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<document id="DFAB9451110473C3920801CC59734D8D" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7839FFA2FEECFE9E98BFFEA1" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra granodeoro Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="46" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="45" updateTime="1698190273211" updateUser="plazi">
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<mods:title id="A8008B18E97FD15F197B1B819C2790D2">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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<mods:name id="12B267E59B31FB86D76099E1D96F0299" type="personal">
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<mods:namePart id="E978E27B4DA4863A5CD19E8C1DDF98A0">Erwin, Terry L.</mods:namePart>
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<mods:title id="34BDC7F48BAEFAF529289F27A0E1AAE9">Zootaxa</mods:title>
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<mods:date id="738705677611C923A6D79F5CC272AE59">2002</mods:date>
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<mods:identifier id="3E67D9E6C9FE40494A81B5829FE42255" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<treatment id="03A287CA7839FFA2FEECFE9E98BFFEA1" ID-DOI="http://doi.org/10.5281/zenodo.6278003" ID-GBIF-Taxon="121065200" ID-Zenodo-Dep="6278003" LSID="urn:lsid:plazi:treatment:03A287CA7839FFA2FEECFE9E98BFFEA1" httpUri="http://treatment.plazi.org/id/03A287CA7839FFA2FEECFE9E98BFFEA1" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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<paragraph id="8BB436DC7839FFA3FEECFE9E9EF8FECA" blockId="44.[264,716,283,350]" box="[264,716,283,310]" pageId="44" pageNumber="45">
<heading id="D0FC81B07839FFA3FEECFE9E9EF8FECA" bold="true" box="[264,716,283,310]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B97FEACE7839FFA3FEECFE9E9EF8FECA" bold="true" box="[264,716,283,310]" pageId="44" pageNumber="45">
<taxonomicName id="4C0B4D5F7839FFA3FEECFE9E9E1BFECA" ID-CoL="B2NH" authority="Erwin" authorityName="Erwin" box="[264,559,283,310]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="granodeoro" status="sp. nov.">
<emphasis id="B97FEACE7839FFA3FEECFE9E9DECFEC9" bold="true" box="[264,472,284,310]" italics="true" pageId="44" pageNumber="45">Agra granodeoro</emphasis>
Erwin
</taxonomicName>
,
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<paragraph id="8BB436DC7839FFA3FEECFEC69DCFFEA1" blockId="44.[264,716,283,350]" box="[264,507,324,350]" pageId="44" pageNumber="45">
(
<figureCitation id="13302A597839FFA3FEF5FEC69D9EFEA1" box="[273,426,324,350]" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="44" pageNumber="45">Figs. 19a, 19</figureCitation>
b, 19c)
</paragraph>
</subSubSection>
<subSubSection id="C31165577839FFA3FEECFE119804FE29" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BB436DC7839FFA3FEECFE119804FE29" blockId="44.[264,1324,403,1030]" pageId="44" pageNumber="45">
<bibRef id="52ABA22A7839FFA3FEECFE119804FE29" pageId="44" pageNumber="45" type="journal article">
<emphasis id="B97FEACE7839FFA3FEECFE119DB5FE52" bold="true" box="[264,385,403,429]" pageId="44" pageNumber="45">
<title id="F90FDDEA7839FFA3FEECFE119D48FE52" box="[264,380,403,429]" jopScore="0.15499471" pageId="44" pageNumber="45" type="_" typeScore="3">
<typeStatus id="54B0887E7839FFA3FEECFE119D48FE52" box="[264,380,403,429]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</title>
.
</emphasis>
<author id="53B6DE397839FFA3FE6AFE169EA8FE52" box="[398,668,403,430]" fns="I" npo="LF" pageId="44" pageNumber="45">
Male,
<emphasis id="B97FEACE7839FFA3FE3AFE119EA8FE52" bold="true" box="[478,668,403,429]" pageId="44" pageNumber="45">
<collectingCountry id="F31C764C7839FFA3FE3AFE119EA8FE52" box="[478,668,403,429]" name="Costa Rica" pageId="44" pageNumber="45">COSTA RICA</collectingCountry>
</emphasis>
</author>
:
<journalOrPublisher id="51CA3C107839FFA3FD4AFE1698F7FE51" box="[686,1219,404,430]" jopScore="3.6123679" pageId="44" pageNumber="45">Cartago, Turrialba, Chirripo, Grano de Oro</journalOrPublisher>
,
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<part id="FFAF9C417839FFA3FB37FE16993AFE51" box="[1235,1294,404,430]" pageId="44" pageNumber="45" type="volume">1120</part>
m
</quantity>
,
<geoCoordinate id="EE3F501B7839FFA3FEECFE3E9D5DFE29" box="[264,361,444,470]" direction="north" orientation="latitude" pageId="44" pageNumber="45" precision="925" value="9.883333">
09°
<pagination id="B2C2C8287839FFA3FED5FE3E9D7BFE29" box="[305,335,444,470]" pageId="44" pageNumber="45" type="pagination">53</pagination>
'N
</geoCoordinate>
<geoCoordinate id="EE3F501B7839FFA3FE94FE3E9DE3FE29" box="[368,471,444,470]" direction="west" orientation="longitude" pageId="44" pageNumber="45" precision="925" value="-83.63333">83°38'W</geoCoordinate>
, September, (P. Campos)(
<collectionCode id="ED1AAE197839FFA3FCECFE3E9F61FE29" box="[776,853,444,470]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="45">INBio</collectionCode>
:
<collectionCode id="ED1AAE197839FFA3FC84FE3E9FA7FE29" LSID="urn:lsid:biocol.org:col:14323" box="[864,915,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="44" pageNumber="45">CRI</collectionCode>
000­935857).
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C31165577839FFA3FEDCFE6198B1FDD9" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8BB436DC7839FFA3FEDCFE6198B1FDD9" blockId="44.[264,1324,403,1030]" pageId="44" pageNumber="45">
<emphasis id="B97FEACE7839FFA3FEDCFE619D84FE02" bold="true" box="[312,432,483,509]" pageId="44" pageNumber="45">Diagnosis</emphasis>
. This is the smallest member of the palmata species­group in
<collectingCountry id="F31C764C7839FFA3FB43FE66991EFE01" box="[1191,1322,484,510]" name="Costa Rica" pageId="44" pageNumber="45">Costa Rica</collectingCountry>
and the only species with a narrow, tapered prothorax, black knees and tarsi.
</paragraph>
</subSubSection>
<subSubSection id="C31165577839FFA3FEDCFDB19FB1FC99" pageId="44" pageNumber="45" type="description">
<paragraph id="8BB436DC7839FFA3FEDCFDB19FB1FC99" blockId="44.[264,1324,403,1030]" pageId="44" pageNumber="45">
<emphasis id="B97FEACE7839FFA3FEDCFDB19DFDFDB2" bold="true" box="[312,457,563,589]" pageId="44" pageNumber="45">Description</emphasis>
. Color and luster: Piceous. Legs rufotestaceous with black knees and tarsi. Antennomeres 1­2 and apex of 3­10 infuscated. Female unknown. Form: Head (
<figureCitation id="13302A597839FFA3FB18FDDE9D07FD61" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="44" pageNumber="45">Fig. 19a</figureCitation>
) behind eyes moderately long, sides tapered to abruptly obtuse angles. Pronotum widest at base, markedly narrowed just posterior to apical collar; disc sparsely punctate. Elytron with apex (
<figureCitation id="13302A597839FFA3FE0CFD569E08FD11" box="[488,572,724,750]" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="44" pageNumber="45">Fig. 19</figureCitation>
b) markedly oblique, finely dentate suturally, sutural margin sinuate. Aedeagus (
<figureCitation id="13302A597839FFA3FE31FD7E9E1BFCE9" box="[469,559,764,790]" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="44" pageNumber="45">Fig. 19</figureCitation>
c) with two ostial membranes separated by a wide sclerotized bridge, endophallus without vestiture or armature; apex of phallus diamond­shaped. Size: moderately large,
<quantity id="4CF39B397839FFA3FE38FCCE9E71FC99" box="[476,581,844,870]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="44" pageNumber="45" unit="mm" value="18.1">18.1 mm</quantity>
in length,
<quantity id="4CF39B397839FFA3FD27FCCE9F21FC99" box="[707,789,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="44" pageNumber="45" unit="mm" value="4.4">4.4mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C31165577839FFA3FEDCFCF19D94FBF9" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BB436DC7839FFA3FEDCFCF19889FC71" blockId="44.[264,1324,403,1030]" box="[312,1213,883,910]" pageId="44" pageNumber="45">
<emphasis id="B97FEACE7839FFA3FEDCFCF19EA4FC72" bold="true" box="[312,656,883,909]" pageId="44" pageNumber="45">Other specimens examined.</emphasis>
The male
<typeStatus id="54B0887E7839FFA3FCE9FCF69F4EFC71" box="[781,890,884,910]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
is the only specimen I saw.
</paragraph>
<paragraph id="8BB436DC7839FFA3FEDCFC199D94FBF9" blockId="44.[264,1324,403,1030]" pageId="44" pageNumber="45">
<emphasis id="B97FEACE7839FFA3FEDCFC199DCAFC4A" bold="true" box="[312,510,923,949]" pageId="44" pageNumber="45">Specific epithet.</emphasis>
The specific epithet,
<emphasis id="B97FEACE7839FFA3FD1CFC1E9FBDFC4A" box="[760,905,924,949]" italics="true" pageId="44" pageNumber="45">
<taxonomicName id="4C0B4D5F7839FFA3FD1CFC1E9FB0FC4A" box="[760,900,924,949]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="granodeoro">granodeoro</taxonomicName>
,
</emphasis>
is used as a noun in apposition, and refers in Spanish to a &quot;golden seed,&quot; in reference to the brassy glint of the elytra and to the
<typeStatus id="54B0887E7839FFA3FEECFC6E9D0EFBF9" box="[264,314,1004,1030]" pageId="44" pageNumber="45">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF7466547839FFA3FEECF97D9D04F8A3" httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="44" pageNumber="45" targetBox="[406,1178,1107,1755]" targetPageId="44">
<paragraph id="8BB436DC7839FFA3FEECF97D9D04F8A3" blockId="44.[264,1324,1791,1884]" pageId="44" pageNumber="45">
<emphasis id="B97FEACE7839FFA3FEECF97D9DADF8E8" bold="true" box="[264,409,1791,1815]" pageId="44" pageNumber="45">FIGURE 19.</emphasis>
<taxonomicName id="4C0B4D5F7839FFA3FE46F8829E0AF8E8" box="[418,574,1792,1815]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="granodeoro">
<emphasis id="B97FEACE7839FFA3FE46F8829E0AF8E8" box="[418,574,1792,1815]" italics="true" pageId="44" pageNumber="45">A. granodeoro</emphasis>
</taxonomicName>
, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0.
</paragraph>
</caption>
<subSubSection id="C31165577838FFA2FEDCFE9998BFFEA1" pageId="45" pageNumber="46" type="description">
<paragraph id="8BB436DC7838FFA2FEDCFE9998BFFEA1" blockId="45.[264,1321,283,350]" pageId="45" pageNumber="46">
<emphasis id="B97FEACE7838FFA2FEDCFE999D4AFECA" bold="true" box="[312,382,283,309]" pageId="45" pageNumber="46">Notes</emphasis>
. The
<typeStatus id="54B0887E7838FFA2FE27FE9E9E04FEC9" box="[451,560,284,310]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
is from the Cordillera Volcánica Central Conservation Area. An illustration of palmata­group stylomere 2 is on the
<taxonomicName id="4C0B4D5F7838FFA2FCB8FEC69839FEA2" box="[860,1037,324,349]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="atriperna">
<emphasis id="B97FEACE7838FFA2FCB8FEC69839FEA2" box="[860,1037,324,349]" italics="true" pageId="45" pageNumber="46">Agra atriperna</emphasis>
</taxonomicName>
web page.
</paragraph>
</subSubSection>
</treatment>
</document>