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<document id="3D25EC18B552A9A98D66EEA6E9D4495B" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD5D11793F68FEEFFDABFD33" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola helmorei" docType="treatment" docVersion="25" lastPageNumber="54" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="53" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C4493C44A461B73C438EB4C757214CCB">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="46532ABF16FD0C5C6339ED6E785428E2">Park, J. - S.</mods:namePart>
<mods:affiliation id="21639695616E29DD154FF0B777A4F9FE">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="E53BB617D4DFE04BF36A55C0EA9917A5">Carlton, C. E.</mods:namePart>
<mods:affiliation id="61872481EFA98CFFB0724627AF431269">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="AB7DDD80193175B7DC595BDBAA5B57F5">The Coleopterists Bulletin</mods:title>
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<mods:part id="1D29686FA2610552452EEAB4793FAC9F">
<mods:date id="42BA5604600CEE8D156145197EF0CD2A">2014</mods:date>
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<mods:title id="666D1A815F22C702F67A15A1BFAEE7BF">mo 13</mods:title>
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<treatment id="03D82A3EBD5D11793F68FEEFFDABFD33" ID-DOI="http://doi.org/10.5281/zenodo.5765455" ID-GBIF-Taxon="191245054" ID-Zenodo-Dep="5765455" LSID="urn:lsid:plazi:treatment:03D82A3EBD5D11793F68FEEFFDABFD33" httpUri="http://treatment.plazi.org/id/03D82A3EBD5D11793F68FEEFFDABFD33" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection id="C36BC8A3BD5D11783F68FEEFFBE9FEAB" box="[813,1087,284,305]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BCE9B28BD5D11783F68FEEFFBE9FEAB" blockId="52.[813,1087,284,332]" box="[813,1087,284,305]" pageId="52" pageNumber="53">
<heading id="D0862C44BD5D11783F68FEEFFBE9FEAB" box="[813,1087,284,305]" centered="true" fontSize="10" level="2" pageId="52" pageNumber="53" reason="2">
<emphasis id="B905473ABD5D11783F68FEEFFBE9FEAB" bold="true" box="[813,1087,284,305]" pageId="52" pageNumber="53">
10.
<taxonomicName id="4C71E0ABBD5D11783F11FEEFFC7DFEAB" ID-CoL="4TYY9" authority="SHARP" box="[852,939,284,305]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783F11FEEFFC7DFEAB" bold="true" box="[852,939,284,305]" italics="true" pageId="52" pageNumber="53">helmorei</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD5D11783FC4FEC4FC3CFED6" box="[897,1002,311,332]" pageId="52" pageNumber="53" type="description">
<paragraph id="8BCE9B28BD5D11783FC4FEC4FC3CFED6" blockId="52.[813,1087,284,332]" box="[897,1002,311,332]" pageId="52" pageNumber="53">(2 species)</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD5D11793E9EFE92FEE7FF68" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BCE9B28BD5D11793E9EFE92FEE7FF68" blockId="52.[707,1193,353,667]" lastBlockId="53.[151,637,194,242]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="B905473ABD5D11783E9EFE92FC92FEEC" bold="true" box="[731,836,353,374]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
Members of the
<taxonomicName id="4C71E0ABBD5D11783FBEFE91FB86FEED" box="[1019,1104,354,375]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783FBEFE91FB86FEED" box="[1019,1104,354,375]" italics="true" pageId="52" pageNumber="53">helmorei</emphasis>
</taxonomicName>
speciesgroup can be distinguished from other
<taxonomicName id="4C71E0ABBD5D11783821FE8FFB7FFE0B" box="[1124,1193,380,401]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B905473ABD5D11783821FE8FFB7FFE0B" box="[1124,1193,380,401]" italics="true" pageId="52" pageNumber="53">Sagola</emphasis>
</taxonomicName>
species-groups by the following combination of characters: body length
<quantity id="4C8936CDBD5D11783FFFFE42FBE1FE5C" box="[954,1079,433,455]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.9" metricValueMin="2.3" pageId="52" pageNumber="53" unit="mm" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.32.9 mm</quantity>
; head subquadrate with dense setae around frontal sulcus (
<figureCitation id="134A87ADBD5D11783E8CFE15FCF8FE61" box="[713,814,486,508]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53">Fig. 29ab</figureCitation>
); antennomere 1 approximately 2.5 times longer than wide with dull surface; anterior and posterior frontal foveae round (
<figureCitation id="134A87ADBD5D11783F8DFDE8FBCFFDAA" box="[968,1049,539,560]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53">Fig. 29e</figureCitation>
); frontal sulcus reaching midpoint of eye (
<figureCitation id="134A87ADBD5D11783FF7FDC5FBD6FDD1" box="[946,1024,566,587]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53">Fig. 29e</figureCitation>
); hind wings welldeveloped; legs elongate and slender (
<figureCitation id="134A87ADBD5D11783876FDA2FB4BFDFC" box="[1075,1181,592,614]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53">Fig. 29ab</figureCitation>
); abdominal tergites IVVI with discal carinae; genitalia small, not exceeding
<quantity id="4C8936CDBD5D11783F8DFD75FBF6FD01" box="[968,1056,646,667]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="52" pageNumber="53" unit="mm" value="0.25">0.25 mm</quantity>
(
<figureCitation id="134A87ADBD5D11783875FD75FB4BFD01" box="[1072,1181,645,667]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53">Fig. 29cd</figureCitation>
); present on North Island, not known from South Island (
<figureCitation id="134A87ADBD5C11793C9BFF2EFEF3FF68" box="[222,293,221,242]" captionStart="Fig" captionStartId="53.[709,741,1662,1681]" captionTargetBox="[688,1170,943,1636]" captionTargetId="figure-573@53.[685,1171,938,1639]" captionTargetPageId="53" captionText="Fig. 30. Known collection localities of New Zealand Sagola species.S. helmorei: black circles;S. wellingtonensis: triangle." figureDoi="http://doi.org/10.5281/zenodo.4907821" httpUri="https://zenodo.org/record/4907821/files/figure.png" pageId="53" pageNumber="54">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0ECBA0BD5D11783C80F98DFDD7F95B" ID-DOI="http://doi.org/10.5281/zenodo.4907819" ID-Zenodo-Dep="4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="52" pageNumber="53" startId="52.[197,229,1662,1681]" targetBox="[188,1181,753,1641]" targetPageId="52">
<paragraph id="8BCE9B28BD5D11783C80F98DFDD7F95B" blockId="52.[173,1194,1662,1729]" pageId="52" pageNumber="53">
<emphasis id="B905473ABD5D11783C80F98DFEDDF90B" bold="true" box="[197,267,1662,1681]" pageId="52" pageNumber="53">Fig. 29.</emphasis>
<taxonomicName id="4C71E0ABBD5D11783D65F98CFE49F90B" box="[288,415,1662,1681]" class="Insecta" family="Staphylinidae" genus="Sagola" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B905473ABD5D11783D65F98CFE8DF90B" box="[288,347,1663,1681]" italics="true" pageId="52" pageNumber="53">Sagola</emphasis>
species
</taxonomicName>
in the
<taxonomicName id="4C71E0ABBD5D11783D99F98CFDF3F90B" box="[476,549,1663,1681]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783D99F98CFDF3F90B" box="[476,549,1663,1681]" italics="true" pageId="52" pageNumber="53">helmorei</emphasis>
</taxonomicName>
species-group. Habitus, dorsal views: a)
<taxonomicName id="4C71E0ABBD5D11783FC2F98CFC30F90B" box="[903,998,1663,1681]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783FC2F98CFC30F90B" box="[903,998,1663,1681]" italics="true" pageId="52" pageNumber="53">S. helmorei</emphasis>
</taxonomicName>
, b)
<taxonomicName id="4C71E0ABBD5D1178384FF98CFB75F90B" box="[1034,1187,1663,1681]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="wellingtonensis">
<emphasis id="B905473ABD5D1178384FF98CFB75F90B" box="[1034,1187,1663,1681]" italics="true" pageId="52" pageNumber="53">S. wellingtonensis</emphasis>
</taxonomicName>
. Aedeagi, dorsal views: c)
<taxonomicName id="4C71E0ABBD5D11783DCDF965FE30F932" box="[392,486,1686,1704]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783DCDF965FE30F932" box="[392,486,1686,1704]" italics="true" pageId="52" pageNumber="53">S. helmorei</emphasis>
</taxonomicName>
, d)
<taxonomicName id="4C71E0ABBD5D11783E4DF965FD76F932" box="[520,672,1686,1704]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="wellingtonensis">
<emphasis id="B905473ABD5D11783E4DF965FD76F932" box="[520,672,1686,1704]" italics="true" pageId="52" pageNumber="53">S. wellingtonensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C71E0ABBD5D11783EEFF965FCDFF932" box="[682,777,1686,1705]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="helmorei">
<emphasis id="B905473ABD5D11783EEFF965FD63F932" box="[682,693,1686,1704]" italics="true" pageId="52" pageNumber="53">S</emphasis>
.
<emphasis id="B905473ABD5D11783E85F965FCDFF932" box="[704,777,1686,1704]" italics="true" pageId="52" pageNumber="53">helmorei</emphasis>
</taxonomicName>
, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C36BC8A3BD5C11793C90FEDAFDABFD33" pageId="53" pageNumber="54" type="key">
<paragraph id="8BCE9B28BD5C11793C90FEDAFE0BFEC6" blockId="53.[213,576,297,350]" pageId="53" pageNumber="54">
<emphasis id="B905473ABD5C11793C90FEDAFE0BFEC6" bold="true" pageId="53" pageNumber="54">
KEY TO SPECIES OF THE
<emphasis id="B905473ABD5C11793D99FEDDFD96FEA4" bold="true" box="[476,576,302,318]" italics="true" pageId="53" pageNumber="54">HELMOREI</emphasis>
SPECIES- GROUP
</emphasis>
</paragraph>
<paragraph id="8BCE9B28BD5C11793CEAFE80FE25FE27" blockId="53.[151,638,371,445]" pageId="53" pageNumber="54">The key is mainly based on male specimens because most female specimens are indistinguishable based on external morphology.</paragraph>
<keyStep id="3085838DBD5C11793CD2FE21FDABFD33" pageId="53" pageNumber="54">
<key id="FFE0CC19BD5C11793CD2FE21FDABFD33" pageId="53" pageNumber="54">
<paragraph id="8BCE9B28BD5C11793CD2FE21FDA8FDAD" blockId="53.[151,638,466,681]" pageId="53" pageNumber="54">
<keyLead id="3080381DBD5C11793CD2FE21FDA8FDAD" pageId="53" pageNumber="54">
1. Ventral surface of temple normal (
<figureCitation id="134A87ADBD5C11793E58FE21FDBAFE7D" box="[541,620,466,487]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="53" pageNumber="54">Fig. 29f</figureCitation>
); antennomeres 68 longer than wide; left paramere divided for one-third length (
<figureCitation id="134A87ADBD5C11793E6EFDFBFDA1FD87" box="[555,631,520,541]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="53" pageNumber="54">Fig. 29c</figureCitation>
) ......................................
<emphasis id="B905473ABD5C11793DD8FDD1FDA8FDAD" bold="true" box="[413,638,546,567]" pageId="53" pageNumber="54">
<taxonomicName id="4C71E0ABBD5C11793DD8FDD1FDDEFDAD" ID-CoL="4TYY9" box="[413,520,546,567]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="helmorei" status="sp. nov.">
<emphasis id="B905473ABD5C11793DD8FDD1FDDEFDAD" bold="true" box="[413,520,546,567]" italics="true" pageId="53" pageNumber="54">S. helmorei</emphasis>
</taxonomicName>
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<paragraph id="8BCE9B28BD5C11793CD2FDB6FDABFD33" blockId="53.[151,638,466,681]" pageId="53" pageNumber="54">
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1. Ventral surface of temple depressed and flat; antennomeres 68 subquadrate; left paramere more deeply divided for two-thirds length (
<figureCitation id="134A87ADBD5C11793C96FD67FEF5FD33" box="[211,291,660,681]" captionStart="Fig" captionStartId="52.[197,229,1662,1681]" captionTargetBox="[188,1181,753,1641]" captionTargetId="figure-229@52.[173,1193,733,1661]" captionTargetPageId="52" captionText="Fig. 29. Sagola species in the helmorei species-group. Habitus, dorsal views: a) S. helmorei, b) S. wellingtonensis. Aedeagi, dorsal views: c) S. helmorei, d) S. wellingtonensis. S. helmorei, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907819" httpUri="https://zenodo.org/record/4907819/files/figure.png" pageId="53" pageNumber="54">Fig. 29d</figureCitation>
) .......
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<taxonomicName id="4C71E0ABBD5C11793D13FD67FDD2FD33" ID-CoL="4TZ4L" box="[342,516,660,681]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B905473ABD5C11793D13FD67FDD2FD33" bold="true" box="[342,516,660,681]" italics="true" pageId="53" pageNumber="54">S. wellingtonensis</emphasis>
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