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<document id="9A8A54E8FF67D88C38F6AAF0155C7CEF" ID-DOI="10.1080/00222930400001574" ID-ISSN="1464-5262" ID-Zenodo-Dep="4657854" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617296447408" checkinUser="carolina" docAuthor="Noreña, Carolina, Damborenea, Cristina &amp; Brusa, Francisco" docDate="2005" docId="03BC5A4BFFB5FFD0FCB173DE0496FD9D" docLanguage="en" docName="JNATHIST.39.18.1457-1468.pdf" docOrigin="Journal of Natural History 39 (18)" docSource="http://dx.doi.org/10.1080/00222930400001574" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Philosyrtis rauli Noreña &amp; Damborenea &amp; Brusa 2005, n. sp." docType="treatment" docVersion="3" lastPageNumber="1466" masterDocId="FF852233FFB2FFDBFD4A7608073CFFDB" masterDocTitle="New freshwater interstitial Otoplanidae (Platyhelminthes: Proseriata) from the Paraná and Uruguay rivers, South America" masterLastPageNumber="1468" masterPageNumber="1457" pageNumber="1462" updateTime="1698899804078" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5A41ECD21F62331505014FB1F646025E">New freshwater interstitial Otoplanidae (Platyhelminthes: Proseriata) from the Paraná and Uruguay rivers, South America</mods:title>
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<mods:namePart id="8484A8C38D2B7B1232474F8DC45A33BE">Noreña, Carolina</mods:namePart>
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<mods:namePart id="9E002B2AEFC23D6BBDB6F4AD8CDF7681">Damborenea, Cristina</mods:namePart>
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<mods:namePart id="96E3FEFB930C8211634730211DC21239">Brusa, Francisco</mods:namePart>
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<mods:title id="D9043750C2C5383795B8FB3C6572C6FF">Journal of Natural History</mods:title>
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<mods:part id="32CBA80E0B35744D0DB8A86AEC31DD2F">
<mods:date id="03C6676A8602DE7BBF33C05A097AE724">2005</mods:date>
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<mods:number id="336CB52CCA1666EB58B7F77692193527">2005-09-30</mods:number>
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<mods:number id="A7908F70BE4FAFC3A48EADF9055CC569">39</mods:number>
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<mods:number id="52E27641C82F0E9E7808A74A55D705F3">18</mods:number>
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<mods:identifier id="063E18D7CE65D2D603C6F8A3D5550BFA" type="DOI">10.1080/00222930400001574</mods:identifier>
<mods:identifier id="97005FBB1842CA4EB92615219B3DDC44" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03BC5A4BFFB5FFD0FCB173DE0496FD9D" LSID="urn:lsid:plazi:treatment:03BC5A4BFFB5FFD0FCB173DE0496FD9D" httpUri="http://treatment.plazi.org/id/03BC5A4BFFB5FFD0FCB173DE0496FD9D" lastPageId="11" lastPageNumber="1466" pageId="7" pageNumber="1462">
<subSubSection id="C30FB8D6FFB5FFDCFCB173DE042BFA35" box="[507,791,1494,1518]" pageId="7" pageNumber="1462" type="nomenclature">
<paragraph id="8BAAEB5DFFB5FFDCFCB173DE042BFA35" blockId="7.[381,916,1431,1550]" box="[507,791,1494,1518]" pageId="7" pageNumber="1462">
<heading id="D0E25C31FFB5FFDCFCB173DE042BFA35" box="[507,791,1494,1518]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="1462" reason="2">
<taxonomicName id="4C1590DEFFB5FFDCFCB173DE05F0FA35" authority="Noreña &amp; Damborenea &amp; Brusa, 2005" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[507,716,1494,1518]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="7" pageNumber="1462" phylum="Platyhelminthes" rank="species" species="rauli" status="sp. nov.">
<emphasis id="B961374FFFB5FFDCFCB173DE05F0FA35" bold="true" box="[507,716,1494,1518]" italics="true" pageId="7" pageNumber="1462">Philosyrtis rauli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2528A34FFB5FFDCFF9C73DE042BFA35" box="[726,791,1494,1518]" pageId="7" pageNumber="1462" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C30FB8D6FFB5FFD0FF0773FE0496FD9D" lastPageId="11" lastPageNumber="1466" pageId="7" pageNumber="1462" type="description">
<paragraph id="8BAAEB5DFFB5FFDCFF0773FE05F9F9D5" blockId="7.[381,916,1431,1550]" box="[589,709,1526,1550]" pageId="7" pageNumber="1462">
(
<figureCitation id="132EF7D8FFB5FFDCFF1F73FE0580F9D5" box="[597,700,1526,1550]" captionStart="Figure 4" captionStartId="9.[125,185,1611,1630]" captionTargetBox="[208,1088,164,1572]" captionTargetId="figure-0@9.[208,1090,164,1575]" captionTargetPageId="9" captionText="Figure 4. Philosyrtis rauli. (a) Ventral view of the body. b, bristles; br, brain; co, copulatory organ, ov, ovary; ph, pharynx; rh, rhabdites; st, statocyst; t, testes; vi, vitellaria; vs, vesicula seminalis. (b) Details of the needles of the copulatory organ." figureDoi="http://doi.org/10.5281/zenodo.4657864" httpUri="https://zenodo.org/record/4657864/files/figure.png" pageId="7" pageNumber="1462">Figure 4</figureCitation>
)
</paragraph>
<paragraph id="8BAAEB5DFFB5FFDCFD37702407E8F99F" blockId="7.[125,212,1580,1604]" box="[125,212,1580,1604]" pageId="7" pageNumber="1462">
<emphasis id="B961374FFFB5FFDCFD37702407E8F99F" box="[125,212,1580,1604]" italics="true" pageId="7" pageNumber="1462">Locality</emphasis>
</paragraph>
<paragraph id="8BAAEB5DFFB5FFDCFD3770550427F94E" blockId="7.[125,1172,1629,1685]" pageId="7" pageNumber="1462">
<materialsCitation id="3B7DE100FFB5FFDCFD3770550413F9AE" box="[125,815,1629,1653]" country="Uruguay" location="Rio Negro Department" pageId="7" pageNumber="1462" specimenCount="1" stateProvince="Rio Negro">
<collectingCountry id="F302ABCDFFB5FFDCFD37705507DEF9AE" box="[125,226,1629,1653]" name="Uruguay" pageId="7" pageNumber="1462">Uruguay</collectingCountry>
river near Las Cañas,
<collectingRegion id="49D125BFFFB5FFDCFF4570550413F9AE" box="[527,815,1629,1653]" country="Uruguay" name="Rio Negro" pageId="7" pageNumber="1462">Rio Negro Department</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B7DE100FFB5FFDCFE097055042BF94E" collectingDate="November 2002, January and February 2003" country="Uruguay" location="Sublittoral" pageId="7" pageNumber="1462" specimenCount="1">
<collectingCountry id="F302ABCDFFB5FFDCFE0970550491F9AE" box="[835,941,1629,1653]" name="Uruguay" pageId="7" pageNumber="1462">Uruguay</collectingCountry>
.
<location id="8ECABD86FFB5FFDCFE8A70550302F9AE" LSID="urn:lsid:plazi:treatment:03BC5A4BFFB5FFD0FCB173DE0496FD9D:8ECABD86FFB5FFDCFE8A70550302F9AE" box="[960,1086,1629,1653]" country="Uruguay" name="Sublittoral" pageId="7" pageNumber="1462">Sublittoral</location>
, sandy sediments,
<collectingDate id="EFEF3475FFB5FFDCFC487075042BF94E" box="[258,791,1661,1685]" isEnumeration="true" pageId="7" pageNumber="1462">
<date id="FFABCD9DFFB5FFDCFC48707506FFF94E" box="[258,451,1661,1685]" pageId="7" pageNumber="1462" value="2002-11">November 2002</date>
,
<date id="FFABCD9DFFB5FFDCFC9A7075042BF94E" box="[464,791,1661,1685]" pageId="7" pageNumber="1462" value="2003-01" valueMax="2003-02-31" valueMin="2003-01-00">January and February 2003</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<caption id="DF6ABBD5FFBAFFD3FDC0706D0693F948" ID-DOI="http://doi.org/10.5281/zenodo.4657862" ID-Zenodo-Dep="4657862" httpUri="https://zenodo.org/record/4657862/files/figure.png" pageId="8" pageNumber="1463" startId="8.[138,198,1637,1656]" targetBox="[401,923,169,1601]" targetPageId="8">
<paragraph id="8BAAEB5DFFBAFFD3FDC0706D0693F948" blockId="8.[138,1185,1637,1683]" pageId="8" pageNumber="1463">
Figure 3.
<taxonomicName id="4C1590DEFFBAFFD3FDA3706D062AF9A3" authorityName="Marcus" authorityYear="1949" box="[233,278,1637,1656]" class="Rhabditophora" family="Otoplanidae" genus="Kata" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="1463" phylum="Platyhelminthes" rank="genus">
<emphasis id="B961374FFFBAFFD3FDA3706D062AF9A3" box="[233,278,1637,1656]" italics="true" pageId="8" pageNumber="1463">Kata</emphasis>
</taxonomicName>
sp. Ventral view of the body. b, bristles; br, brain; co, copulatory organ, ov, ovary; ph, pharynx; st, statocyst; t, testes; vi, vitellaria.
</paragraph>
</caption>
<caption id="DF6ABBD5FFBBFFD2FD377043061EF948" ID-DOI="http://doi.org/10.5281/zenodo.4657864" ID-Zenodo-Dep="4657864" httpUri="https://zenodo.org/record/4657864/files/figure.png" pageId="9" pageNumber="1464" startId="9.[125,185,1611,1630]" targetBox="[208,1088,164,1572]" targetPageId="9">
<paragraph id="8BAAEB5DFFBBFFD2FD377043061EF948" blockId="9.[125,1172,1611,1683]" pageId="9" pageNumber="1464">
Figure 4.
<taxonomicName id="4C1590DEFFBBFFD2FD9770430658F985" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[221,356,1611,1630]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="9" pageNumber="1464" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFBBFFD2FD9770430658F985" box="[221,356,1611,1630]" italics="true" pageId="9" pageNumber="1464">Philosyrtis rauli</emphasis>
</taxonomicName>
. (a) Ventral view of the body. b, bristles; br, brain; co, copulatory organ, ov, ovary; ph, pharynx; rh, rhabdites; st, statocyst; t, testes; vi, vitellaria; vs, vesicula seminalis. (b) Details of the needles of the copulatory organ.
</paragraph>
</caption>
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<emphasis id="B961374FFFB8FFD1FDC0768007D4FF7B" box="[138,232,136,160]" italics="true" pageId="10" pageNumber="1465">Material</emphasis>
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC076B10578FF2A" blockId="10.[138,1185,185,241]" pageId="10" pageNumber="1465">
Ten individuals (mature and immature) studied alive, six immature and two mature specimens sagittally sectioned (
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).
</paragraph>
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<emphasis id="B961374FFFB8FFD1FDC07722061EFE99" box="[138,290,298,322]" italics="true" pageId="10" pageNumber="1465">Type material</emphasis>
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<materialsCitation id="3B7DE100FFB8FFD1FDC07752059DFE49" country="Argentina" location="Helminthological Collection" municipality="Museo de La Plata" pageId="10" pageNumber="1465" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AE55FFFFB8FFD1FDC0775207C6FEA9" box="[138,250,346,370]" pageId="10" pageNumber="1465" type="holotype">Holotype</typeStatus>
: one sagittally sectioned specimen deposited in the
<location id="8ECABD86FFB8FFD1FE1A775203A1FEA9" LSID="urn:lsid:plazi:treatment:03BC5A4BFFB5FFD0FCB173DE0496FD9D:8ECABD86FFB8FFD1FE1A775203A1FEA9" box="[848,1181,346,370]" country="Argentina" municipality="Museo de La Plata" name="Helminthological Collection" pageId="10" pageNumber="1465">Helminthological Collection</location>
,
<collectingMunicipality id="6BCE7127FFB8FFD1FDC077720655FE49" box="[138,361,378,402]" pageId="10" pageNumber="1465">Museo de La Plata</collectingMunicipality>
,
<collectingCountry id="F302ABCDFFB8FFD1FC3F777206DBFE49" box="[373,487,378,402]" name="Argentina" pageId="10" pageNumber="1465">Argentina</collectingCountry>
(cat. no. 5313)
</materialsCitation>
.
<materialsCitation id="3B7DE100FFB8FFD1FFE6777203A1FE69" collectionCode="MNCN" country="Argentina" location="Museo Nacional de Ciencias Naturales de Madrid" pageId="10" pageNumber="1465" specimenCode="MNCN 4.01" specimenCount="1" typeStatus="paratype">
<typeStatus id="54AE55FFFFB8FFD1FFE67772042AFE49" box="[684,790,378,402]" pageId="10" pageNumber="1465" type="paratype">Paratype</typeStatus>
: one sagittally sectioned specimen deposited in the
<location id="8ECABD86FFB8FFD1FC0B7792044CFE69" LSID="urn:lsid:plazi:treatment:03BC5A4BFFB5FFD0FCB173DE0496FD9D:8ECABD86FFB8FFD1FC0B7792044CFE69" box="[321,880,410,434]" country="Argentina" name="Museo Nacional de Ciencias Naturales de Madrid" pageId="10" pageNumber="1465">Museo Nacional de Ciencias Naturales de Madrid</location>
(cat. no.
<specimenCode id="DBB34326FFB8FFD1FE9D77920351FE69" box="[983,1133,410,434]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="10" pageNumber="1465" type="Museum">MNCN 4.01</specimenCode>
/46)
</materialsCitation>
.
</paragraph>
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<emphasis id="B961374FFFB8FFD1FDC077E307C7FDD8" box="[138,251,491,515]" italics="true" pageId="10" pageNumber="1465">Etymology</emphasis>
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC07414054EFDEF" blockId="10.[138,626,540,564]" box="[138,626,540,564]" pageId="10" pageNumber="1465">This species is dedicated to Raúl Codina.</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC07464063FFD5F" blockId="10.[138,259,620,644]" box="[138,259,620,644]" pageId="10" pageNumber="1465">
<emphasis id="B961374FFFB8FFD1FDC07464063FFD5F" box="[138,259,620,644]" italics="true" pageId="10" pageNumber="1465">Description</emphasis>
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC07495031DFC6F" blockId="10.[138,1185,669,1172]" pageId="10" pageNumber="1465">
The living animals are
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long and about
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thick, fast-moving and strongly spastic. Colourless and transparent. The cephalic lobe is not clearly set off, but it is provided with long sensory bristles (
<figureCitation id="132EF7D8FFB8FFD1FF6D74D505A7FD2E" box="[551,667,733,757]" captionStart="Figure 4" captionStartId="9.[125,185,1611,1630]" captionTargetBox="[208,1088,164,1572]" captionTargetId="figure-0@9.[208,1090,164,1575]" captionTargetPageId="9" captionText="Figure 4. Philosyrtis rauli. (a) Ventral view of the body. b, bristles; br, brain; co, copulatory organ, ov, ovary; ph, pharynx; rh, rhabdites; st, statocyst; t, testes; vi, vitellaria; vs, vesicula seminalis. (b) Details of the needles of the copulatory organ." figureDoi="http://doi.org/10.5281/zenodo.4657864" httpUri="https://zenodo.org/record/4657864/files/figure.png" pageId="10" pageNumber="1465">Figure 4a</figureCitation>
). Adhesive papillae and bristles on the lateral side of the body at more or less regular distances, more abundant at the posterior end. Numerous needle-like rhabdites grouped behind the encapsulated brain, anterior to the pharynx and between the testes, forming two parallel rows. Isolated rhabdites over the entire body. Epidermis unciliated dorsally, ventrally well developed, ciliated and forming a creeping sole. Pharynx internally ciliated, before the middle of the body. Gut extending from the brain to the posterior end of the body, behind the vesicula seminalis.
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDEF75B50562FB4F" blockId="10.[138,1185,669,1172]" pageId="10" pageNumber="1465">
Ovaries situated at the basis of the pharynx. Two lateral rows of yolk follicles running from the pharynx to the posterior end. Only one pair of testicular follicles in adult animals, and up to three pairs in juveniles, situated just behind the brain and in front of the ovaries. Male copulatory apparatus consisting of an ovoid vesicula seminalis, the prostatic glands and the cuticular apparatus. The cuticular apparatus consisting of eight needles
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long and
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thick. The needles are proximally straight with an elongated, sickle-like terminal end (
<figureCitation id="132EF7D8FFB8FFD1FC6572740694FB4F" box="[303,424,1148,1172]" captionStart="Figure 4" captionStartId="9.[125,185,1611,1630]" captionTargetBox="[208,1088,164,1572]" captionTargetId="figure-0@9.[208,1090,164,1575]" captionTargetPageId="9" captionText="Figure 4. Philosyrtis rauli. (a) Ventral view of the body. b, bristles; br, brain; co, copulatory organ, ov, ovary; ph, pharynx; rh, rhabdites; st, statocyst; t, testes; vi, vitellaria; vs, vesicula seminalis. (b) Details of the needles of the copulatory organ." figureDoi="http://doi.org/10.5281/zenodo.4657864" httpUri="https://zenodo.org/record/4657864/files/figure.png" pageId="10" pageNumber="1465">Figure 4b</figureCitation>
). Bursa absent.
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC072B5032FFA8F" blockId="10.[138,1185,1213,1364]" pageId="10" pageNumber="1465">
<emphasis id="B961374FFFB8FFD1FDC072B5069CFB0E" box="[138,416,1213,1237]" italics="true" pageId="10" pageNumber="1465">
Biology.
<taxonomicName id="4C1590DEFFB8FFD1FDBB72B5069CFB0E" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[241,416,1213,1237]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="species" species="rauli">Philosyrtis rauli</taxonomicName>
</emphasis>
was found on a beach with sandy sediment near Las Cañas. According to the abiotic factors measured in the sample areas (
<tableCitation id="C697DEE6FFB8FFD1FECD72D504D8FB2E" box="[903,996,1245,1269]" captionStart="Table I" captionStartId="4.[138,192,1000,1019]" captionText="Table I. Temperature, pH, conductivity, TDS, salinity and dissolved oxygen values of sampled localities." pageId="10" pageNumber="1465">Table I</tableCitation>
), the ecological characterization is estimated as follows: rheobiotic, thermophilic (it is unquestionably a mesotypical species with thermophilic tendencies that depend on its adaptation to warm environments), stenobiotic (poly-oxibiotic), mesoionic, freshwater inhabitant.
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDC07375058CF9EF" blockId="10.[138,1185,1405,1685]" pageId="10" pageNumber="1465">
<emphasis id="B961374FFFB8FFD1FDC073750687FA4E" box="[138,443,1405,1429]" italics="true" pageId="10" pageNumber="1465">
Discussion.
<taxonomicName id="4C1590DEFFB8FFD1FC5A73750687FA4E" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[272,443,1405,1429]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="species" species="rauli">Philosyrtis rauli</taxonomicName>
</emphasis>
is a typical representative of the subfamily
<taxonomicName id="4C1590DEFFB8FFD1FEF5737507F3FA6E" authority="(Ax 1956)" baseAuthorityName="Ax" baseAuthorityYear="1956" class="Rhabditophora" family="Otoplanidae" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="subFamily" subFamily="Paratoplaninae">
Paratoplaninae (
<bibRefCitation id="EF8496ACFFB8FFD1F934737507FBFA6E" author="Ax P" pageId="10" pageNumber="1465" pagination="499 - 796" refId="ref4396" refString="Ax P. 1956. Monographie der Otoplanidae (Turbellaria). Morphologie und Systematik. Akademie der Wissenschaften und der Literatur in Mainz, Abhandlungen der Mathematisch-naturwissenschaftlichen Klasse 13: 499 - 796." type="journal article" year="1956">Ax 1956</bibRefCitation>
)
</taxonomicName>
and shares the following features with genus
<taxonomicName id="4C1590DEFFB8FFD1FFA87395046CFA6E" authorityName="Girad" authorityYear="1904" box="[738,848,1437,1461]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="genus">
<emphasis id="B961374FFFB8FFD1FFA87395046CFA6E" box="[738,848,1437,1461]" italics="true" pageId="10" pageNumber="1465">Philosyrtis</emphasis>
</taxonomicName>
: depressed epithelium in the regions of the creeping sole, dorsally unciliated epithelium; long, finger-like papillae; pharynx in the middle of the body; two rows of testicular follicles and paired ovary in front of the pharynx; two rows of yolk follicles behind the pharynx; cuticular apparatus with needles, but without stylet; absence of a bursa.
</paragraph>
<paragraph id="8BAAEB5DFFB8FFD1FDEF70350585F94E" blockId="10.[138,1185,1405,1685]" pageId="10" pageNumber="1465">
The genus
<taxonomicName id="4C1590DEFFB8FFD1FC63703506ABF98E" authorityName="Girad" authorityYear="1904" box="[297,407,1597,1621]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="genus">
<emphasis id="B961374FFFB8FFD1FC63703506ABF98E" box="[297,407,1597,1621]" italics="true" pageId="10" pageNumber="1465">Philosyrtis</emphasis>
</taxonomicName>
includes eight species, three of them occurring in South America:
<taxonomicName id="4C1590DEFFB8FFD1FDC070560532F9AE" authority="Ax and Ax, 1974" authorityName="Ax and Ax" authorityYear="1974" box="[138,526,1629,1653]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="species" species="sanjuanensis">
<emphasis id="B961374FFFB8FFD1FDC070560608F9AE" box="[138,308,1629,1653]" italics="true" pageId="10" pageNumber="1465">P. sanjuanensis</emphasis>
Ax and Ax, 1974
</taxonomicName>
,
<taxonomicName id="4C1590DEFFB8FFD1FF567056048DF9AE" authority="Ax and Ax, 1974" authorityName="Ax and Ax" authorityYear="1974" box="[540,945,1629,1653]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="species" species="santacruzensis">
<emphasis id="B961374FFFB8FFD1FF56705605E5F9AE" box="[540,729,1629,1653]" italics="true" pageId="10" pageNumber="1465">P. santacruzensis</emphasis>
Ax and Ax, 1974
</taxonomicName>
(
<collectingRegion id="49D125BFFFB8FFD1FE88705503A1F9AE" box="[962,1181,1629,1653]" country="Ecuador" name="Galapagos" pageId="10" pageNumber="1465">Galápagos Islands</collectingRegion>
,
<collectingCountry id="F302ABCDFFB8FFD1FDC0707507D3F94E" box="[138,239,1661,1685]" name="Ecuador" pageId="10" pageNumber="1465">Ecuador</collectingCountry>
) and
<taxonomicName id="4C1590DEFFB8FFD1FC7970760589F94E" authority="Marcus, 1950" authorityName="Marcus" authorityYear="1950" box="[307,693,1661,1685]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="10" pageNumber="1465" phylum="Platyhelminthes" rank="species" species="eumeca">
<emphasis id="B961374FFFB8FFD1FC7970760698F94E" box="[307,420,1662,1685]" italics="true" pageId="10" pageNumber="1465">P. eumeca</emphasis>
<bibRefCitation id="EF8496ACFFB8FFD1FCE77075056DF94E" author="Marcus E" box="[429,593,1661,1685]" pageId="10" pageNumber="1465" pagination="5 - 191" refId="ref4825" refString="Marcus E. 1950. Turbellaria Brasileiros (8). Boletins da Faculdade de Filosofia, Ciencias e Letras. Universidade de Sao Paulo. Zoologia 15: 5 - 191." type="journal article" year="1950">Marcus, 1950</bibRefCitation>
(
<collectingCountry id="F302ABCDFFB8FFD1FF2870750591F94E" box="[610,685,1661,1685]" name="Brazil" pageId="10" pageNumber="1465">Brazil</collectingCountry>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BAAEB5DFFB9FFD0FDD27681067BFE1F" blockId="11.[125,1173,137,582]" pageId="11" pageNumber="1466">
The main difference between
<taxonomicName id="4C1590DEFFB9FFD0FF4576810583FF7A" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[527,703,137,161]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FF4576810583FF7A" box="[527,703,137,161]" italics="true" pageId="11" pageNumber="1466">Philosyrtis rauli</emphasis>
</taxonomicName>
and other South American
<taxonomicName id="4C1590DEFFB9FFD0F96C768103A8FF7A" authorityName="Girad" authorityYear="1904" box="[1062,1172,137,161]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="genus">
<emphasis id="B961374FFFB9FFD0F96C768103A8FF7A" box="[1062,1172,137,161]" italics="true" pageId="11" pageNumber="1466">Philosyrtis</emphasis>
</taxonomicName>
species lies in the cuticular apparatus. Whereas
<taxonomicName id="4C1590DEFFB9FFD0FF9076A204B5FF1A" authorityName="Ax and Ax" authorityYear="1974" box="[730,905,169,193]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="sanjuanensis">
<emphasis id="B961374FFFB9FFD0FF9076A204B5FF1A" box="[730,905,169,193]" italics="true" pageId="11" pageNumber="1466">P. sanjuanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1590DEFFB9FFD0FE9876A203A8FF1A" authorityName="Ax and Ax" authorityYear="1974" box="[978,1172,169,193]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="santacruzensis">
<emphasis id="B961374FFFB9FFD0FE9876A203A8FF1A" box="[978,1172,169,193]" italics="true" pageId="11" pageNumber="1466">P. santacruzensis</emphasis>
</taxonomicName>
bear two groups of needles, one dorsal and one ventral, and
<taxonomicName id="4C1590DEFFB9FFD0FE1E76C204FAFF3A" authorityName="Marcus" authorityYear="1950" box="[852,966,202,225]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="eumeca">
<emphasis id="B961374FFFB9FFD0FE1E76C204FAFF3A" box="[852,966,202,225]" italics="true" pageId="11" pageNumber="1466">P. eumeca</emphasis>
</taxonomicName>
has three groups of needles with different structure and localization,
<taxonomicName id="4C1590DEFFB9FFD0FFBD76E20473FED9" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[759,847,234,258]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FFBD76E20473FED9" box="[759,847,234,258]" italics="true" pageId="11" pageNumber="1466">P. rauli</emphasis>
</taxonomicName>
shows a semicircle with a single
<typeStatus id="54AE55FFFFB9FFD0FD87770207C2FEF9" box="[205,254,266,290]" pageId="11" pageNumber="1466">type</typeStatus>
of needle. A similar cuticular apparatus can be found in
<taxonomicName id="4C1590DEFFB9FFD0FE857703061DFE99" authority="Sopott, 1972" authorityName="Sopott" authorityYear="1972" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rotundicephala">
<emphasis id="B961374FFFB9FFD0FE85770303A8FEF9" box="[975,1172,266,290]" italics="true" pageId="11" pageNumber="1466">P. rotundicephala</emphasis>
<bibRefCitation id="EF8496ACFFB9FFD0FD377722061DFE99" author="Sopott B" box="[125,289,298,322]" pageId="11" pageNumber="1466" pagination="167 - 236" refId="ref5087" refString="Sopott B. 1972. Systematik und Okologie der Proseriaten (Turbellaria) der deutschen Nordseekuste. Mikrofauna des Meeresbodens 13: 167 - 236." type="journal article" year="1972">Sopott, 1972</bibRefCitation>
</taxonomicName>
, but this species differs from
<taxonomicName id="4C1590DEFFB9FFD0FFE17723043BFE99" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[683,775,298,322]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FFE17723043BFE99" box="[683,775,298,322]" italics="true" pageId="11" pageNumber="1466">P. rauli</emphasis>
</taxonomicName>
in the number of needles (10 needles in
<taxonomicName id="4C1590DEFFB9FFD0FC49774306FBFEB8" authorityName="Sopott" authorityYear="1972" box="[259,455,331,355]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rotundicephala">
<emphasis id="B961374FFFB9FFD0FC49774306FBFEB8" box="[259,455,331,355]" italics="true" pageId="11" pageNumber="1466">P. rotundicephala</emphasis>
</taxonomicName>
, eight needles in
<taxonomicName id="4C1590DEFFB9FFD0FFED774305C3FEB8" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[679,767,331,355]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FFED774305C3FEB8" box="[679,767,331,355]" italics="true" pageId="11" pageNumber="1466">P. rauli</emphasis>
</taxonomicName>
) and in the shape of their distal end. Another difference between the two species is the number of testicular follicles, only one pair in
<taxonomicName id="4C1590DEFFB9FFD0FC24778406F2FE7F" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[366,462,396,420]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FC24778406F2FE7F" box="[366,462,396,420]" italics="true" pageId="11" pageNumber="1466">P. rauli</emphasis>
</taxonomicName>
(character shared with
<taxonomicName id="4C1590DEFFB9FFD0FE59778404B3FE78" authorityName="Marcus" authorityYear="1950" box="[787,911,396,419]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="eumeca">
<emphasis id="B961374FFFB9FFD0FE59778404B3FE78" box="[787,911,396,419]" italics="true" pageId="11" pageNumber="1466">P. eumeca</emphasis>
</taxonomicName>
) and four pairs in
<emphasis id="B961374FFFB9FFD0FD3777A5067BFE1F" box="[125,327,428,452]" italics="true" pageId="11" pageNumber="1466">
<taxonomicName id="4C1590DEFFB9FFD0FD3777A5067FFE1F" authorityName="Sopott" authorityYear="1972" box="[125,323,428,452]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rotundicephala">P. rotundicephala</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BAAEB5DFFB9FFD0FDD277C50496FD9D" blockId="11.[125,1173,137,582]" pageId="11" pageNumber="1466">
<taxonomicName id="4C1590DEFFB9FFD0FDD277C50693FE3E" authorityName="Sopott" authorityYear="1972" box="[152,431,461,485]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rotundicephala">
<emphasis id="B961374FFFB9FFD0FDD277C50693FE3E" box="[152,431,461,485]" italics="true" pageId="11" pageNumber="1466">Philosyrtis rotundicephala</emphasis>
</taxonomicName>
is a marine species from northern Europe, while
<taxonomicName id="4C1590DEFFB9FFD0FEBD77C60370FE3E" authorityName="Noreña &amp; Damborenea &amp; Brusa" authorityYear="2005" box="[1015,1100,461,485]" class="Rhabditophora" family="Otoplanidae" genus="Philosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="rauli">
<emphasis id="B961374FFFB9FFD0FEBD77C60370FE3E" box="[1015,1100,461,485]" italics="true" pageId="11" pageNumber="1466">P. rauli</emphasis>
</taxonomicName>
is the only freshwater species of the genus, from interstitial habitats of the
<collectingCountry id="F302ABCDFFB9FFD0FED977E504C4FDDE" box="[915,1016,493,517]" name="Uruguay" pageId="11" pageNumber="1466">Uruguay</collectingCountry>
river. Within the
<taxonomicName id="4C1590DEFFB9FFD0FDFA74060662FDFD" baseAuthorityName="Ax" baseAuthorityYear="1956" box="[176,350,526,550]" class="Rhabditophora" family="Otoplanidae" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1465" phylum="Platyhelminthes" rank="subFamily" subFamily="Paratoplaninae">Paratoplaninae</taxonomicName>
only three species of
<taxonomicName id="4C1590DEFFB9FFD0FF31740605C0FDFD" box="[635,764,526,550]" class="Rhabditophora" family="Otoplanidae" genus="Pseudosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="genus">
<emphasis id="B961374FFFB9FFD0FF31740605C0FDFD" box="[635,764,526,550]" italics="true" pageId="11" pageNumber="1466">Pseudosyrtis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1590DEFFB9FFD0FE5F74060484FDFD" authorityName="Ax" authorityYear="1956" baseAuthorityName="Ax" baseAuthorityYear="1951" box="[789,952,526,550]" class="Rhabditophora" family="Otoplanidae" genus="Pseudosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="subterranea">
<emphasis id="B961374FFFB9FFD0FE5F74060484FDFD" box="[789,952,526,550]" italics="true" pageId="11" pageNumber="1466">P. subterranea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1590DEFFB9FFD0FE8574060366FDFD" authorityName="Ax" authorityYear="1956" baseAuthorityName="Gieysztor" baseAuthorityYear="1938" box="[975,1114,526,550]" class="Rhabditophora" family="Otoplanidae" genus="Pseudosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="fluviatilis">
<emphasis id="B961374FFFB9FFD0FE8574060366FDFD" box="[975,1114,526,550]" italics="true" pageId="11" pageNumber="1466">P. fluviatilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1590DEFFB9FFD0FD3774270609FD9D" authorityName="Riemann" authorityYear="1965" box="[125,309,558,582]" class="Rhabditophora" family="Otoplanidae" genus="Pseudosyrtis" kingdom="Animalia" order="Seriata" pageId="11" pageNumber="1466" phylum="Platyhelminthes" rank="species" species="neiswestnovae">
<emphasis id="B961374FFFB9FFD0FD3774270609FD9D" box="[125,309,558,582]" italics="true" pageId="11" pageNumber="1466">P. neiswestnovae</emphasis>
</taxonomicName>
) have been recorded for brackish/freshwater habitats.
</paragraph>
</subSubSection>
</treatment>
</document>