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<document id="15AA8E5B7C45B818A413B81FE2A03820" ID-DOI="10.11646/zootaxa.3872.2.1" ID-GBIF-Dataset="e9b8d555-d661-4f1a-86e8-f2a0acc38798" ID-ISSN="1175-5326" ID-Zenodo-Dep="253224" ID-ZooBank="F9F81CC8-E033-46B7-B73B-9FB777DF4116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460926510372" checkinUser="plazi" docAuthor="Ponder, Winston F., Fukuda, Hiroshi &amp; Hallan, Anders" docDate="2014" docId="03CDAD6557653C05FF05FB0EB9C8AB4F" docLanguage="en" docName="zt03872p153.pdf" docOrigin="Zootaxa 3872 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coliracemata mortoni Ponder, Fukuda &amp; Hallan, 2014, n. sp." docType="treatment" docUuid="86E58BC1-1EB8-4B38-A7C6-0AEA33630A50" docUuidSource="ZooBank" docVersion="9" lastPageNumber="134" masterDocId="FFF4D51D57783C24FF92FFB2BB3EA911" masterDocTitle="A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)" masterLastPageNumber="153" masterPageNumber="101" pageNumber="130" updateTime="1698362634141" updateUser="plazi">
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<mods:title id="69F4DDE4209B033A49F53CA4156FD1D5">A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)</mods:title>
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<mods:namePart id="4141AC79BB229BE42AF12AAC7B473D41">Ponder, Winston F.</mods:namePart>
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<mods:namePart id="F19F8177DF4E69B8798BED62447033A4">Fukuda, Hiroshi</mods:namePart>
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<mods:namePart id="09FDD1BD9E02B053FB9FE5A9CF208F21">Hallan, Anders</mods:namePart>
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<mods:date id="D1D70AC5C8AC5284C6642D6C557B3800">2014</mods:date>
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<treatment id="03CDAD6557653C05FF05FB0EB9C8AB4F" ID-DOI="http://doi.org/10.5281/zenodo.5631021" ID-GBIF-Taxon="119589573" ID-Zenodo-Dep="5631021" LSID="urn:lsid:zoobank.org:act:86E58BC1-1EB8-4B38-A7C6-0AEA33630A50" httpUri="http://treatment.plazi.org/id/03CDAD6557653C05FF05FB0EB9C8AB4F" lastPageId="33" lastPageNumber="134" pageId="29" pageNumber="130">
<subSubSection id="C37E4FF857653C39FF05FB0EBA5CADE6" pageId="29" pageNumber="130" type="nomenclature">
<paragraph id="8BDB1C7357653C39FF05FB0EBAD4ADC7" blockId="29.[151,490,1212,1271]" box="[151,490,1212,1238]" pageId="29" pageNumber="130">
<heading id="D093AB1F57653C39FF05FB0EBAD4ADC7" bold="true" box="[151,490,1212,1238]" fontSize="11" level="1" pageId="29" pageNumber="130" reason="1">
<emphasis id="B910C06157653C39FF05FB0EBAD4ADC7" bold="true" box="[151,490,1212,1238]" pageId="29" pageNumber="130">
<taxonomicName id="4C6467F057653C39FF05FB0EBA9FADC7" ID-CoL="X2L9" box="[151,417,1212,1238]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="mortoni" status="sp. nov.">
<emphasis id="B910C06157653C39FF05FB0EBA9FADC7" bold="true" box="[151,417,1212,1238]" italics="true" pageId="29" pageNumber="130">Coliracemata mortoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57653C39FE3AFB0EBAD4ADC7" box="[424,490,1212,1238]" pageId="29" pageNumber="130" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDB1C7357653C39FF05FB6CBA5CADE6" blockId="29.[151,490,1212,1271]" box="[151,354,1246,1271]" pageId="29" pageNumber="130">
<figureCitation id="135F00F657653C39FF05FB6CBA05ADE6" box="[151,315,1246,1271]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStart-5="FIGURE 21" captionStart-6="FIGURE 22" captionStartId-0="30.[151,250,1217,1239]" captionStartId-1="30.[151,250,1976,1998]" captionStartId-2="31.[151,250,518,540]" captionStartId-3="31.[151,250,1519,1541]" captionStartId-4="32.[151,250,1768,1790]" captionStartId-5="33.[151,250,1716,1738]" captionStartId-6="34.[151,250,1251,1273]" captionTargetBox-0="[214,1370,253,1203]" captionTargetBox-1="[208,1350,1354,1961]" captionTargetBox-2="[262,1329,199,489]" captionTargetBox-3="[212,1360,615,1486]" captionTargetBox-4="[161,1427,204,1740]" captionTargetBox-5="[150,1436,663,1683]" captionTargetBox-6="[182,1402,441,1239]" captionTargetId-0="figure@30.[214,1370,253,1203]" captionTargetId-1="figure@30.[202,1358,1346,1966]" captionTargetId-2="figure@31.[249,1338,193,496]" captionTargetId-3="figure@31.[210,1367,611,1493]" captionTargetId-4="figure@32.[152,1435,193,1746]" captionTargetId-5="figure@33.[137,1449,658,1694]" captionTargetId-6="figure@34.[182,1403,441,1241]" captionTargetPageId-0="30" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="31" captionTargetPageId-4="32" captionTargetPageId-5="33" captionTargetPageId-6="34" captionText-0="FIGURE 16. Holotypes of new species of Coliracemata n. gen. (from left towards right: dorsal, lateral and ventral view respectively). A C, Coliracemata mortoni n. sp. (AMS C. 269811; shell diameter 1.82 mm); D F, Coliracemata katurana n. sp. (AMS C. 445414; shell diameter 1.06 mm); G I, Coliracemata clarkae n. sp. (AMS C. 462962; shell diameter 1.36 mm)." captionText-1="FIGURE 17. Shells of Coliracemata mortoni (AMS C. 460381; A C) and Coliracemata katurana (AMS C. 445414; D F) with dorsal, lateral and ventral views given from left towards right." captionText-2="FIGURE 18. External and internal view (left to right) of opercula. A, B, Coliracemata mortoni (AMS C. 460381); C, D, Coliracemata katurana (AMS C. 445414). Scale bars: 100 µm." captionText-3="FIGURE 19. Head-foot and gut characters of Coliracemata mortoni (AMS C. 460381). A, head, dorsal view; B, posterior end of foot, dorsal view; C, external view of intestine and rectum; D, stomach. Scale bars: 200 µm. Abbreviations: ac, anterior chamber; ct, cephalic tentacle; fp, faecal pellet; int, intestine; o, oesophagus; pc, posterior chamber; r, rectum; ss, style sac." captionText-4="FIGURE 20. Cephalic and penial characters of Coliracemata mortoni. Mong Tseng Wai, Hong Kong (AMS C. 463401). A C, cephalic tentacle showing distinct compound cilia on distal tip; D, apocrine glands situated on penis; E, F, penis with clusters of apocrine glands." captionText-5="FIGURE 21. Radulae of Coliracemata mortoni (AMS C. 460381) and Coliracemata katurana (AMS C. 445414). Coliracemata mortoni: A, central, lateral and marginal teeth; B, central and lateral teeth. Coliracemata katurana: C, central, lateral and marginal teeth; D, central and lateral teeth." captionText-6="FIGURE 22. Reproductive anatomy of Coliracemata mortoni (AMS C. 460381). A, male head and penis; B, penis, ventral view; C, penis showing separate groups of apocrine glands; D, female reproductive system shown from anterior (left) to posterior (right); E, female reproductive system shown from posterior (left) to anterior (right). Scale bars: 200 µm. Abbreviations: ag, albumen gland; bc, bursa copulatrix; cag, clusters of apocrine glands; cb, cut base of penis; cg, capsule gland; co, coiled oviduct; ct, cephalic tentacle; fa, female aperture; gg, groups of apocrine glands; gm, glandular mass; pd, penial duct; ppw, position of posterior wall of mantle cavity; sr, seminal receptacle." httpUri-0="https://zenodo.org/record/253240/files/figure.png" httpUri-1="https://zenodo.org/record/253241/files/figure.png" httpUri-2="https://zenodo.org/record/253242/files/figure.png" httpUri-3="https://zenodo.org/record/253243/files/figure.png" httpUri-4="https://zenodo.org/record/253244/files/figure.png" httpUri-5="https://zenodo.org/record/253245/files/figure.png" httpUri-6="https://zenodo.org/record/253246/files/figure.png" pageId="29" pageNumber="130">Figures 1622</figureCitation>
,
<figureCitation id="135F00F657653C39FED4FB6CBA5CADE6" box="[326,354,1246,1271]" captionStart="FIGURE 26" captionStartId="37.[151,250,1778,1800]" captionTargetBox="[257,1327,694,1762]" captionTargetId="figure@37.[257,1327,694,1762]" captionTargetPageId="37" captionText="FIGURE 26. Type localities of all species of Coliracemata n. gen., Colenuda n. gen., and Coleglabra n. gen." httpUri="https://zenodo.org/record/253250/files/figure.png" pageId="29" pageNumber="130">26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857653C39FF05FAADBA81ACD9" pageId="29" pageNumber="130" type="etymology">
<paragraph id="8BDB1C7357653C39FF05FAADB896AC4C" blockId="29.[151,1437,1311,2021]" pageId="29" pageNumber="130">
<emphasis id="B910C06157653C39FF05FAADBA20AC29" bold="true" box="[151,286,1311,1336]" pageId="29" pageNumber="130">Etymology.</emphasis>
Named for Professor Brian Morton as a small recognition of his enormous contributions to marine biological and malacological studies on the
<collectingCountry id="F3735CE357653C39FD10FAF7B838AC4C" box="[642,774,1349,1373]" name="Hong Kong" pageId="29" pageNumber="130">Hong Kong</collectingCountry>
marine fauna.
</paragraph>
<paragraph id="8BDB1C7357653C39FF55FAD5BA81ACD9" blockId="29.[151,1437,1311,2021]" pageId="29" pageNumber="130">
<emphasis id="B910C06157653C39FF55FAD5BADAAC91" bold="true" box="[199,484,1383,1408]" pageId="29" pageNumber="130">
<typeStatus id="54DFA2D157653C39FF55FAD5BA32AC91" box="[199,268,1383,1408]" pageId="29" pageNumber="130">Types</typeStatus>
and
<typeStatus id="54DFA2D157653C39FED9FAD5BA40AC91" box="[331,382,1383,1408]" pageId="29" pageNumber="130">type</typeStatus>
locality.
</emphasis>
<typeStatus id="54DFA2D157653C39FE7FFAD5B964AC91" box="[493,602,1383,1408]" pageId="29" pageNumber="130" type="holotype">Holotype</typeStatus>
: Under wood, front edge of mangrove. Mong Tseng Wai, Southern Deep Bay,
<collectingCountry id="F3735CE357653C39FF40FA3FBA64ACB4" box="[210,346,1421,1445]" name="Hong Kong" pageId="29" pageNumber="130">Hong Kong</collectingCountry>
. 22˚29 N, 114˚00E,
<date id="FFDA3AB357653C39FDC7FA3EB9D0ACB5" box="[597,750,1420,1445]" pageId="29" pageNumber="130" value="1983-04-16">16 Apr. 1983</date>
. Coll. W. F. Ponder (AMS C.269811).
<typeStatus id="54DFA2D157653C39FB38FA3FBE25ACB4" box="[1194,1307,1421,1445]" pageId="29" pageNumber="130" type="paratype">Paratypes</typeStatus>
: Same data (AMS C.460381, 5 spms).
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857653C39FF55FA66BBDFAF6D" pageId="29" pageNumber="130" type="materials_examined">
<paragraph id="8BDB1C7357653C39FF55FA66B967ACFC" blockId="29.[151,1437,1311,2021]" box="[199,601,1492,1517]" pageId="29" pageNumber="130">
<emphasis id="B910C06157653C39FF55FA66BA8EACFC" bold="true" box="[199,432,1492,1517]" pageId="29" pageNumber="130">Material examined.</emphasis>
<typeStatus id="54DFA2D157653C39FE25FA67BAD1ACFC" box="[439,495,1493,1517]" pageId="29" pageNumber="130">Type</typeStatus>
material.
</paragraph>
<paragraph id="8BDB1C7357653C39FF55FA4ABBDFAF6D" blockId="29.[151,1437,1311,2021]" pageId="29" pageNumber="130">
<collectingCountry id="F3735CE357653C39FF55FA4ABA6DAF01" box="[199,339,1528,1552]" name="Hong Kong" pageId="29" pageNumber="130">Hong Kong</collectingCountry>
: Same data as
<typeStatus id="54DFA2D157653C39FD95FA4AB909AF01" box="[519,567,1528,1552]" pageId="29" pageNumber="130">type</typeStatus>
material (AMS C.460380, 20+ spms); Tsim Bei Tsui near Lau Fau Shan, seaward edge in mangroves,
<geoCoordinate id="EE507AB457653C39FE71F9AEB96FAF25" box="[483,593,1564,1588]" direction="north" orientation="latitude" pageId="29" pageNumber="130" precision="925" value="22.466667">22° 28' N</geoCoordinate>
,
<geoCoordinate id="EE507AB457653C39FDF8F9AEB9DAAF25" box="[618,740,1564,1589]" direction="east" orientation="longitude" pageId="29" pageNumber="130" precision="925" value="113.98333">113° 59' E</geoCoordinate>
,
<date id="FFDA3AB357653C39FD60F9AEB8B2AF25" box="[754,908,1564,1589]" pageId="29" pageNumber="130" value="1992-04-15">15 Apr. 1992</date>
. Coll. W.F. Ponder (AMS C.463401, 2 spms); Tsim Bei Tsui, Deep Bay, mangroves,
<geoCoordinate id="EE507AB457653C39FDDBF9F2B98BAF46" box="[585,693,1600,1624]" direction="north" orientation="latitude" pageId="29" pageNumber="130" precision="925" value="22.466667">22° 28' N</geoCoordinate>
,
<geoCoordinate id="EE507AB457653C39FD5EF9F2B87AAF46" box="[716,836,1599,1624]" direction="east" orientation="longitude" pageId="29" pageNumber="130" precision="925" value="113.98333">113° 59' E</geoCoordinate>
,
<date id="FFDA3AB357653C39FCC2F98DB8D6AF49" box="[848,1000,1599,1624]" pageId="29" pageNumber="130" value="1983-04-15">15 Apr. 1983</date>
. Coll. W.F. Ponder (AMS C.463405, 3 spms).
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857653C39FF55F935BE12AFB1" box="[199,1324,1671,1696]" pageId="29" pageNumber="130" type="distribution">
<paragraph id="8BDB1C7357653C39FF55F935BE12AFB1" blockId="29.[151,1437,1311,2021]" box="[199,1324,1671,1696]" pageId="29" pageNumber="130">
<emphasis id="B910C06157653C39FF55F935BA5EAFB1" bold="true" box="[199,352,1671,1696]" pageId="29" pageNumber="130">Distribution.</emphasis>
Known only from
<collectingCountry id="F3735CE357653C39FDA5F93AB983AFB1" box="[567,701,1672,1696]" name="Hong Kong" pageId="29" pageNumber="130">Hong Kong</collectingCountry>
, but is presumably more widespread in southern
<collectingCountry id="F3735CE357653C39FB70F935BE17AFB1" box="[1250,1321,1671,1696]" name="China" pageId="29" pageNumber="130">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857653C05FF55F91EB87BA82E" lastPageId="33" lastPageNumber="134" pageId="29" pageNumber="130" type="description">
<paragraph id="8BDB1C7357653C3AFF55F91EB8DEA9C4" blockId="29.[151,1437,1311,2021]" lastBlockId="30.[151,1436,151,213]" lastPageId="30" lastPageNumber="131" pageId="29" pageNumber="130">
<emphasis id="B910C06157653C39FF55F91EBA67AFD4" bold="true" box="[199,345,1708,1733]" pageId="29" pageNumber="130">Description.</emphasis>
<emphasis id="B910C06157653C39FECDF91EBAA8AFD4" box="[351,406,1708,1733]" italics="true" pageId="29" pageNumber="130">Shell</emphasis>
. Minute (up to
<quantity id="4C9CB19657653C39FDD1F91EB9A7AFD5" box="[579,665,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="29" pageNumber="130" unit="mm" value="1.9">1.9 mm</quantity>
in maximum diameter;
<tableCitation id="C6E629C857653C39FC31F91EB8C8AFD5" box="[931,1014,1708,1733]" captionStart="TABLE 3" captionStartId="13.[151,239,656,679]" captionText="TABLE 3. Shell dimensions of type specimens of species of Clenchiellidae." pageId="29" pageNumber="130">Table 3</tableCitation>
), spire flat (
<figureCitation id="135F00F657653C39FBEEF91EBFF0AFD4" box="[1148,1230,1708,1733]" captionStart="FIGURE 16" captionStartId="30.[151,250,1217,1239]" captionTargetBox="[214,1370,253,1203]" captionTargetId="figure@30.[214,1370,253,1203]" captionTargetPageId="30" captionText="FIGURE 16. Holotypes of new species of Coliracemata n. gen. (from left towards right: dorsal, lateral and ventral view respectively). A C, Coliracemata mortoni n. sp. (AMS C. 269811; shell diameter 1.82 mm); D F, Coliracemata katurana n. sp. (AMS C. 445414; shell diameter 1.06 mm); G I, Coliracemata clarkae n. sp. (AMS C. 462962; shell diameter 1.36 mm)." httpUri="https://zenodo.org/record/253240/files/figure.png" pageId="29" pageNumber="130">Figs 16</figureCitation>
AC and 17AC). Protoconch depressed to slightly elevated above spire, about 1.7 whorls, surface of protoconch I sculptured with about three well-spaced very fine spiral threads, terminated by weak varix; protoconch II of one whorl, smooth, terminated by weak varix. Teleoconch of about 1.6 convex whorls, sculptured with distinct, minutely wavy spiral lirae in middle of dorsal part of shell, inner third of dorsal whorls lacking distinct spirals, interspaces about twice width of lirae, about 45 lirae and a few finer spiral threads on outer side of whorl at end of penultimate whorl; finer, close-spaced commarginal growth lines intersect spirals, resulting in netted appearance where stronger spirals occur. Periphery evenly convex, spiral weak to subobsolete on peripheral area. Base evenly convex, with wavy spiral lirae, umbilicus wide (nearly half width of base), distinct spiral sculpture lacking in umbilicus. Sutures deeply impressed. Aperture near circular, with simple, slightly thickened peristome, external varix absent. Colour yellow-brown to dark brown due to periostracum and deposits; shell white.
</paragraph>
<caption id="DF1B4CFB57663C3AFF05FB73BEBBAC03" httpUri="https://zenodo.org/record/253240/files/figure.png" pageId="30" pageNumber="131" targetBox="[214,1370,253,1203]" targetPageId="30">
<paragraph id="8BDB1C7357663C3AFF05FB73BEBBAC03" blockId="30.[151,1436,1217,1298]" pageId="30" pageNumber="131">
<emphasis id="B910C06157663C3AFF05FB73BA1DADC6" bold="true" box="[151,291,1217,1239]" pageId="30" pageNumber="131">FIGURE 16.</emphasis>
Holotypes of new species of
<taxonomicName id="4C6467F057663C3AFDF1FB73B9D1ADC6" box="[611,751,1217,1239]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B910C06157663C3AFDF1FB73B9D1ADC6" box="[611,751,1217,1239]" italics="true" pageId="30" pageNumber="131">Coliracemata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57663C3AFD6AFB73B87DADC6" box="[760,835,1217,1239]" pageId="30" pageNumber="131" rank="genus">
<emphasis id="B910C06157663C3AFD6AFB73B87DADC6" bold="true" box="[760,835,1217,1239]" pageId="30" pageNumber="131">n. gen.</emphasis>
</taxonomicNameLabel>
(from left towards right: dorsal, lateral and ventral view respectively).
<emphasis id="B910C06157663C3AFEBAFB6DBA66ADE4" bold="true" box="[296,344,1247,1269]" pageId="30" pageNumber="131">AC</emphasis>
,
<taxonomicName id="4C6467F057663C3AFEF5FB6DB974ADE4" box="[359,586,1247,1269]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="mortoni" status="sp. nov.">
<emphasis id="B910C06157663C3AFEF5FB6DB974ADE4" box="[359,586,1247,1269]" italics="true" pageId="30" pageNumber="131">Coliracemata mortoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57663C3AFDC0FB6DB9B3ADE4" box="[594,653,1247,1269]" pageId="30" pageNumber="131" rank="species">
<emphasis id="B910C06157663C3AFDC0FB6DB9B3ADE4" bold="true" box="[594,653,1247,1269]" pageId="30" pageNumber="131">n. sp.</emphasis>
</taxonomicNameLabel>
(AMS C.269811; shell diameter 1.82 mm);
<emphasis id="B910C06157663C3AFBC5FB6DBFBDADE4" bold="true" box="[1111,1155,1247,1269]" pageId="30" pageNumber="131">DF</emphasis>
,
<taxonomicName id="4C6467F057663C3AFB03FB6DBEBEADE4" box="[1169,1408,1247,1269]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="katurana" status="sp. nov.">
<emphasis id="B910C06157663C3AFB03FB6DBEBEADE4" box="[1169,1408,1247,1269]" italics="true" pageId="30" pageNumber="131">Coliracemata katurana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57663C3AFA15FB6DBB8BAC03" pageId="30" pageNumber="131" rank="species">
<emphasis id="B910C06157663C3AFA15FB6DBB8BAC03" bold="true" pageId="30" pageNumber="131">n. sp.</emphasis>
</taxonomicNameLabel>
(AMS C.445414; shell diameter 1.06 mm);
<emphasis id="B910C06157663C3AFDE4FB4EB99EAC03" bold="true" box="[630,672,1276,1298]" pageId="30" pageNumber="131">GI</emphasis>
,
<taxonomicName id="4C6467F057663C3AFD3FFB4EB8B5AC03" box="[685,907,1276,1298]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="clarkae" status="sp. nov.">
<emphasis id="B910C06157663C3AFD3FFB4EB8B5AC03" box="[685,907,1276,1298]" italics="true" pageId="30" pageNumber="131">Coliracemata clarkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57663C3AFC03FB4EB8F4AC03" box="[913,970,1276,1298]" pageId="30" pageNumber="131" rank="species">
<emphasis id="B910C06157663C3AFC03FB4EB8F4AC03" bold="true" box="[913,970,1276,1298]" pageId="30" pageNumber="131">n. sp.</emphasis>
</taxonomicNameLabel>
(AMS C.462962; shell diameter 1.36 mm).
</paragraph>
</caption>
<caption id="DF1B4CFB57663C3AFF05F80AB835AEFC" httpUri="https://zenodo.org/record/253241/files/figure.png" pageId="30" pageNumber="131" targetBox="[208,1350,1354,1961]" targetPageId="30">
<paragraph id="8BDB1C7357663C3AFF05F80AB835AEFC" blockId="30.[151,1435,1976,2029]" pageId="30" pageNumber="131">
<emphasis id="B910C06157663C3AFF05F80ABA21AEDF" bold="true" box="[151,287,1976,1998]" pageId="30" pageNumber="131">FIGURE 17.</emphasis>
Shells of
<taxonomicName id="4C6467F057663C3AFE16F80AB95BAEDF" box="[388,613,1976,1998]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="mortoni">
<emphasis id="B910C06157663C3AFE16F80AB95BAEDF" box="[388,613,1976,1998]" italics="true" pageId="30" pageNumber="131">Coliracemata mortoni</emphasis>
</taxonomicName>
(AMS C.460381;
<emphasis id="B910C06157663C3AFCB2F80AB86FAEDF" bold="true" box="[800,849,1976,1998]" pageId="30" pageNumber="131">AC</emphasis>
) and
<taxonomicName id="4C6467F057663C3AFC18F80ABF46AEDF" box="[906,1144,1976,1998]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="131" phylum="Mollusca" rank="species" species="katurana">
<emphasis id="B910C06157663C3AFC18F80ABF46AEDF" box="[906,1144,1976,1998]" italics="true" pageId="30" pageNumber="131">Coliracemata katurana</emphasis>
</taxonomicName>
(AMS C.445414;
<emphasis id="B910C06157663C3AFAA0F80ABE5EAEDF" bold="true" box="[1330,1376,1976,1998]" pageId="30" pageNumber="131">DF</emphasis>
) with dorsal, lateral and ventral views given from left towards right.
</paragraph>
</caption>
<caption id="DF1B4CFB57673C3BFF05FDB4B82CAB2B" httpUri="https://zenodo.org/record/253242/files/figure.png" pageId="31" pageNumber="132" targetBox="[262,1329,199,489]" targetPageId="31">
<paragraph id="8BDB1C7357673C3BFF05FDB4B82CAB2B" blockId="31.[151,1436,518,570]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF05FDB4BA22AB0D" bold="true" box="[151,284,518,540]" pageId="31" pageNumber="132">FIGURE 18</emphasis>
. External and internal view (left to right) of opercula.
<emphasis id="B910C06157673C3BFCFAFDB4B89FAB0D" bold="true" box="[872,929,518,540]" pageId="31" pageNumber="132">A, B,</emphasis>
<taxonomicName id="4C6467F057673C3BFC3EFDB4BFAFAB0D" box="[940,1169,518,540]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="132" phylum="Mollusca" rank="species" species="mortoni">
<emphasis id="B910C06157673C3BFC3EFDB4BFAFAB0D" box="[940,1169,518,540]" italics="true" pageId="31" pageNumber="132">Coliracemata mortoni</emphasis>
</taxonomicName>
(AMS C.460381);
<emphasis id="B910C06157673C3BFAF3FDB4BEA2AB0D" bold="true" box="[1377,1436,518,540]" pageId="31" pageNumber="132">C, D,</emphasis>
<taxonomicName id="4C6467F057673C3BFF05FD96BABAAB2B" box="[151,388,548,570]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="132" phylum="Mollusca" rank="species" species="katurana">
<emphasis id="B910C06157673C3BFF05FD96BABAAB2B" box="[151,388,548,570]" italics="true" pageId="31" pageNumber="132">Coliracemata katurana</emphasis>
</taxonomicName>
(AMS C.445414). Scale bars: 100 µm.
</paragraph>
</caption>
<caption id="DF1B4CFB57673C3BFF05FA5DBE4FAF52" httpUri="https://zenodo.org/record/253243/files/figure.png" pageId="31" pageNumber="132" targetBox="[212,1360,615,1486]" targetPageId="31">
<paragraph id="8BDB1C7357673C3BFF05FA5DBE4FAF52" blockId="31.[151,1436,1519,1603]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF05FA5DBA1EAF14" bold="true" box="[151,288,1519,1541]" pageId="31" pageNumber="132">FIGURE 19.</emphasis>
Head-foot and gut characters of
<taxonomicName id="4C6467F057673C3BFDE2FA5DB86CAF17" box="[624,850,1519,1542]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="132" phylum="Mollusca" rank="species" species="mortoni">
<emphasis id="B910C06157673C3BFDE2FA5DB86CAF17" box="[624,850,1519,1542]" italics="true" pageId="31" pageNumber="132">Coliracemata mortoni</emphasis>
</taxonomicName>
(AMS C.460381).
<emphasis id="B910C06157673C3BFB8AFA5DBF0EAF14" bold="true" box="[1048,1072,1519,1541]" pageId="31" pageNumber="132">A,</emphasis>
head, dorsal view;
<emphasis id="B910C06157673C3BFB6AFA5DBE31AF14" bold="true" box="[1272,1295,1519,1541]" pageId="31" pageNumber="132">B,</emphasis>
posterior end of foot, dorsal view;
<emphasis id="B910C06157673C3BFEE6F9BCBAB3AF35" bold="true" box="[372,397,1550,1572]" pageId="31" pageNumber="132">C,</emphasis>
external view of intestine and rectum;
<emphasis id="B910C06157673C3BFCB8F9BCB87CAF35" bold="true" box="[810,834,1550,1572]" pageId="31" pageNumber="132">D,</emphasis>
stomach. Scale bars: 200 µm. Abbreviations: ac, anterior chamber; ct, cephalic tentacle; fp, faecal pellet; int, intestine; o, oesophagus; pc, posterior chamber; r, rectum; ss, style sac.
</paragraph>
</caption>
<paragraph id="8BDB1C7357673C3BFF55F9D8B9F9AFFF" blockId="31.[151,1437,1642,2027]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF55F9D8BA7AAF92" box="[199,324,1642,1667]" italics="true" pageId="31" pageNumber="132">Operculum</emphasis>
. Horny, near circular, of 45 slowly increasing whorls, each sculptured externally with fine, oblique wrinkles (
<figureCitation id="135F00F657673C3BFEFEF93CBAFFAFB6" box="[364,449,1678,1703]" captionStart="FIGURE 18" captionStartId="31.[151,250,518,540]" captionTargetBox="[262,1329,199,489]" captionTargetId="figure@31.[249,1338,193,496]" captionTargetPageId="31" captionText="FIGURE 18. External and internal view (left to right) of opercula. A, B, Coliracemata mortoni (AMS C. 460381); C, D, Coliracemata katurana (AMS C. 445414). Scale bars: 100 µm." httpUri="https://zenodo.org/record/253242/files/figure.png" pageId="31" pageNumber="132">Fig. 18</figureCitation>
A, B). Interior with narrow, raised edge to muscle attachment area close to weakly thickened columellar edge. Small white (calcareous?) sub-spiral projection in middle part, with thickened ridge between projection and outer edge of muscle scar.
</paragraph>
<paragraph id="8BDB1C7357673C3BFF55F948BEB8AE26" blockId="31.[151,1437,1642,2027]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF55F948BA07AE02" box="[199,313,1786,1811]" italics="true" pageId="31" pageNumber="132">Head-foot</emphasis>
. Similar to that of
<taxonomicName id="4C6467F057673C3BFD9EF948B83CAE02" box="[524,770,1786,1811]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="132" phylum="Mollusca" rank="species" species="clarkae">
<emphasis id="B910C06157673C3BFD9EF948B83CAE02" box="[524,770,1786,1811]" italics="true" pageId="31" pageNumber="132">Coliracemata clarkae</emphasis>
</taxonomicName>
but unpigmented except for dense-white spots on head and a few on basal half of tentacles and foot. Posterior end of foot tapering and rounded posteriorly (
<figureCitation id="135F00F657673C3BFB5CF8ACBE1DAE26" box="[1230,1315,1822,1847]" captionStart="FIGURE 19" captionStartId="31.[151,250,1519,1541]" captionTargetBox="[212,1360,615,1486]" captionTargetId="figure@31.[210,1367,611,1493]" captionTargetPageId="31" captionText="FIGURE 19. Head-foot and gut characters of Coliracemata mortoni (AMS C. 460381). A, head, dorsal view; B, posterior end of foot, dorsal view; C, external view of intestine and rectum; D, stomach. Scale bars: 200 µm. Abbreviations: ac, anterior chamber; ct, cephalic tentacle; fp, faecal pellet; int, intestine; o, oesophagus; pc, posterior chamber; r, rectum; ss, style sac." httpUri="https://zenodo.org/record/253243/files/figure.png" pageId="31" pageNumber="132">Figs 19</figureCitation>
and
<figureCitation id="135F00F657673C3BFAC8F8ADBE44AE26" box="[1370,1402,1823,1847]" captionStart="FIGURE 20" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[161,1427,204,1740]" captionTargetId="figure@32.[152,1435,193,1746]" captionTargetPageId="32" captionText="FIGURE 20. Cephalic and penial characters of Coliracemata mortoni. Mong Tseng Wai, Hong Kong (AMS C. 463401). A C, cephalic tentacle showing distinct compound cilia on distal tip; D, apocrine glands situated on penis; E, F, penis with clusters of apocrine glands." httpUri="https://zenodo.org/record/253244/files/figure.png" pageId="31" pageNumber="132">20</figureCitation>
).
</paragraph>
<paragraph id="8BDB1C7357673C3BFF55F8F0B917AE4A" blockId="31.[151,1437,1642,2027]" box="[199,553,1858,1883]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF55F8F0BA7DAE4A" box="[199,323,1858,1883]" italics="true" pageId="31" pageNumber="132">
<taxonomicName id="4C6467F057673C3BFF55F8F0BA7EAE4A" box="[199,320,1858,1883]" class="Bryopsida" family="Hypnaceae" genus="Ctenidium" kingdom="Plantae" order="Hypnales" pageId="31" pageNumber="132" phylum="Bryophyta" rank="genus">Ctenidium</taxonomicName>
.
</emphasis>
About 20 filaments.
</paragraph>
<paragraph id="8BDB1C7357673C3BFF55F8D4B88BAED6" blockId="31.[151,1437,1642,2027]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF55F8D4BA21AE6E" box="[199,287,1894,1919]" italics="true" pageId="31" pageNumber="132">
<taxonomicName id="4C6467F057673C3BFF55F8D4BA22AE6E" box="[199,284,1894,1919]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="31" pageNumber="132" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
.
</emphasis>
Typical of family. Cusp formulae variable on central and lateral teeth; 25+1+25, 12+1+14, 1417, 1214, median cusps of central and lateral teeth long (about twice as long [or slightly more] as adjacent cusps), narrow, pointed; other cusps on all teeth slender, pointed (
<figureCitation id="135F00F657673C3BFC8DF81CB84EAED6" box="[799,880,1966,1991]" captionStart="FIGURE 21" captionStartId="33.[151,250,1716,1738]" captionTargetBox="[150,1436,663,1683]" captionTargetId="figure@33.[137,1449,658,1694]" captionTargetPageId="33" captionText="FIGURE 21. Radulae of Coliracemata mortoni (AMS C. 460381) and Coliracemata katurana (AMS C. 445414). Coliracemata mortoni: A, central, lateral and marginal teeth; B, central and lateral teeth. Coliracemata katurana: C, central, lateral and marginal teeth; D, central and lateral teeth." httpUri="https://zenodo.org/record/253245/files/figure.png" pageId="31" pageNumber="132">Fig. 21</figureCitation>
A, B).
</paragraph>
<paragraph id="8BDB1C7357673C3BFF55F861BFC1AEFA" blockId="31.[151,1437,1642,2027]" box="[199,1279,2002,2027]" pageId="31" pageNumber="132">
<emphasis id="B910C06157673C3BFF55F861BBC6AEFB" box="[199,248,2003,2026]" italics="true" pageId="31" pageNumber="132">Gut.</emphasis>
Anterior oesophagus with two very strong folds; rectum with two tight loops (
<figureCitation id="135F00F657673C3BFBF8F860BF85AEFA" box="[1130,1211,2002,2027]" captionStart="FIGURE 19" captionStartId="31.[151,250,1519,1541]" captionTargetBox="[212,1360,615,1486]" captionTargetId="figure@31.[210,1367,611,1493]" captionTargetPageId="31" captionText="FIGURE 19. Head-foot and gut characters of Coliracemata mortoni (AMS C. 460381). A, head, dorsal view; B, posterior end of foot, dorsal view; C, external view of intestine and rectum; D, stomach. Scale bars: 200 µm. Abbreviations: ac, anterior chamber; ct, cephalic tentacle; fp, faecal pellet; int, intestine; o, oesophagus; pc, posterior chamber; r, rectum; ss, style sac." httpUri="https://zenodo.org/record/253243/files/figure.png" pageId="31" pageNumber="132">Fig. 19</figureCitation>
C, D).
</paragraph>
<caption id="DF1B4CFB57583C04FF05F95ABA00AE2A" httpUri="https://zenodo.org/record/253244/files/figure.png" pageId="32" pageNumber="133" targetBox="[161,1427,204,1740]" targetPageId="32">
<paragraph id="8BDB1C7357583C04FF05F95ABA00AE2A" blockId="32.[151,1436,1768,1851]" pageId="32" pageNumber="133">
<emphasis id="B910C06157583C04FF05F95ABA1EAFEF" bold="true" box="[151,288,1768,1790]" pageId="32" pageNumber="133">FIGURE 20.</emphasis>
Cephalic and penial characters of
<taxonomicName id="4C6467F057583C04FDEDF95AB85FAFEF" box="[639,865,1768,1790]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="133" phylum="Mollusca" rank="species" species="mortoni">
<emphasis id="B910C06157583C04FDEDF95AB85FAFEF" box="[639,865,1768,1790]" italics="true" pageId="32" pageNumber="133">Coliracemata mortoni</emphasis>
</taxonomicName>
. Mong Tseng Wai, Hong Kong (AMS C.463401).
<emphasis id="B910C06157583C04FAF6F95ABEA5AFEF" bold="true" box="[1380,1435,1768,1790]" pageId="32" pageNumber="133">AC,</emphasis>
cephalic tentacle showing distinct compound cilia on distal tip;
<emphasis id="B910C06157583C04FC84F8B4B811AE0D" bold="true" box="[790,815,1798,1820]" pageId="32" pageNumber="133">D,</emphasis>
apocrine glands situated on penis;
<emphasis id="B910C06157583C04FB1DF8B4BFFEAE0D" bold="true" box="[1167,1216,1798,1820]" pageId="32" pageNumber="133">E, F,</emphasis>
penis with clusters of apocrine glands.
</paragraph>
</caption>
<paragraph id="8BDB1C7357583C04FF55F8D6B890AED2" blockId="32.[151,1436,1890,2023]" pageId="32" pageNumber="133">
<emphasis id="B910C06157583C04FF55F8D6BA3BAE6A" box="[199,261,1892,1915]" italics="true" pageId="32" pageNumber="133">Penis</emphasis>
. Short, wide distally and proximally (
<figureCitation id="135F00F657583C04FD59F8D0B81BAE6A" box="[715,805,1890,1915]" captionStart="FIGURE 20" captionStartId="32.[151,250,1768,1790]" captionTargetBox="[161,1427,204,1740]" captionTargetId="figure@32.[152,1435,193,1746]" captionTargetPageId="32" captionText="FIGURE 20. Cephalic and penial characters of Coliracemata mortoni. Mong Tseng Wai, Hong Kong (AMS C. 463401). A C, cephalic tentacle showing distinct compound cilia on distal tip; D, apocrine glands situated on penis; E, F, penis with clusters of apocrine glands." httpUri="https://zenodo.org/record/253244/files/figure.png" pageId="32" pageNumber="133">Figs 20</figureCitation>
and
<figureCitation id="135F00F657583C04FCF4F8D1B8B8AE6A" box="[870,902,1891,1915]" captionStart="FIGURE 22" captionStartId="34.[151,250,1251,1273]" captionTargetBox="[182,1402,441,1239]" captionTargetId="figure@34.[182,1403,441,1241]" captionTargetPageId="34" captionText="FIGURE 22. Reproductive anatomy of Coliracemata mortoni (AMS C. 460381). A, male head and penis; B, penis, ventral view; C, penis showing separate groups of apocrine glands; D, female reproductive system shown from anterior (left) to posterior (right); E, female reproductive system shown from posterior (left) to anterior (right). Scale bars: 200 µm. Abbreviations: ag, albumen gland; bc, bursa copulatrix; cag, clusters of apocrine glands; cb, cut base of penis; cg, capsule gland; co, coiled oviduct; ct, cephalic tentacle; fa, female aperture; gg, groups of apocrine glands; gm, glandular mass; pd, penial duct; ppw, position of posterior wall of mantle cavity; sr, seminal receptacle." httpUri="https://zenodo.org/record/253246/files/figure.png" pageId="32" pageNumber="133">22</figureCitation>
). Five to six distinct clusters of many small, apocrine glands on dorsal edge of middle portion; proximal clusters large, distal ones close to tip small. Tip narrow to wide, blunt, with no distinct papilla. Penial duct slightly undulating.
</paragraph>
<paragraph id="8BDB1C7357583C05FF55F87CBAD7A80C" blockId="32.[151,1436,1890,2023]" lastBlockId="33.[151,1437,151,606]" lastPageId="33" lastPageNumber="134" pageId="32" pageNumber="133">
<emphasis id="B910C06157583C04FF55F87CBA1EAEF6" box="[199,288,1998,2023]" italics="true" pageId="32" pageNumber="133">Oviduct</emphasis>
. Albumen gland slightly shorter than capsule gland (
<figureCitation id="135F00F657583C04FCEAF87CB8F5AEF6" box="[888,971,1998,2023]" captionStart="FIGURE 22" captionStartId="34.[151,250,1251,1273]" captionTargetBox="[182,1402,441,1239]" captionTargetId="figure@34.[182,1403,441,1241]" captionTargetPageId="34" captionText="FIGURE 22. Reproductive anatomy of Coliracemata mortoni (AMS C. 460381). A, male head and penis; B, penis, ventral view; C, penis showing separate groups of apocrine glands; D, female reproductive system shown from anterior (left) to posterior (right); E, female reproductive system shown from posterior (left) to anterior (right). Scale bars: 200 µm. Abbreviations: ag, albumen gland; bc, bursa copulatrix; cag, clusters of apocrine glands; cb, cut base of penis; cg, capsule gland; co, coiled oviduct; ct, cephalic tentacle; fa, female aperture; gg, groups of apocrine glands; gm, glandular mass; pd, penial duct; ppw, position of posterior wall of mantle cavity; sr, seminal receptacle." httpUri="https://zenodo.org/record/253246/files/figure.png" pageId="32" pageNumber="133">Fig. 22</figureCitation>
D, E). Coiled oviduct narrow, with single S-shaped coil with indentations. Seminal receptacles short, about equal in size. Ventral channel gently curved. Bursa small, slender, lying on left side of oviduct gland, about 1/5 behind posterior wall of mantle cavity; posterior part slightly wider than anterior; extremely short bursal duct on left side of capsule gland. Genital opening simple, on anterior end of bursal duct.
</paragraph>
<paragraph id="8BDB1C7357593C05FF55FE9BB87BA82E" blockId="33.[151,1437,151,606]" box="[199,837,295,320]" pageId="33" pageNumber="134">
<emphasis id="B910C06157593C05FF55FE9BBA48A851" box="[199,374,297,320]" italics="true" pageId="33" pageNumber="134">Nervous system</emphasis>
. Similar to that of
<taxonomicName id="4C6467F057593C05FDDBFE95B800A851" box="[585,830,295,320]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="clarkae">
<emphasis id="B910C06157593C05FDDBFE95B800A851" box="[585,830,295,320]" italics="true" pageId="33" pageNumber="134">Coliracemata clarkae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857593C05FF55FEFEB9C8AB4F" pageId="33" pageNumber="134" type="discussion">
<paragraph id="8BDB1C7357593C05FF55FEFEBA8CAB09" blockId="33.[151,1437,151,606]" pageId="33" pageNumber="134">
<emphasis id="B910C06157593C05FF55FEFEBA05A874" bold="true" box="[199,315,332,357]" pageId="33" pageNumber="134">Remarks.</emphasis>
This species has a similar umbilicus and probably a lower spire than
<taxonomicName id="4C6467F057593C05FBC5FEFEBFCFA874" box="[1111,1265,332,357]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157593C05FBC5FEFEBFCFA874" box="[1111,1265,332,357]" italics="true" pageId="33" pageNumber="134">Coliracemata</emphasis>
</taxonomicName>
?
<taxonomicName id="4C6467F057593C05FA94FEFFBEA2A875" box="[1286,1436,333,356]" class="Gastropoda" family="Valvatidae" genus="Valvata" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="microscopica">
<emphasis id="B910C06157593C05FA94FEFFBEA2A875" box="[1286,1436,333,356]" italics="true" pageId="33" pageNumber="134">microscopica</emphasis>
</taxonomicName>
(see below) and is also similar to
<taxonomicName id="4C6467F057593C05FD83FEC2B99BA899" box="[529,677,367,392]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="katurana">
<emphasis id="B910C06157593C05FD83FEC2B99BA899" box="[529,677,367,392]" italics="true" pageId="33" pageNumber="134">Co. katurana</emphasis>
</taxonomicName>
in shell characters but is much larger.
<taxonomicName id="4C6467F057593C05FBC5FEC2BFE1A899" box="[1111,1247,368,392]" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="mortoni">
<emphasis id="B910C06157593C05FBC5FEC2BFE1A899" box="[1111,1247,368,392]" italics="true" pageId="33" pageNumber="134">Co. mortoni</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6467F057593C05FAE6FEC2BBD5A8BC" class="Gastropoda" family="Clenchiellidae" genus="Coliracemata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="clarkae">
<emphasis id="B910C06157593C05FAE6FEC2BBD5A8BC" italics="true" pageId="33" pageNumber="134">Co. clarkae</emphasis>
</taxonomicName>
in being larger, having more prominent spiral sculpture and a lower spire. The only other similar species found in the same general vicinity is
<taxonomicName id="4C6467F057593C05FDD7FE05B9C9A8C1" box="[581,759,439,464]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="minutissima">
<emphasis id="B910C06157593C05FDD7FE05B9C9A8C1" box="[581,759,439,464]" italics="true" pageId="33" pageNumber="134">Cl. minutissima</emphasis>
</taxonomicName>
, which differs most obviously in having a strong apertural varix. Anatomically it differs from the two congeneric species described herein in details of penial, and female genital tract morphology.
</paragraph>
<paragraph id="8BDB1C7357593C05FF55FD96B9C8AB4F" blockId="33.[151,1437,151,606]" pageId="33" pageNumber="134">
This species may possibly have been combined as part of Tongs (1986)
<taxonomicName id="4C6467F057593C05FC6FFD96BF40AB2C" box="[1021,1150,548,573]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157593C05FC6FFD96BF40AB2C" box="[1021,1150,548,573]" italics="true" pageId="33" pageNumber="134">Clenchiella</emphasis>
</taxonomicName>
sp. although she figured and described
<taxonomicName id="4C6467F057593C05FEAAFDF7B91FAB4C" box="[312,545,581,606]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="134" phylum="Mollusca" rank="species" species="varicosa" status="sp. nov.">
<emphasis id="B910C06157593C05FEAAFDF7B91FAB4C" box="[312,545,581,606]" italics="true" pageId="33" pageNumber="134">Clenchiella varicosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2237D1A57593C05FDBAFDF7B959AB4F" box="[552,615,581,606]" pageId="33" pageNumber="134" rank="species">
<emphasis id="B910C06157593C05FDBAFDF7B959AB4F" bold="true" box="[552,615,581,606]" pageId="33" pageNumber="134">n. sp.</emphasis>
</taxonomicNameLabel>
(see above).
</paragraph>
</subSubSection>
</treatment>
</document>