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<document id="0B26C1DFE06DD99D96404201D749D9AB" ID-DOI="10.5252/geodiversitas2021v43a22" ID-ISSN="1638-9395" ID-Zenodo-Dep="5764379" ID-ZooBank="urn:lsid:zoobank.org:pub:6D1643AD-66A5-4678-9397-CE08E610D641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1638890191441" checkinUser="felipe" docAuthor="Vasseur, Raphaël &amp; Lathuilière, Bernard" docDate="2021" docId="03A287A2485131209AAD630FFE70F83C" docLanguage="en" docName="Geodiversitas.43.22.1187-1291.pdf" docOrigin="Geodiversitas 43 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Araiophyllum liasicum Beauvais 1986" docType="treatment" docVersion="6" lastPageNumber="1203" masterDocId="FF9BFFDA484331329A586119FFC1FFD0" masterDocTitle="Pliensbachian corals from the Western Tethys" masterLastPageNumber="1291" masterPageNumber="1187" pageNumber="1203" updateTime="1699441156090" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="72BA1047BB58224BEDF2E2B7A9983E25">Pliensbachian corals from the Western Tethys</mods:title>
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<mods:namePart id="01AE441A12C34F54DA96B2ED6CCFB2DC">Vasseur, Raphaël</mods:namePart>
<mods:affiliation id="F9D964B7658071C39A35E882CE117C33">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<mods:namePart id="94152D18A8CADF9DCF540A817A23D40E">Lathuilière, Bernard</mods:namePart>
<mods:affiliation id="3AAE98D169513273D17EB23BEDEEC68E">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<mods:title id="ACFCB122CD369751B1A978CA152F44DD">Geodiversitas</mods:title>
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<mods:date id="72E3CE1072F819B4EBC00F63D4C8E894">2021</mods:date>
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<taxonomicName id="4C0B4D37485131209AAD630FFD57FDE0" authority="Beauvais, 1986" authorityName="Beauvais" authorityYear="1986" box="[245,662,534,561]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="species" species="liasicum">
<emphasis id="B97FEAA6485131209AAD630FFE2DFDE1" bold="true" box="[245,492,534,561]" italics="true" pageId="18" pageNumber="1203">Araiophyllum liasicum</emphasis>
<bibRefCitation id="EF9A4B45485131209BAB630FFD57FDE0" author="BEAUVAIS L." box="[499,662,534,560]" pageId="18" pageNumber="1203" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986</bibRefCitation>
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<paragraph id="8BB436B4485131209BC6632FFE2CFD81" blockId="18.[245,662,534,593]" box="[414,493,566,593]" pageId="18" pageNumber="1203">
(
<figureCitation id="13302A31485131209BFF632FFE25FD80" box="[423,484,566,592]" captionStart="FIG" captionStartId="19.[132,143,1940,1957]" captionTargetBox="[185,1397,215,1875]" captionTargetId="figure-16@19.[185,1398,215,1875]" captionTargetPageId="19" captionText="FIG. 8. — Araiophyllum liasicum Beauvais,1986: A, transverse section in the holotype MNHN.F.A32023; B, lateral view of a corallite of the holotype MNHN.F.A32023; C, transverse section of a corallite of the holotype MNHN.F.A32023 (skeleton in dark);D, longitudinal section of a corallite of the holotype MNHN.F.A32023; E, detail in transverse section of the axial part of one corallite of the holotype MNHN.F.A32023; F, transverse section of one corallite of the holotype MNHN.F.A32023 (skeleton in dark). Scale bars: A, 4 cm; B, 1 cm; C, 2 mm; D, E, 1 mm; F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5764399" httpUri="https://zenodo.org/record/5764399/files/figure.png" pageId="18" pageNumber="1203">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311653F485131209ADC6376FEB1FD72" pageId="18" pageNumber="1203" type="reference_group">
<paragraph id="8BB436B4485131209ADC6376FEB1FD72" blockId="18.[132,774,623,674]" pageId="18" pageNumber="1203">
<treatmentCitationGroup id="AB1B119A485131209ADC6376FEB1FD72" pageId="18" pageNumber="1203">
<treatmentCitation id="0AAA10A5485131209ADC6376FDD4FD57" author="BEAUVAIS L." box="[132,533,623,647]" page="55" pageId="18" pageNumber="1203" year="1986">
<taxonomicName id="4C0B4D37485131209ADC6376FDD4FD57" authority="Beauvais, 1986: 55" authorityName="Beauvais" authorityPageNumber="55" authorityYear="1986" box="[132,533,623,647]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="species" species="liasicum">
<emphasis id="B97FEAA6485131209ADC6376FE96FD57" box="[132,343,623,647]" italics="true" pageId="18" pageNumber="1203">Araiophyllum liasicum</emphasis>
<bibRefCitation id="EF9A4B45485131209B046376FDD4FD57" author="BEAUVAIS L." box="[348,533,623,647]" pageId="18" pageNumber="1203" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986: 55</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 12, fig. 3. —
<treatmentCitation id="0AAA10A54851312098E56376FEDBFD71" author="VASSEUR R." page="124" pageId="18" pageNumber="1203" year="2018">
<bibRefCitation id="EF9A4B454851312098E56376FEDBFD71" author="VASSEUR R." pageId="18" pageNumber="1203" refId="ref55769" refString="VASSEUR R. 2018. - Extinctions et recouvrements de coraux au cours de la crise Pliensbachien-Toarcien. These de doctorat, Universite de Lorraine, France, 542 p." type="book" year="2018">Vasseur 2018: 124-125</bibRefCitation>
</treatmentCitation>
, fig. 3.8.
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</paragraph>
</subSubSection>
<subSubSection id="C311653F485131209ADC63A3FE5FFC15" pageId="18" pageNumber="1203" type="materials_examined">
<paragraph id="8BB436B4485131209ADC63A3FE5FFC15" blockId="18.[132,775,697,965]" pageId="18" pageNumber="1203">
<materialsCitation id="3B633CE9485131209ADC63A3FE5FFC15" accessionNumber="A32023" collectionCode="TYPE, MNHN" country="Morocco" location="Pliensbachian" municipality="High Atlas" pageId="18" pageNumber="1203" specimenCount="4" typeStatus="holotype">
<collectionCode id="ED1AAE71485131209ADC63A3FF74FD1F" box="[132,181,698,721]" pageId="18" pageNumber="1203">TYPE</collectionCode>
MATERIAL. —
<typeStatus id="54B08816485131209B1F63A0FE65FD01" box="[327,420,697,721]" pageId="18" pageNumber="1203" type="holotype">Holotype</typeStatus>
<collectionCode id="ED1AAE71485131209BF363A3FE3EFD01" box="[427,511,698,721]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="18" pageNumber="1203">MNHN</collectionCode>
.F.
<accessionNumber id="9458AB5748513120984D63A0FDA5FD01" box="[533,612,697,721]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A32023" pageId="18" pageNumber="1203" type="EnaNcbi">A32023</accessionNumber>
(
<bibRefCitation id="EF9A4B4548513120982B63A0FF5EFD3B" author="BEAUVAIS L." pageId="18" pageNumber="1203" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais 1986: 55</bibRefCitation>
, pl. 12, fig. 3).
<typeStatus id="54B08816485131209ADC621DFF74FCC9" box="[132,181,772,795]" pageId="18" pageNumber="1203">TYPE</typeStatus>
HORIZON. —
<location id="8ED4606F485131209B1B621AFE0CFCCB" LSID="urn:lsid:plazi:treatment:03A287A2485131209AAD630FFE70F83C:8ED4606F485131209B1B621AFE0CFCCB" box="[323,461,771,795]" country="Morocco" municipality="High Atlas" name="Pliensbachian" pageId="18" pageNumber="1203">Pliensbachian</location>
.
<typeStatus id="54B08816485131209ADC622AFF74FC99" box="[132,181,819,842]" pageId="18" pageNumber="1203">TYPE</typeStatus>
LOCALITY. — Jebel Bou Dahar,
<collectingMunicipality id="6BD0ACCE485131209BA8622AFD98FC9B" box="[496,601,819,843]" pageId="18" pageNumber="1203">High Atlas</collectingMunicipality>
of
<collectingCountry id="F31C7624485131209822622AFD14FC9A" box="[634,725,819,842]" name="Morocco" pageId="18" pageNumber="1203">Morocco</collectingCountry>
. STUDIED SAMPLE. —
<typeStatus id="54B08816485131209B0C627BFE75FCAA" box="[340,436,866,890]" pageId="18" pageNumber="1203" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED1AAE71485131209BE5627AFDD3FCAA" box="[445,530,867,890]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="18" pageNumber="1203">MNHN</collectionCode>
.F.
<accessionNumber id="9458AB5748513120987F627BFDB7FCAA" box="[551,630,866,890]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A32023" pageId="18" pageNumber="1203" type="EnaNcbi">A32023</accessionNumber>
. GEOGRAPHIC AND STRATIGRAPHIC RANGES. —
<location id="8ED4606F485131209838628BFD2BFC7A" LSID="urn:lsid:plazi:treatment:03A287A2485131209AAD630FFE70F83C:8ED4606F485131209838628BFD2BFC7A" box="[608,746,914,938]" country="Morocco" municipality="High Atlas" name="Pliensbachian" pageId="18" pageNumber="1203">Pliensbachian</location>
of
<location id="8ED4606F485131209ADE62B5FEE9FC14" LSID="urn:lsid:plazi:treatment:03A287A2485131209AAD630FFE70F83C:8ED4606F485131209ADE62B5FEE9FC14" box="[134,296,940,964]" country="Morocco" municipality="High Atlas" name="Jebel Bou Dahar" pageId="18" pageNumber="1203">Jebel Bou Dahar</location>
(
<collectingCountry id="F31C7624485131209B6D62B4FE52FC15" box="[309,403,941,965]" name="Morocco" pageId="18" pageNumber="1203">Morocco</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311653F485131209ADC62ECFE70F83C" pageId="18" pageNumber="1203" type="description">
<paragraph id="8BB436B4485131209ADC62ECFE9FFA3D" blockId="18.[131,777,1012,1613]" pageId="18" pageNumber="1203">
DESCRIPTION (modified after
<bibRefCitation id="EF9A4B45485131209B9E62EDFDA3FBDE" author="BEAUVAIS L." box="[454,610,1012,1038]" pageId="18" pageNumber="1203" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais 1986</bibRefCitation>
) Phaceloid colony with subcylindrical corallites multiplying by fissiparous intracalicular budding. The external part of the corallites is covered by an epitheca
<emphasis id="B97FEAA6485131209871654DFD87FBBE" box="[553,582,1108,1134]" italics="true" pageId="18" pageNumber="1203">s.l.</emphasis>
that reveals some regularly granulated costae. The calicular plateau has been observed on a very few corallites because of a very poor preservation but it shows the characteristic aspect described by
<bibRefCitation id="EF9A4B45485131209AF065CDFEEEFB3E" author="CUIF J. P." box="[168,303,1235,1262]" pageId="18" pageNumber="1203" pagination="157 - 180" refId="ref50032" refString="CUIF J. P. 1975. - Caracteres morphologiques, microstructuraux et systematiques des Pachythecalidae, nouvelle famille de Madreporaires triasiques. Geobios 8: 157 - 180. https: // doi. org / 10.1016 / S 0016 - 6995 (75) 80035 - 0" type="journal article" year="1975">Cuif (1975)</bibRefCitation>
for the genus: a hemispherical shape corresponding to a particular orientation of trabecular axes. Radial elements are totally porous, maybe a few less than in Upper Jurassic microsolenids. However, pores appear more abundant in the axial zone than in the periphery in transverse section because of the orientation of trabecular axes. Moniliform distal edge. Pennulae are numerous. Synapticulae abundant. Endotheca not observed. Weak, parietal papillose columella.
</paragraph>
<paragraph id="8BB436B4485131209AC364EAFEEBF99D" blockId="18.[131,777,1012,1613]" pageId="18" pageNumber="1203">
Calicular diameter:
<quantity id="4CF39B51485131209B2864EAFE31F9DD" box="[368,496,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="18" pageNumber="1203" unit="mm" value="8.0" valueMax="12.0" valueMin="4.0">4 to 12 mm</quantity>
Number of septa: 92 to 120 Septal density: 12 per
<quantity id="4CF39B51485131209B9A670AFDC5F9FD" box="[450,516,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="1203" unit="mm" value="2.0">2 mm</quantity>
Trabecular density: 4 to 5 per
<quantity id="4CF39B51485131209AB9672AFEE7F99D" box="[225,294,1587,1613]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="1203" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BB436B4485131209ADC676BFE13F95B" blockId="18.[130,777,1650,2028]" box="[132,466,1650,1676]" pageId="18" pageNumber="1203">SIMILARITIES AND DIFFERENCES</paragraph>
<paragraph id="8BB436B4485131209ADC678BFE59F8FC" blockId="18.[130,777,1650,2028]" pageId="18" pageNumber="1203">
This species appears very close to the genus
<taxonomicName id="4C0B4D37485131209808678AFF7DF91C" authority="Koby, 1887" authorityName="Koby" authorityYear="1887" class="Anthozoa" family="Comoseriidae" genus="Dermoseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6485131209808678AFD00F97C" box="[592,705,1683,1708]" italics="true" pageId="18" pageNumber="1203">Dermoseris</emphasis>
Koby, 1887
</taxonomicName>
that shows as
<taxonomicName id="4C0B4D37485131209B3167ABFE3BF91C" authorityName="Cuif" authorityYear="1975" box="[361,506,1714,1740]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6485131209B3167ABFE3BF91C" box="[361,506,1714,1740]" italics="true" pageId="18" pageNumber="1203">Araiophyllum</emphasis>
</taxonomicName>
subhemispheric calices.
<taxonomicName id="4C0B4D37485131209ADE67CBFEAEF93C" authorityName="Beauvais" authorityYear="1986" box="[134,367,1746,1772]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="species" species="liasicum">
<emphasis id="B97FEAA6485131209ADE67CBFEAEF93C" box="[134,367,1746,1772]" italics="true" pageId="18" pageNumber="1203">Araiophyllum liasicum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0B4D37485131209BAE67CBFDB4F93C" authorityName="Cuif" authorityYear="1975" box="[502,629,1746,1772]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="species" species="triasicum">
<emphasis id="B97FEAA6485131209BAE67CBFDB4F93C" box="[502,629,1746,1772]" italics="true" pageId="18" pageNumber="1203">A. triasicum</emphasis>
</taxonomicName>
by the bigger size of the corallites, closer to the dimensions of the Upper Jurassic genus
<taxonomicName id="4C0B4D37485131209B78660BFE50F8FB" authorityName="Koby" authorityYear="1887" box="[288,401,1810,1835]" class="Anthozoa" family="Comoseriidae" genus="Dermoseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6485131209B78660BFE50F8FB" box="[288,401,1810,1835]" italics="true" pageId="18" pageNumber="1203">Dermoseris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB436B4485131209AC36628FE70F83C" blockId="18.[130,777,1650,2028]" pageId="18" pageNumber="1203">
The more significant difference between the two genera would be the different microstructure of pennulae between
<taxonomicName id="4C0B4D37485131209ADD6668FED6F85B" authorityName="Cuif" authorityYear="1975" box="[133,279,1905,1931]" class="Anthozoa" family="Comoseriidae" genus="Araiophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6485131209ADD6668FED6F85B" box="[133,279,1905,1931]" italics="true" pageId="18" pageNumber="1203">Araiophyllum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B4D37485131209B0D666BFE08F85B" authorityName="Koby" authorityYear="1887" box="[341,457,1906,1931]" class="Anthozoa" family="Comoseriidae" genus="Dermoseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="1203" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6485131209B0D666BFE08F85B" box="[341,457,1906,1931]" italics="true" pageId="18" pageNumber="1203">Dermoseris</emphasis>
</taxonomicName>
: despite the low number of observations, it seems the latter shows regular granulations on the curved edges of pennulae and a more regular alternating disposition (see
<bibRefCitation id="EF9A4B45485131209B6966C8FE60F83C" author="CUIF J. P." box="[305,417,2001,2028]" pageId="18" pageNumber="1203" pagination="157 - 180" refId="ref50032" refString="CUIF J. P. 1975. - Caracteres morphologiques, microstructuraux et systematiques des Pachythecalidae, nouvelle famille de Madreporaires triasiques. Geobios 8: 157 - 180. https: // doi. org / 10.1016 / S 0016 - 6995 (75) 80035 - 0" type="journal article" year="1975">Cuif 1975</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>