treatments-xml/data/03/A2/87/03A287C10D21FFC8FF1AAD83FE16FD2B.xml

<document ID-DOI="10.11646/zootaxa.5263.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804383" ID-ZooBank="5BE2011B-8F58-415C-B788-EBFB8E5B6DB1" checkinTime="1680760970283" checkinUser="plazi" docAuthor="Mendes, Diego Matheus De Mello, Sobral, Rafael &amp; Neto, Alberto Moreira Da Silva" docDate="2023" docId="03A287C10D21FFC8FF1AAD83FE16FD2B" docLanguage="en" docName="zootaxa.5263.3.7.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megatympanon speculatum Piza 1958" docType="treatment" docVersion="2" lastPageNumber="435" masterDocId="FF9BFFB90D23FFCDFF8DAE7CFF9EFFAE" masterDocTitle="The Austral Predator Katydid-new species of Megatympanon Piza, 1958 (Orthoptera: Tettigoniidae: Listroscelidinae: Terpandrini) from the Brazilian Atlantic Rainforest" masterLastPageNumber="440" masterPageNumber="430" pageNumber="432" updateTime="1680761394069" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>The Austral Predator Katydid-new species of Megatympanon Piza, 1958 (Orthoptera: Tettigoniidae: Listroscelidinae: Terpandrini) from the Brazilian Atlantic Rainforest</mods:title>
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<mods:namePart>Mendes, Diego Matheus De Mello</mods:namePart>
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<mods:namePart>Sobral, Rafael</mods:namePart>
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<mods:affiliation>Instituto Nacional de Pesquisas da Amazônia, INPA, Programa de Pós-graduação em Entomologia, Caixa Postal 2223, 69080 - 971, Manaus, Amazonas, Brazil. rafaelsobralves @ gmail. com; https: // orcid. org / 0000 - 0002 - 4975 - 1193</mods:affiliation>
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<mods:namePart>Neto, Alberto Moreira Da Silva</mods:namePart>
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<mods:affiliation>Instituto Nacional de Pesquisas da Amazônia, INPA, Programa de Pós-graduação em Entomologia, Caixa Postal 2223, 69080 - 971, Manaus, Amazonas, Brazil. rafaelsobralves @ gmail. com; https: // orcid. org / 0000 - 0002 - 4975 - 1193 &amp; Universidade Estadual de Feira de Santana, 44036 - 900, Feira de Santana, Bahia, Brazil.</mods:affiliation>
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<subSubSection box="[151,621,1022,1049]" pageId="2" pageNumber="432" type="nomenclature">
<paragraph blockId="2.[151,621,1022,1084]" box="[151,621,1022,1049]" pageId="2" pageNumber="432">
<heading bold="true" box="[151,621,1022,1049]" fontSize="11" level="1" pageId="2" pageNumber="432" reason="1">
<taxonomicName ID-CoL="72XM2" authority="Piza, 1958" authorityName="Piza" authorityYear="1958" box="[151,621,1022,1049]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis bold="true" box="[151,621,1022,1049]" pageId="2" pageNumber="432">
<emphasis bold="true" box="[151,487,1022,1049]" italics="true" pageId="2" pageNumber="432">Megatympanon speculatum</emphasis>
<bibRefCitation author="Piza Jr., S. de" box="[493,621,1022,1048]" pageId="2" pageNumber="432" pagination="237 - 241" refId="ref4958" refString="Piza Jr., S. de T. (1958) A primeira especie neotropical da familia &quot; Tympanophoridae &quot; (Orthoptera, Tettigonioidea). Revista Brasileira de Biologia, 18 (2), 237 - 241." type="journal article" year="1958">Piza, 1958</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,353,1058,1084]" pageId="2" pageNumber="432" type="description">
<paragraph blockId="2.[151,621,1022,1084]" box="[151,353,1058,1084]" pageId="2" pageNumber="432">
<figureCitation box="[151,285,1058,1084]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[152,255,916,940]" captionStartId-1="3.[152,255,1896,1920]" captionStartId-2="4.[152,255,755,779]" captionStartId-3="4.[152,255,2002,2026]" captionTargetBox-0="[209,1366,240,890]" captionTargetBox-1="[166,1420,181,1873]" captionTargetBox-2="[173,1414,181,731]" captionTargetBox-3="[162,1426,875,1979]" captionTargetId-0="figure-26@2.[209,1378,235,890]" captionTargetId-1="figure-31@3.[166,1420,181,1873]" captionTargetId-2="figure-19@4.[173,1414,181,731]" captionTargetId-3="figure-69@4.[162,1426,875,1979]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURE 1. Pictorial key for species of Megatympanon Piza, 1958." captionText-1="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." captionText-2="FIGURE 3. Megatympanon speculatum, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein." captionText-3="FIGURE 4. Megatympanon speculatum, stridulatory file of male. A: left file; B: right file." figureDoi-0="http://doi.org/10.5281/zenodo.7804387" figureDoi-1="http://doi.org/10.5281/zenodo.7804389" figureDoi-2="http://doi.org/10.5281/zenodo.7804393" figureDoi-3="http://doi.org/10.5281/zenodo.7804395" httpUri-0="https://zenodo.org/record/7804387/files/figure.png" httpUri-1="https://zenodo.org/record/7804389/files/figure.png" httpUri-2="https://zenodo.org/record/7804393/files/figure.png" httpUri-3="https://zenodo.org/record/7804395/files/figure.png" pageId="2" pageNumber="432">Figures 1–4</figureCitation>
and 
<figureCitation box="[339,353,1058,1084]" captionStart="FIGURE 9" captionStartId="9.[152,255,1006,1030]" captionTargetBox="[151,1436,181,982]" captionTargetId="figure-31@9.[151,1436,181,982]" captionTargetPageId="9" captionText="FIGURE 9. Map of Megatympanon species geographical records. Area of the Serra do Mar coastal forests modified from WWF." figureDoi="http://doi.org/10.5281/zenodo.7804408" httpUri="https://zenodo.org/record/7804408/files/figure.png" pageId="2" pageNumber="432">9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="432" type="reference_group">
<paragraph blockId="2.[151,1436,1121,1176]" pageId="2" pageNumber="432">
<taxonomicName authority="Piza, 1958" authorityName="Piza" authorityYear="1958" box="[151,546,1121,1145]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[151,430,1121,1145]" italics="true" pageId="2" pageNumber="432">Megatympanon speculatum</emphasis>
<bibRefCitation author="Piza Jr., S. de" box="[436,546,1121,1145]" pageId="2" pageNumber="432" pagination="237 - 241" refId="ref4958" refString="Piza Jr., S. de T. (1958) A primeira especie neotropical da familia &quot; Tympanophoridae &quot; (Orthoptera, Tettigonioidea). Revista Brasileira de Biologia, 18 (2), 237 - 241." type="journal article" year="1958">Piza, 1958</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation author="Riek, E. F." box="[556,670,1121,1145]" pageId="2" pageNumber="432" pagination="161 - 171" refId="ref4999" refString="Riek, E. F. (1976) The Australian genus Tympanophora White (Orthoptera: Tettigoniidae: Tympanophorinae). Australian Journal of Entomology, 15, 161 - 171. https: // doi. org / 10.1111 / j. 1440 - 6055.1976. tb 01687. x" type="journal article" year="1976">Riek, 1976</bibRefCitation>
; 
<bibRefCitation author="Paschoal, A. D. &amp; Barros, O. N. F." box="[680,935,1121,1146]" pageId="2" pageNumber="432" pagination="237 - 251" refId="ref4901" refString="Paschoal, A. D. &amp; Barros, O. N. F. (1977) Catalogo dos tipos depositados no Museu de Zoologia da Escola Superior de Agricultura &quot; Luiz de Queiroz &quot;, I: Insecta. Revista de Agricultura (Piracicaba), 52 (4), 237 - 251." type="journal article" year="1977">Paschoal &amp; Barros, 1977</bibRefCitation>
; 
<bibRefCitation author="Rentz, D. C. F." box="[945,1070,1122,1146]" pageId="2" pageNumber="432" refId="ref5106" refString="Rentz, D. C. F. (2001) The Listroscelidinae, Tympanophorinae, Meconematinae and Microtettigoniinae. A Monograph of the Tettigoniidae of Australia. Vol. 3. CSIRO Publishing, Melbourne, 524 pp. https: // doi. org / 10.1071 / 9780643105324" type="book" year="2001">Rentz, 2001</bibRefCitation>
; 
<bibRefCitation author="Chamorro-Rengifo, J. &amp; Braun, H." box="[1080,1433,1121,1146]" pageId="2" pageNumber="432" pagination="41 - 66" refId="ref4455" refString="Chamorro-Rengifo, J. &amp; Braun, H. (2010) The Tettigoniidae (Orthoptera) described by Salvador de Toledo Piza Jr. and deposited in the collection of the University of Sao Paulo, Escola Superior de Agricultura &quot; Luiz de Queiroz, Brazil. Zootaxa, 2635 (1), 41 - 66. https: // doi. org / 10.11646 / zootaxa. 2635.1.3" type="journal article" year="2010">Chamorro-Rengifo &amp; Braun, 2010</bibRefCitation>
; Chamorro-Rengifo, Braun &amp; Lopes-Andrade, 2014; Fialho 
<emphasis box="[804,855,1152,1176]" italics="true" pageId="2" pageNumber="432">et. al</emphasis>
, 2014; Cigliano 
<emphasis box="[1025,1076,1152,1176]" italics="true" pageId="2" pageNumber="432">et. al</emphasis>
, 2023.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="432" type="diagnosis">
<paragraph blockId="2.[151,1437,1217,2036]" pageId="2" pageNumber="432">
<emphasis bold="true" box="[151,274,1217,1243]" pageId="2" pageNumber="432">Diagnosis.</emphasis>
Pronotal disc posteriorly forming a round projection with big black triangular spot, in dorsal view (
<figureCitation captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Fig. 2C</figureCitation>
). Tegmina narrow with rounded apex (
<figureCitation box="[635,705,1253,1279]" captionStart="FIGURE 3" captionStartId="4.[152,255,755,779]" captionTargetBox="[173,1414,181,731]" captionTargetId="figure-19@4.[173,1414,181,731]" captionTargetPageId="4" captionText="FIGURE 3. Megatympanon speculatum, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7804393" httpUri="https://zenodo.org/record/7804393/files/figure.png" pageId="2" pageNumber="432">Fig. 3</figureCitation>
); total length of tegmina almost two times longer than body, in lateral view (
<figureCitation box="[296,386,1289,1315]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Fig. 2A</figureCitation>
). Vein MA with one branch reaching apex of tegmina (
<figureCitation box="[991,1058,1289,1315]" captionStart="FIGURE 3" captionStartId="4.[152,255,755,779]" captionTargetBox="[173,1414,181,731]" captionTargetId="figure-19@4.[173,1414,181,731]" captionTargetPageId="4" captionText="FIGURE 3. Megatympanon speculatum, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7804393" httpUri="https://zenodo.org/record/7804393/files/figure.png" pageId="2" pageNumber="432">Fig. 3</figureCitation>
). Cercus curved inwards in dorsal view, conical in lateral view, apically with tip small and curved inward (
<figureCitation box="[976,1093,1325,1351]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Figs. 2I–J</figureCitation>
). Subgenital plate trapezoidal, laterally concave and posteriorly projected with medial notch narrow and Y-shaped (
<figureCitation box="[1097,1213,1361,1387]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Figs. 2I–J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="435" pageId="2" pageNumber="432" type="description">
<paragraph blockId="2.[151,1437,1217,2036]" pageId="2" pageNumber="432">
<emphasis bold="true" box="[199,419,1397,1423]" pageId="2" pageNumber="432">Description. Male.</emphasis>
We include here only characteristics that were not used in the original description made by 
<bibRefCitation author="Piza Jr., S. de" box="[151,281,1433,1460]" pageId="2" pageNumber="432" pagination="237 - 241" refId="ref4958" refString="Piza Jr., S. de T. (1958) A primeira especie neotropical da familia &quot; Tympanophoridae &quot; (Orthoptera, Tettigonioidea). Revista Brasileira de Biologia, 18 (2), 237 - 241." type="journal article" year="1958">Piza (1958)</bibRefCitation>
or in the redescription by 
<bibRefCitation author="Fialho, V. S. &amp; Chamorro-Rengifo, J. &amp; Lopes-Andrade, C. &amp; Yotoko, K." box="[577,796,1433,1459]" pageId="2" pageNumber="432" pagination="1 - 42" refId="ref4753" refString="Fialho, V. S., Chamorro-Rengifo, J., Lopes-Andrade, C. &amp; Yotoko, K. (2014) Systematics of spiny predatory katydids (Tettigoniidae: Listroscelidinae) from the Brazilian Atlantic forest based on morphology and molecular data. PLoS ONE, 9 (8), 1 - 42. https: // doi. org / 10.1371 / journal. pone. 0103758" type="journal article" year="2014">
Fialho 
<emphasis box="[655,711,1433,1459]" italics="true" pageId="2" pageNumber="432">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[151,1437,1217,2036]" pageId="2" pageNumber="432">
<emphasis box="[199,280,1469,1495]" italics="true" pageId="2" pageNumber="432">Thorax</emphasis>
. Pronotal disc rectangular, without carina, anteriorly straight, slightly convex laterally and posteriorly, forming a round projection with big black triangular spot, in dorsal view (
<figureCitation box="[1000,1090,1505,1531]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Fig. 2C</figureCitation>
); Dorsal-lateral suture slightly curved (
<figureCitation box="[243,376,1541,1567]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Figs. 2D–E</figureCitation>
). Furcal suture bifurcated, V-shaped and concolorous with pronotal disc (
<figureCitation box="[1196,1284,1541,1567]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="2" pageNumber="432">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,1217,2036]" pageId="2" pageNumber="432">
<emphasis box="[199,267,1578,1603]" italics="true" pageId="2" pageNumber="432">Wings</emphasis>
. Tegmina narrow with rounded apex (
<figureCitation box="[700,769,1577,1603]" captionStart="FIGURE 3" captionStartId="4.[152,255,755,779]" captionTargetBox="[173,1414,181,731]" captionTargetId="figure-19@4.[173,1414,181,731]" captionTargetPageId="4" captionText="FIGURE 3. Megatympanon speculatum, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7804393" httpUri="https://zenodo.org/record/7804393/files/figure.png" pageId="2" pageNumber="432">Fig. 3</figureCitation>
). Vein R slightly sinuous anteriorly and curved posteriorly, with many branches almost straight and parallel reaching anterior margin of tegmina; main branch of Vein R reaching apex of tegmina. Vein MA with one branch, anteriorly straight and posteriorly slightly curved, reaching apex of tegmina; Vein MP with four bifurcations at main branch and seven branches reaching posterior margin of tegmina. Vein CuA narrow, anteriorly curved, surrounding area of second mirror cell, medially straight and posteriorly curved; vein CuA with branches slightly curved, parallel and turned to apex, except by first branch curved to base of tegmina; vein CuP curved, wide, surrounding posterior margin of first mirror cell. Vein AA narrow, surrounding area of first mirror cell. Vein Sc sinuous with several short ramifications (
<figureCitation box="[1117,1185,1829,1855]" captionStart="FIGURE 3" captionStartId="4.[152,255,755,779]" captionTargetBox="[173,1414,181,731]" captionTargetId="figure-19@4.[173,1414,181,731]" captionTargetPageId="4" captionText="FIGURE 3. Megatympanon speculatum, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.7804393" httpUri="https://zenodo.org/record/7804393/files/figure.png" pageId="2" pageNumber="432">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="2.[151,1437,1217,2036]" pageId="2" pageNumber="432">
Left stridulatory file curved with apex straight and narrow; apical teeth short and close to each other; medial and basal teeth long and wide (
<figureCitation box="[453,544,1901,1927]" captionStart="FIGURE 4" captionStartId="4.[152,255,2002,2026]" captionTargetBox="[162,1426,875,1979]" captionTargetId="figure-69@4.[162,1426,875,1979]" captionTargetPageId="4" captionText="FIGURE 4. Megatympanon speculatum, stridulatory file of male. A: left file; B: right file." figureDoi="http://doi.org/10.5281/zenodo.7804395" httpUri="https://zenodo.org/record/7804395/files/figure.png" pageId="2" pageNumber="432">Fig. 4A</figureCitation>
). Total length of left stridulatory file 
<quantity box="[961,1028,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="432" unit="mm" value="4.0">4 mm</quantity>
, greater vein width of 0.5 mm and a total of 68 teeth (
<figureCitation box="[347,438,1937,1963]" captionStart="FIGURE 4" captionStartId="4.[152,255,2002,2026]" captionTargetBox="[162,1426,875,1979]" captionTargetId="figure-69@4.[162,1426,875,1979]" captionTargetPageId="4" captionText="FIGURE 4. Megatympanon speculatum, stridulatory file of male. A: left file; B: right file." figureDoi="http://doi.org/10.5281/zenodo.7804395" httpUri="https://zenodo.org/record/7804395/files/figure.png" pageId="2" pageNumber="432">Fig. 4A</figureCitation>
). Right stridulatory file sinuous with apex straight and narrowing gradually; apical teeth short and close to each other; medial teeth big and sparse; basal teeth similar to medial teeth but shorter and closer to each other (
<figureCitation box="[311,400,2009,2035]" captionStart="FIGURE 4" captionStartId="4.[152,255,2002,2026]" captionTargetBox="[162,1426,875,1979]" captionTargetId="figure-69@4.[162,1426,875,1979]" captionTargetPageId="4" captionText="FIGURE 4. Megatympanon speculatum, stridulatory file of male. A: left file; B: right file." figureDoi="http://doi.org/10.5281/zenodo.7804395" httpUri="https://zenodo.org/record/7804395/files/figure.png" pageId="2" pageNumber="432">Fig. 4B</figureCitation>
). Total length file of 4.3 mm, widest vein of 0.5 mm and total of 74 teeth (
<figureCitation box="[1232,1331,2009,2035]" captionStart="FIGURE 4" captionStartId="4.[152,255,2002,2026]" captionTargetBox="[162,1426,875,1979]" captionTargetId="figure-69@4.[162,1426,875,1979]" captionTargetPageId="4" captionText="FIGURE 4. Megatympanon speculatum, stridulatory file of male. A: left file; B: right file." figureDoi="http://doi.org/10.5281/zenodo.7804395" httpUri="https://zenodo.org/record/7804395/files/figure.png" pageId="2" pageNumber="432">Figs. 4B</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7804389" ID-Zenodo-Dep="7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="3" pageNumber="433" startId="3.[152,255,1896,1920]" targetBox="[166,1420,181,1873]" targetPageId="3">
<paragraph blockId="3.[151,1437,1896,2029]" pageId="3" pageNumber="433">
<emphasis bold="true" box="[152,281,1896,1921]" pageId="3" pageNumber="433">FIGURE 2.</emphasis>
<taxonomicName authorityName="Piza" authorityYear="1958" box="[289,569,1897,1921]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[289,569,1897,1921]" italics="true" pageId="3" pageNumber="433">Megatympanon speculatum</emphasis>
</taxonomicName>
, male. A: 
<emphasis box="[678,753,1897,1921]" italics="true" pageId="3" pageNumber="433">habitus</emphasis>
, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7804393" ID-Zenodo-Dep="7804393" httpUri="https://zenodo.org/record/7804393/files/figure.png" pageId="4" pageNumber="434" startId="4.[152,255,755,779]" targetBox="[173,1414,181,731]" targetPageId="4">
<paragraph blockId="4.[151,1437,755,851]" pageId="4" pageNumber="434">
<emphasis bold="true" box="[152,282,755,779]" pageId="4" pageNumber="434">FIGURE 3.</emphasis>
<taxonomicName authorityName="Piza" authorityYear="1958" box="[291,572,755,779]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[291,572,755,779]" italics="true" pageId="4" pageNumber="434">Megatympanon speculatum</emphasis>
</taxonomicName>
, right tegmina of male in dorsal view. Abbreviations: AP: Posterior anal vein; AA: anterior anal vein; CuA: anterior cubital vein; CuP: posterior cubital vein; MA: anterior median vein; MP: posterior median vein; R: radial vein; Sc: subcostal vein.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7804395" ID-Zenodo-Dep="7804395" box="[152,1074,2002,2027]" httpUri="https://zenodo.org/record/7804395/files/figure.png" pageId="4" pageNumber="434" startId="4.[152,255,2002,2026]" targetBox="[162,1426,875,1979]" targetPageId="4">
<paragraph blockId="4.[152,1074,2002,2027]" box="[152,1074,2002,2027]" pageId="4" pageNumber="434">
<emphasis bold="true" box="[152,280,2002,2027]" pageId="4" pageNumber="434">FIGURE 4.</emphasis>
<taxonomicName authorityName="Piza" authorityYear="1958" box="[287,566,2003,2027]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[287,566,2003,2027]" italics="true" pageId="4" pageNumber="434">Megatympanon speculatum</emphasis>
</taxonomicName>
, stridulatory file of male. A: left file; B: right file.
</paragraph>
</caption>
<paragraph blockId="5.[151,1437,151,645]" pageId="5" pageNumber="435">
<emphasis box="[199,305,151,177]" italics="true" pageId="5" pageNumber="435">Abdomen</emphasis>
. Cercus curved inwards in dorsal view, conical in lateral view, apically with tip small and curved inward; apex of cercus exceeding base of stylus; cercus with several short bristles (
<figureCitation box="[1105,1232,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="5" pageNumber="435">Figs. 2I–K</figureCitation>
). Subgenital plate trapezoidal, laterally concave and posteriorly projected with medial notch narrow and Y-shaped (
<figureCitation box="[1224,1348,223,249]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="5" pageNumber="435">Figs. 2I–K</figureCitation>
). Stylus short and triangular (
<figureCitation box="[386,511,259,285]" captionStart="FIGURE 2" captionStartId="3.[152,255,1896,1920]" captionTargetBox="[166,1420,181,1873]" captionTargetId="figure-31@3.[166,1420,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Megatympanon speculatum, male. A: habitus, dorsal view; B: head, frontal view; C: head and pronotum, dorsal view; D: head and pronotum, lateral view; E: Thoracic sternites, ventral view; F: foreleg, lateral view; G: midleg, lateral view; H: stridulatory area, dorsal view; I–K: Terminalia in ventral, dorsal and lateral view respectively. Abbreviations: Mes: mesobasisternum; Met: metabasisternum; Cer: cerci; Sty: styli; Pl: subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.7804389" httpUri="https://zenodo.org/record/7804389/files/figure.png" pageId="5" pageNumber="435">Figs. 2I–K</figureCitation>
).
</paragraph>
<paragraph blockId="5.[151,1437,151,645]" pageId="5" pageNumber="435">
<emphasis bold="true" box="[199,465,294,321]" pageId="5" pageNumber="435">Geographical records.</emphasis>
<materialsCitation country="Brazil" location="Sao Paulo" pageId="5" pageNumber="435" specimenCount="1" stateProvince="Sao Paulo">
<collectingCountry box="[472,545,295,321]" name="Brazil" pageId="5" pageNumber="435">Brazil</collectingCountry>
: 
<collectingRegion box="[555,671,295,321]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="435">São Paulo</collectingRegion>
, 
<collectingRegion box="[682,832,295,321]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="435">Minas Gerais</collectingRegion>
(new record), 
<collectingRegion box="[997,1159,295,321]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="435">Rio de Janeiro</collectingRegion>
and 
<collectingRegion box="[1214,1373,295,321]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="435">Espírito Santo</collectingRegion>
(new record) (
<figureCitation box="[247,316,331,357]" captionStart="FIGURE 9" captionStartId="9.[152,255,1006,1030]" captionTargetBox="[151,1436,181,982]" captionTargetId="figure-31@9.[151,1436,181,982]" captionTargetPageId="9" captionText="FIGURE 9. Map of Megatympanon species geographical records. Area of the Serra do Mar coastal forests modified from WWF." figureDoi="http://doi.org/10.5281/zenodo.7804408" httpUri="https://zenodo.org/record/7804408/files/figure.png" pageId="5" pageNumber="435">Fig. 9</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="435" type="materials_examined">
<paragraph blockId="5.[151,1437,151,645]" pageId="5" pageNumber="435">
<emphasis bold="true" box="[199,438,366,392]" pageId="5" pageNumber="435">Material examined.</emphasis>
<materialsCitation collectingDate="1963-02-25" collectionCode="MZSP" collectorName="M. Werner &amp; H. Reichardt" country="Brazil" location="Est. Biol. Boraceria" municipality="Salesopolis" pageId="5" pageNumber="435" specimenCode="MZSP 5356" specimenCount="1" stateProvince="Sao Paulo">
<collectingCountry box="[450,552,367,393]" name="Brazil" pageId="5" pageNumber="435">BRASIL</collectingCountry>
, 
<collectingRegion box="[569,690,367,393]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="435">São Paulo</collectingRegion>
, 
<collectingMunicipality box="[706,836,367,393]" pageId="5" pageNumber="435">Salesópolis</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03A287C10D21FFC8FF1AAD83FE16FD2B:8ED4600C0D26FFC8FCD8AF13FBDEFE27" box="[853,1088,367,393]" country="Brazil" municipality="Salesopolis" name="Est. Biol. Boraceria" pageId="5" pageNumber="435" stateProvince="Sao Paulo">Est. Biol. Boracéria</location>
, 
<date box="[1104,1220,366,393]" pageId="5" pageNumber="435" value="1963-02-25">
<collectingDate box="[1104,1220,366,393]" pageId="5" pageNumber="435" value="1963-02-25">25.ii.1963</collectingDate>
</date>
, 
<collectorName box="[1237,1363,367,393]" pageId="5" pageNumber="435">M. Werner</collectorName>
&amp; 
<collectorName pageId="5" pageNumber="435">H. Reichardt</collectorName>
<emphasis box="[271,305,403,429]" italics="true" pageId="5" pageNumber="435">leg</emphasis>
. / 
<taxonomicName authority="Piza" authorityName="Piza" authorityYear="1958" box="[341,712,403,429]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[341,654,403,429]" italics="true" pageId="5" pageNumber="435">Megatympanon speculatum</emphasis>
Piza
</taxonomicName>
<emphasis box="[723,764,403,429]" italics="true" pageId="5" pageNumber="435">det.</emphasis>
/ 
<specimenCode box="[793,932,402,429]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP 5356</specimenCode>
(1J 
<collectionCode box="[997,1075,403,429]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1961-02-24" collectionCode="MZSP" collectorName="Rabello" country="Brazil" elevation="850" location="Idem" pageId="5" pageNumber="435" specimenCode="MZSP 5354" specimenCount="1" stateProvince="Sao Paulo">
<emphasis box="[1098,1154,403,429]" italics="true" pageId="5" pageNumber="435">
<location LSID="urn:lsid:plazi:treatment:03A287C10D21FFC8FF1AAD83FE16FD2B:8ED4600C0D26FFC8FBC7AFEFFB1CFE03" box="[1098,1154,403,429]" country="Brazil" name="Idem" pageId="5" pageNumber="435" stateProvince="Sao Paulo">Idem</location>
</emphasis>
, 
<quantity box="[1172,1238,402,429]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="435" unit="m" value="850.0">
<elevation box="[1172,1238,402,429]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="435" unit="m" value="850.0">850m</elevation>
</quantity>
, 
<date box="[1253,1369,403,429]" pageId="5" pageNumber="435" value="1961-02-24">
<collectingDate box="[1253,1369,403,429]" pageId="5" pageNumber="435" value="1961-02-24">24.ii.1961</collectingDate>
</date>
, luz, 
<collectorName box="[151,238,439,465]" pageId="5" pageNumber="435">Rabello</collectorName>
leg. / 
<taxonomicName authority="Piza" authorityName="Piza" authorityYear="1958" box="[305,666,439,465]" class="Insecta" family="Tettigoniidae" genus="Megatympanon" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="speculatum">
<emphasis box="[305,613,439,465]" italics="true" pageId="5" pageNumber="435">Megatympanon speculatum</emphasis>
Piza
</taxonomicName>
<emphasis box="[673,714,439,465]" italics="true" pageId="5" pageNumber="435">det.</emphasis>
/ 
<specimenCode box="[733,867,438,465]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP 5354</specimenCode>
(1J 
<collectionCode box="[923,1001,439,465]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1989-03-18" collectingDateMax="1989-03-24" collectingDateMin="1989-03-18" collectionCode="MZSP" collectorName="A. Mesa &amp; A. Van der Laan &amp; H. Costa &amp; L. Nadai &amp; E. Zeffa &amp; M. C. Bianchi &amp; Colecao Alejo Mesa" country="Brazil" location="Boraceia" municipality="Idem" pageId="5" pageNumber="435" specimenCode="MZSP 6063" specimenCount="1" stateProvince="Sao Paulo">
<emphasis box="[1020,1076,439,465]" italics="true" pageId="5" pageNumber="435">
<collectingMunicipality box="[1020,1076,439,465]" pageId="5" pageNumber="435">Idem</collectingMunicipality>
</emphasis>
, 
<location LSID="urn:lsid:plazi:treatment:03A287C10D21FFC8FF1AAD83FE16FD2B:8ED4600C0D26FFC8FBCCAFCBFB39FE7F" box="[1089,1191,439,465]" country="Brazil" municipality="Idem" name="Boraceia" pageId="5" pageNumber="435" stateProvince="Sao Paulo">Boracéia</location>
, 
<date box="[1201,1362,439,465]" pageId="5" pageNumber="435" value="1989-03-18" valueMax="1989-03-24" valueMin="1989-03-18">
<collectingDate box="[1201,1362,439,465]" pageId="5" pageNumber="435" value="1989-03-18" valueMax="1989-03-24" valueMin="1989-03-18">
18-24.iii. 
<quantity box="[1301,1362,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.98985" pageId="5" pageNumber="435" unit="m" value="1989.85">1989</quantity>
</collectingDate>
</date>
, 850m alt., 
<collectorName box="[198,294,475,501]" pageId="5" pageNumber="435">A. Mesa</collectorName>
, 
<collectorName box="[303,486,475,501]" pageId="5" pageNumber="435">A. Van der Laan</collectorName>
, 
<collectorName box="[496,596,475,501]" pageId="5" pageNumber="435">H. Costa</collectorName>
, 
<collectorName box="[606,706,475,501]" pageId="5" pageNumber="435">L. Nadai</collectorName>
, 
<collectorName box="[716,806,475,501]" pageId="5" pageNumber="435">E. Zeffa</collectorName>
&amp; 
<collectorName box="[840,990,475,501]" pageId="5" pageNumber="435">M.C. Bianchi</collectorName>
<emphasis box="[996,1038,475,501]" italics="true" pageId="5" pageNumber="435">leg.</emphasis>
/ 
<collectorName box="[1057,1283,475,501]" pageId="5" pageNumber="435">Coleção Alejo Mesa</collectorName>
/ 
<specimenCode box="[1302,1436,475,501]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP 6063</specimenCode>
</materialsCitation>
(1J 
<collectionCode box="[203,281,511,537]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="435" type="Museum">MZSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="435" type="description">
<paragraph blockId="5.[151,1437,151,645]" pageId="5" pageNumber="435">
<emphasis bold="true" box="[199,456,546,573]" pageId="5" pageNumber="435">Measurements (mm).</emphasis>
J: TL: 36,1–34,5; TegL: 67,4 –66,7; TegH: 14,1–13,4; WF: 6,8–5,7; PL: 12,0–11,4; 
<collectingCountry box="[151,191,583,609]" name="Philippines" pageId="5" pageNumber="435">PH</collectingCountry>
: 6,4–6,3; FF: 12,9–12,2; FT: 12,6–12,0; MF: 12,4–12,0; MT: 13,4–12,9; HF: 33,6–33,3; HT: 35,6–33,5; Lplac: 5,7–4,8; LC: 3,8–3,5.
</paragraph>
</subSubSection>
</treatment>
</document>