413D79FD13FE0FFD6F" blockId="13.[184,605,683,757]" box="[316,473,736,757]" pageId="13" pageNumber="14">
(
<figureCitation id="134A87ADBD6411413D01FD13FE42FD6F" box="[324,404,736,757]" captionStart="Fig" captionStartId="8.[197,229,1638,1657]" captionTargetBox="[190,1185,642,1617]" captionTargetId="figure-178@8.[173,1193,624,1622]" captionTargetPageId="8" captionText="Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907747" httpUri="https://zenodo.org/record/4907747/files/figure.png" pageId="13" pageNumber="14">Figs. 3e</figureCitation>
,
<figureCitation id="134A87ADBD6411413DE4FD13FE6FFD6F" box="[417,441,736,757]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="13" pageNumber="14">4e</figureCitation>
,
<figureCitation id="134A87ADBD6411413D80FD13FE07FD6F" box="[453,465,736,757]" captionStart="Fig" captionStartId="10.[731,763,905,924]" captionTargetBox="[710,1193,188,881]" captionTargetId="figure-608@10.[707,1194,183,884]" captionTargetPageId="10" captionText="Fig. 5. Known collection localities of New Zealand Sagola species. S. hirtalis: black circles; S. terricola: triangles; S. convexa: black squares; S. coromandelensis: star; S. northlandensis: white circle; S. taranakiensis: white square." figureDoi="http://doi.org/10.5281/zenodo.4907753" httpUri="https://zenodo.org/record/4907753/files/figure.png" pageId="13" pageNumber="14">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6411413CEAFCF9FDB0FC3E" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BCE9B28BD6411413CEAFCF9FDB0FC3E" blockId="13.[151,639,778,1729]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413CEAFCF9FE90FC85" bold="true" box="[175,326,778,799]" pageId="13" pageNumber="14">Type Material.</emphasis>
<materialsCitation id="3B199175BD6411413D11FCF9FDB0FC3E" ID-GBIF-Occurrence="3415153805" collectingDate="1974-10-10" collectionCode="NZAC" country="New Zealand" location="Northland" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Northland" typeStatus="holotype">
<emphasis id="B905473ABD6411413D11FCF9FEC2FCA0" bold="true" pageId="13" pageNumber="14">
<typeStatus id="54CA258ABD6411413D11FCF9FE65FC85" box="[340,435,778,799]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F366DBB8BD6411413DFBFCF9FDAFFC85" box="[446,633,778,799]" name="New Zealand" pageId="13" pageNumber="14">NEW ZEALAND</collectingCountry>
:
<collectingRegion id="49B555CABD6411413CD2FCD6FED8FCA0" box="[151,270,805,826]" country="New Zealand" name="Northland" pageId="13" pageNumber="14">Northland</collectingRegion>
:
</emphasis>
<specimenCount id="9D7750A1BD6411413D5AFCD6FEE8FCA1" box="[287,318,805,827]" pageId="13" pageNumber="14" type="male">1♂</specimenCount>
(
<collectionCode id="ED6003EDBD6411413D15FCD6FE4DFCA0" box="[336,411,805,826]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="13" pageNumber="14" type="Other institutional type">NZAC</collectionCode>
), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand: ND: Omahuta S.F., Kauri Res., litter 74/81
<date id="FFCFBDE8BD6411413D4CFC86FE53FC10" box="[265,389,885,906]" pageId="13" pageNumber="14" value="1974-10-10">
<collectingDate id="EF8B4400BD6411413D4CFC86FE53FC10" box="[265,389,885,906]" pageId="13" pageNumber="14" value="1974-10-10">10 Oct 1974</collectingDate>
</date>
, JC Watt”, “
<typeStatus id="54CA258ABD6411413E44FC86FDA8FC10" box="[513,638,885,906]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C71E0ABBD6411413CD2FC7CFD80FC3E" authority="Park and Carlton 2013" authorityName="Park and Carlton" authorityYear="2013" box="[151,598,911,932]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="northlandensis">
<emphasis id="B905473ABD6411413CD2FC7CFEA4FC3E" bold="true" box="[151,370,911,932]" italics="true" pageId="13" pageNumber="14">Sagola northlandensis</emphasis>
Park and Carlton 2013
</taxonomicName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6411413CEAFC59FD93FC43" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BCE9B28BD6411413CEAFC59FD93FC43" blockId="13.[151,639,778,1729]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413CEAFC59FEF4FC25" bold="true" box="[175,290,938,959]" pageId="13" pageNumber="14">Etymology.</emphasis>
The species name is based on the biotic region of the
<typeStatus id="54CA258ABD6411413D1FFC37FE55FC43" box="[346,387,964,985]" pageId="13" pageNumber="14">type</typeStatus>
locality,
<collectingRegion id="49B555CABD6411413D9BFC37FD97FC43" box="[478,577,964,985]" country="New Zealand" name="Northland" pageId="13" pageNumber="14">Northland</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6411413CEAFC2CFE4BFB34" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BCE9B28BD6411413CEAFC2CFE4BFB34" blockId="13.[151,639,778,1729]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413CEAFC2CFECEFC6E" bold="true" box="[175,280,991,1012]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
This species can be distinguished from the other species of the
<taxonomicName id="4C71E0ABBD6411413D8AFC09FDC1FB95" authorityName="Broun, 1893 b:" authorityYear="1050" box="[463,535,1018,1039]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hirtalis">
<emphasis id="B905473ABD6411413D8AFC09FDC1FB95" box="[463,535,1018,1039]" italics="true" pageId="13" pageNumber="14">hirtalis</emphasis>
</taxonomicName>
group by the following combination of characters: antennomere 1 elongate, approximately 3 times longer than wide; antennomere 4 elongate; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6411413CEAFB47FC0FFF68" pageId="13" pageNumber="14" type="description">
<paragraph id="8BCE9B28BD6411413CEAFB47FC0FFF68" blockId="13.[151,639,778,1729]" lastBlockId="13.[685,1172,194,321]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413CEAFB47FE54FB53" bold="true" box="[175,386,1204,1225]" pageId="13" pageNumber="14">Description of Male.</emphasis>
Length 3.0 mm. Body brown and elytra, legs, maxillary palpi yellowish brown (
<figureCitation id="134A87ADBD6411413CA0FB1AFEF9FB64" box="[229,303,1257,1278]" captionStart="Fig" captionStartId="8.[197,229,1638,1657]" captionTargetBox="[190,1185,642,1617]" captionTargetId="figure-178@8.[173,1193,624,1622]" captionTargetPageId="8" captionText="Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907747" httpUri="https://zenodo.org/record/4907747/files/figure.png" pageId="13" pageNumber="14">Fig. 3e</figureCitation>
).
<emphasis id="B905473ABD6411413D01FB1AFE54FB64" bold="true" box="[324,386,1257,1278]" pageId="13" pageNumber="14">Head:</emphasis>
As long as wide, widest across eyes (
<figureCitation id="134A87ADBD6411413D5BFAF0FEBCFA82" box="[286,362,1283,1304]" captionStart="Fig" captionStartId="8.[197,229,1638,1657]" captionTargetBox="[190,1185,642,1617]" captionTargetId="figure-178@8.[173,1193,624,1622]" captionTargetPageId="8" captionText="Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907747" httpUri="https://zenodo.org/record/4907747/files/figure.png" pageId="13" pageNumber="14">Fig. 3e</figureCitation>
). Antennomere 1 approximately 3 times longer than wide, 23 longer than wide, 4 elongate, approximately 1.5 times longer than wide, 5 longer than wide, 610 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple.
<emphasis id="B905473ABD6411413E69FA4EFDABFA48" bold="true" box="[556,637,1469,1490]" pageId="13" pageNumber="14">Thorax:</emphasis>
Prosternum as long as wide, widest at one-third length. Elytra rectangular (
<figureCitation id="134A87ADBD6411413DECFA01FE25F99D" box="[425,499,1522,1543]" captionStart="Fig" captionStartId="8.[197,229,1638,1657]" captionTargetBox="[190,1185,642,1617]" captionTargetId="figure-178@8.[173,1193,624,1622]" captionTargetPageId="8" captionText="Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907747" httpUri="https://zenodo.org/record/4907747/files/figure.png" pageId="13" pageNumber="14">Fig. 3e</figureCitation>
). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide.
<emphasis id="B905473ABD6411413D0CF9D4FE65F9A6" bold="true" box="[329,435,1575,1596]" pageId="13" pageNumber="14">Abdomen:</emphasis>
Tergite IV with pair of transverse patches of microtrichia reaching middle.
<emphasis id="B905473ABD6411413CB7F9AEFEB2F9E8" bold="true" box="[242,356,1629,1650]" pageId="13" pageNumber="14">Aedeagus:</emphasis>
Median lobe longer than parameres with semicircular apical lobe (
<figureCitation id="134A87ADBD6411413E6EF984FDA5F916" box="[555,627,1655,1676]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="13" pageNumber="14">Fig. 4e</figureCitation>
). Phallobase of median lobe asymmetrical and triangular (
<figureCitation id="134A87ADBD6411413CB1F95FFE94F95B" box="[244,322,1708,1729]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="13" pageNumber="14">Fig. 4e</figureCitation>
). Right paramere divided into 2 branches from base, and minor branch divided into 2 long branches (
<figureCitation id="134A87ADBD6411413FC2FF2EFC18FF68" box="[903,974,221,242]" captionStart="Fig" captionStartId="9.[175,207,1144,1163]" captionTargetBox="[166,1166,194,1123]" captionTargetId="figure-247@9.[151,1171,183,1135]" captionTargetPageId="9" captionText="Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907749" httpUri="https://zenodo.org/record/4907749/files/figure.png" pageId="13" pageNumber="14">Fig. 4e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD6411413E80FF04FB46FEDB" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BCE9B28BD6411413E80FF04FB5AFE96" blockId="13.[685,1172,194,321]" box="[709,1164,247,268]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413E80FF04FC9EFE96" bold="true" box="[709,840,247,268]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingRegion id="49B555CABD6411413F10FF04FC63FE96" box="[853,949,247,268]" country="New Zealand" name="Northland" pageId="13" pageNumber="14">Northland</collectingRegion>
(
<figureCitation id="134A87ADBD6411413F81FF04FBD7FE96" box="[964,1025,247,268]" captionStart="Fig" captionStartId="10.[731,763,905,924]" captionTargetBox="[710,1193,188,881]" captionTargetId="figure-608@10.[707,1194,183,884]" captionTargetPageId="10" captionText="Fig. 5. Known collection localities of New Zealand Sagola species. S. hirtalis: black circles; S. terricola: triangles; S. convexa: black squares; S. coromandelensis: star; S. northlandensis: white circle; S. taranakiensis: white square." figureDoi="http://doi.org/10.5281/zenodo.4907753" httpUri="https://zenodo.org/record/4907753/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
: white circle).
</paragraph>
<paragraph id="8BCE9B28BD6411413E80FEE1FB46FEDB" blockId="13.[685,1172,194,321]" pageId="13" pageNumber="14">
<emphasis id="B905473ABD6411413E80FEE1FCCCFEBD" bold="true" box="[709,794,274,295]" pageId="13" pageNumber="14">Habitat.</emphasis>
The
<typeStatus id="54CA258ABD6411413F10FEE1FC7EFEBD" box="[853,936,274,295]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and only known specimen of this species was collected by sifting litter.
</paragraph>
</subSubSection>
</treatment>
</document>