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<document id="8CE95C49F96337F78DC20756E5CC0E79" ID-DOI="10.11646/zootaxa.5353.5.6" ID-GBIF-Dataset="95bf5e71-404a-4757-9c89-50e4ffc91c95" ID-ISSN="1175-5326" ID-Zenodo-Dep="10010328" ID-ZooBank="80C94B73-A72D-4C9A-BC36-AF95AF341E81" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1697229704863" checkinUser="plazi" docAuthor="Choi, Jinyeong &amp; Lee, Seunghwan" docDate="2023" docId="03A28797FF82FFF3FF4AFB39FC71FBF2" docLanguage="en" docName="zootaxa.5353.5.6.pdf" docOrigin="Zootaxa 5353 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.5.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discochiton bambusae Choi &amp; Lee 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="484" masterDocId="FF9BFFEFFF83FFF6FFDDFFF0FFB5FFC7" masterDocTitle="A new species of Discochiton Hodgson &amp; Williams (Hemiptera: Coccomorpha: Coccidae) on bamboo from Myanmar" masterLastPageNumber="487" masterPageNumber="479" pageNumber="480" updateTime="1697479236921" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7200EB5C4B11E63B7872DD32E0D72818">A new species of Discochiton Hodgson &amp; Williams (Hemiptera: Coccomorpha: Coccidae) on bamboo from Myanmar</mods:title>
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<mods:namePart id="099798E2C559F63923B6F0192CC4063B">Choi, Jinyeong</mods:namePart>
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<mods:affiliation id="E9126C14461897AAB7C8BB44038A92F1">Evolution, Cell Biology, and Symbiosis Unit, Okinawa Institute of Science and Technology Graduate University, Okinawa 904 - 0495, Japan</mods:affiliation>
<mods:nameIdentifier id="4D0017989358CBDC6BB5BBA99B3B6A7E" type="email">jinyeong.choi@oist.jp</mods:nameIdentifier>
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<mods:namePart id="92D0610CD837B1D5258E83F3B7E857C1">Lee, Seunghwan</mods:namePart>
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<mods:affiliation id="DEA8B94077D55BE69637FDB9EB889A2F">Department of Agricultural Biotechnology, Seoul National University, Seoul 151 - 921, Republic of Korea &amp; Research Institute of Agriculture and Life Sciences, Seoul National University, Seoul 151 - 921, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="36FECD205C258B6F82B68233C2AC6695" type="email">seung@snu.ac.kr</mods:nameIdentifier>
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<mods:title id="08A436213410F1A13A5B4A5816AD7D7E">Zootaxa</mods:title>
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<treatment id="03A28797FF82FFF3FF4AFB39FC71FBF2" ID-DOI="http://doi.org/10.5281/zenodo.10010325" ID-GBIF-Taxon="213996125" ID-Zenodo-Dep="10010325" LSID="urn:lsid:plazi:treatment:03A28797FF82FFF3FF4AFB39FC71FBF2" httpUri="http://treatment.plazi.org/id/03A28797FF82FFF3FF4AFB39FC71FBF2" lastPageId="5" lastPageNumber="484" pageId="1" pageNumber="480">
<subSubSection id="C311650AFF82FFF7FF4AFB39FD39FB24" box="[151,652,1225,1252]" pageId="1" pageNumber="480" type="nomenclature">
<paragraph id="8BB43681FF82FFF7FF4AFB39FD39FB24" blockId="1.[151,652,1225,1287]" box="[151,652,1225,1252]" pageId="1" pageNumber="480">
<heading id="D0FC81EDFF82FFF7FF4AFB39FD39FB24" bold="true" box="[151,652,1225,1252]" fontSize="11" level="1" pageId="1" pageNumber="480" reason="1">
<emphasis id="B97FEA93FF82FFF7FF4AFB39FD39FB24" bold="true" box="[151,652,1225,1252]" pageId="1" pageNumber="480">
<taxonomicName id="4C0B4D02FF82FFF7FF4AFB39FDF6FB24" authority="Choi &amp; Lee" authorityName="Choi &amp; Lee" authorityYear="2023" box="[151,579,1225,1252]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="bambusae" status="sp. n.">
<emphasis id="B97FEA93FF82FFF7FF4AFB39FE1FFB23" bold="true" box="[151,426,1225,1252]" italics="true" pageId="1" pageNumber="480">Discochiton bambusae</emphasis>
Choi &amp; Lee
</taxonomicName>
<taxonomicNameLabel id="A24C57E8FF82FFF7FD94FB39FD39FB24" box="[585,652,1225,1251]" pageId="1" pageNumber="480" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C311650AFF82FFF7FF4AFB1DFEBEFAC0" box="[151,267,1261,1287]" pageId="1" pageNumber="480" type="description">
<paragraph id="8BB43681FF82FFF7FF4AFB1DFEBEFAC0" blockId="1.[151,652,1225,1287]" box="[151,267,1261,1287]" pageId="1" pageNumber="480">
(
<figureCitation id="13302A04FF82FFF7FF42FB1DFEB6FAC0" box="[159,259,1261,1287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[152,255,1512,1536]" captionStartId-1="3.[152,255,1513,1537]" captionStartId-2="4.[152,255,1856,1880]" captionStartId-3="6.[152,255,1943,1967]" captionTargetBox-0="[164,1423,181,1488]" captionTargetBox-1="[164,1423,310,1489]" captionTargetBox-2="[210,1378,181,1832]" captionTargetBox-3="[207,1379,181,1919]" captionTargetId-0="figure-21@2.[164,1423,181,1488]" captionTargetId-1="figure-79@3.[164,1423,309,1490]" captionTargetId-2="figure-21@4.[210,1378,181,1832]" captionTargetId-3="figure-21@6.[207,1379,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionText-0="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." captionText-1="FIGURE 2. Discochiton bambusae Choi &amp; Lee sp. n. A, slide-mounted adult female; B, stigmatic cleft and spines; C, anal plates; D, marginal setae; E, antenna; F, prothoracic leg; G, mesothoracic leg; H, metathoracic leg; I, long dorsal seta; J, short dorsal seta; K, Clear areas (CA14); L, clear areas and associated preopercular pores; M, sclerotized plates forming ventral band; N, submarginal sclerotized ridges. Scale lines for A = 5 mm; K= 1 mm; M = 0.5 mm; B, C, E, F, G, H, L = 100 μm; D = 50 μm; I, J, N = 10 μm." captionText-2="FIGURE 3. Discochiton bambusae Choi &amp; Lee sp. n., adult female.A, reticulations; B, marginal radial lines indicated by pores and setae; C, marginal seta; D, stigmatic cleft and spines, and protrusions on outer margins of stigmatic cleft; E, submarginal sclerotized ridges; F, dorsal setae; G, preopercular pore; H, clear areas surrounded by preopercular pores; I, dorsal pores with a ductule and an unsclerotized pale area; J, anal plates; K, multilocular disc-pore; L, sclerotized plates forming a band; M, metathoracic leg; N, mesothoracic leg; O, prothoracic leg; P, spiracular disc-pore; Q, antenna." captionText-3="FIGURE 4. Discochiton bambusae Choi &amp; Lee sp. n., first-instar nymph.A, stigmatic spines; B, anal plate (left), and anal ring and anal fold (right); C, marginal seta; D, dorsal microduct; E, tarsus and claw with digitules and setae; F, submarginal seta; G, ventral microduct; H, spiracular disc-pore." figureDoi-0="http://doi.org/10.5281/zenodo.10010330" figureDoi-1="http://doi.org/10.5281/zenodo.10010333" figureDoi-2="http://doi.org/10.5281/zenodo.10010335" figureDoi-3="http://doi.org/10.5281/zenodo.10010339" httpUri-0="https://zenodo.org/record/10010330/files/figure.png" httpUri-1="https://zenodo.org/record/10010333/files/figure.png" httpUri-2="https://zenodo.org/record/10010335/files/figure.png" httpUri-3="https://zenodo.org/record/10010339/files/figure.png" pageId="1" pageNumber="480">Figs 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311650AFF82FFF7FF4AFAC5FE13FA7C" pageId="1" pageNumber="480" type="materials_examined">
<paragraph id="8BB43681FF82FFF7FF4AFAC5FE13FA7C" blockId="1.[151,1437,1333,1612]" pageId="1" pageNumber="480">
<emphasis id="B97FEA93FF82FFF7FF4AFAC5FE4BFA88" bold="true" box="[151,510,1333,1359]" pageId="1" pageNumber="480">
Material examined.
<materialsCitation id="3B633CDCFF82FFF7FE55FAC5FBFAFAB4" collectingDate="2018-05-25" collectionCode="SNU" collectorName="Jinyeong Choi" country="Myanmar" latitude="21.380001" location="Ywangan" longitude="96.47" municipality="Ywangan" pageId="1" pageNumber="480" specimenCount="1" stateProvince="Shan State" typeStatus="holotype">
<typeStatus id="54B08823FF82FFF7FE55FAC5FE4CFA88" box="[392,505,1333,1359]" pageId="1" pageNumber="480">Holotype</typeStatus>
:
<specimenCount id="9D0DFD08FF82FFF7FDD8FAC5FD10FA88" box="[517,677,1333,1359]" pageId="1" pageNumber="480">1 adult female</specimenCount>
mounted on a slide,
<collectingCountry id="F31C7611FF82FFF7FC4DFAC6FC4AFA97" box="[912,1023,1334,1360]" name="Myanmar" pageId="1" pageNumber="480">Myanmar</collectingCountry>
,
<collectingRegion id="49CFF863FF82FFF7FBD4FAC5FB34FA88" box="[1033,1153,1333,1359]" country="Myanmar" name="Shan" pageId="1" pageNumber="480">Shan State</collectingRegion>
/
<collectingMunicipality id="6BD0ACFBFF82FFF7FB4BFAC6FAB4FA97" box="[1174,1281,1334,1360]" pageId="1" pageNumber="480">Ywangan</collectingMunicipality>
/
<geoCoordinate id="EE3F5046FF82FFF7FACAFAC7FA29FA95" box="[1303,1436,1335,1362]" degrees="21" direction="north" minutes="22" orientation="latitude" pageId="1" pageNumber="480" precision="15" seconds="48" value="21.380001">21°2248″N</geoCoordinate>
<geoCoordinate id="EE3F5046FF82FFF7FF4AFAABFEACFAB1" box="[151,281,1371,1398]" degrees="96" direction="east" minutes="47" orientation="longitude" pageId="1" pageNumber="480" seconds="87" value="96.47">96°4787″E</geoCoordinate>
/ coll.
<collectorName id="26FE5357FF82FFF7FEA8FAA9FDA9FAB4" box="[373,540,1369,1395]" pageId="1" pageNumber="480">Jinyeong Choi</collectorName>
/
<date id="FFB51041FF82FFF7FDE1FAA9FD1FFAB3" box="[572,682,1369,1396]" pageId="1" pageNumber="480" value="2018-05-25">
<collectingDate id="EFF1E9A9FF82FFF7FDE1FAA9FD1FFAB3" box="[572,682,1369,1396]" pageId="1" pageNumber="480" value="2018-05-25">25.v.2018</collectingDate>
</date>
/
<taxonomicName id="4C0B4D02FF82FFF7FD17FAA9FCCDFAB4" box="[714,888,1369,1395]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="1" pageNumber="480" phylum="Tracheophyta" rank="species" species="tulda">
<emphasis id="B97FEA93FF82FFF7FD17FAA9FCCDFAB4" box="[714,888,1369,1395]" italics="true" pageId="1" pageNumber="480">Bambusa tulda</emphasis>
</taxonomicName>
(
<taxonomicName id="4C0B4D02FF82FFF7FC56FAA9FC5FFAB4" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[907,1002,1369,1395]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="1" pageNumber="480" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) (
<collectionCode id="ED1AAE44FF82FFF7FBD8FAA9FBF6FAB4" box="[1029,1091,1369,1395]" pageId="1" pageNumber="480">SNU</collectionCode>
).
</materialsCitation>
</emphasis>
<materialsCitation id="3B633CDCFF82FFF7FB86FAA9FCA2FA50" collectingDate="2018-05-25" collectionCode="SNU" collectorName="Jinyeong Choi" country="Myanmar" latitude="21.380001" location="Ywangan" municipality="Ywangan" pageId="1" pageNumber="480" specimenCount="9" stateProvince="Shan State" typeStatus="paratype">
<emphasis id="B97FEA93FF82FFF7FB86FAA9FB69FAB4" bold="true" box="[1115,1244,1369,1395]" pageId="1" pageNumber="480">
<typeStatus id="54B08823FF82FFF7FB86FAA9FB62FAB4" box="[1115,1239,1369,1395]" pageId="1" pageNumber="480">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D0DFD08FF82FFF7FB35FAA9FA29FAB4" box="[1256,1436,1369,1395]" pageId="1" pageNumber="480">9 adult females</specimenCount>
mounted singly on slides, same data as holotype (
<collectionCode id="ED1AAE44FF82FFF7FD13FA8DFCB9FA50" box="[718,780,1405,1431]" pageId="1" pageNumber="480">SNU</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B633CDCFF82FFF7FCFDFA8DFE13FA7C" collectingDate="2018-05-25" collectionCode="SNU" collectorName="Jinyeong Choi" country="Myanmar" latitude="21.380001" location="Ywangan" municipality="Ywangan" pageId="1" pageNumber="480" specimenCount="5" stateProvince="Shan State" typeStatus="holotype">
<specimenCount id="9D0DFD08FF82FFF7FCFDFA8DFC9BFA5F" box="[800,814,1405,1432]" pageId="1" pageNumber="480">5</specimenCount>
first-instar nymphs mounted together on 1 slide, same data as holotype (
<collectionCode id="ED1AAE44FF82FFF7FE81FA51FE2FFA7C" box="[348,410,1441,1467]" pageId="1" pageNumber="480">SNU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311650AFF82FFF7FF1AFA35FB96F98C" pageId="1" pageNumber="480" type="diagnosis">
<paragraph id="8BB43681FF82FFF7FF1AFA35FB96F98C" blockId="1.[151,1437,1333,1612]" pageId="1" pageNumber="480">
<emphasis id="B97FEA93FF82FFF7FF1AFA35FEF7FA18" bold="true" box="[199,322,1477,1503]" pageId="1" pageNumber="480">Diagnosis.</emphasis>
Legs with all segments much reduced in length. Sclerotized plates present on venter, forming bands or patches submedially on body. Dorsal abdominal clear areas present in 4 pairs. Antenna 57 segmented. Each stigmatic cleft with 1016 stigmatic spines. Dorsal marginal radial lines numbering about 22 around anterior margin of head, and each side with about 6 between stigmatic clefts and 18 on abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C311650AFF82FFF3FF4AF96DFB74FD0A" lastPageId="5" lastPageNumber="484" pageId="1" pageNumber="480" type="description">
<paragraph id="8BB43681FF82FFF7FF4AF96DFE9DF970" blockId="1.[151,296,1693,1719]" box="[151,296,1693,1719]" pageId="1" pageNumber="480">
<heading id="D0FC81EDFF82FFF7FF4AF96DFE9DF970" bold="true" box="[151,296,1693,1719]" fontSize="11" level="1" pageId="1" pageNumber="480" reason="1">
<emphasis id="B97FEA93FF82FFF7FF4AF96DFE9DF970" bold="true" box="[151,296,1693,1719]" pageId="1" pageNumber="480">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB43681FF82FFF7FF4AF915FF62F848" blockId="1.[151,1437,1765,2008]" pageId="1" pageNumber="480">
<emphasis id="B97FEA93FF82FFF7FF4AF915FE25F938" bold="true" box="[151,400,1765,1791]" pageId="1" pageNumber="480">Unmounted material</emphasis>
(
<figureCitation id="13302A04FF82FFF7FE43F915FE6AF938" box="[414,479,1765,1791]" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1</figureCitation>
). Body of adult female flat and broadly oval. Body camouflaged on bamboo, greenish or yellowish (
<figureCitation id="13302A04FF82FFF7FEECF8F9FE1FF8E4" box="[305,426,1801,1827]" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1B, C</figureCitation>
). Dorsal surface sclerotized, translucent and reticulated, with many indentations (
<figureCitation id="13302A04FF82FFF7FAEAF8F9FA3BF8E4" box="[1335,1422,1801,1827]" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1C</figureCitation>
). Ventral surface also slightly sclerotized, almost transparent but with white translucent areas submedially on body (
<figureCitation id="13302A04FF82FFF7FF42F8A1FF42F8AC" box="[159,247,1873,1899]" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1D</figureCitation>
); central concave brood space also present (
<figureCitation id="13302A04FF82FFF7FD3FF8A1FC8EF8AC" box="[738,827,1873,1899]" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1D</figureCitation>
). First-instar nymphs reddish, elongated oval (
<figureCitation id="13302A04FF82FFF7FA9FF8A1FF7EF848" captionStart="FIGURE 1" captionStartId="2.[152,255,1512,1536]" captionTargetBox="[164,1423,181,1488]" captionTargetId="figure-21@2.[164,1423,181,1488]" captionTargetPageId="2" captionText="FIGURE 1. Discochiton bambusae Choi &amp; Lee sp. n. A, the host plant, Bambusa tulda; B, population of adult females on a culm of B. tulda; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female." figureDoi="http://doi.org/10.5281/zenodo.10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="1" pageNumber="480">Fig. 1E, F, G</figureCitation>
).
</paragraph>
<paragraph id="8BB43681FF82FFF7FF1AF869FE6DF810" blockId="1.[151,1437,1765,2008]" pageId="1" pageNumber="480">
<emphasis id="B97FEA93FF82FFF7FF1AF869FDA7F874" bold="true" box="[199,530,1945,1971]" pageId="1" pageNumber="480">Slide-mounted adult female</emphasis>
(
<figureCitation id="13302A04FF82FFF7FDFFF869FDDCF874" box="[546,617,1945,1971]" captionStart="FIGURE 2" captionStartId="3.[152,255,1513,1537]" captionTargetBox="[164,1423,310,1489]" captionTargetId="figure-79@3.[164,1423,309,1490]" captionTargetPageId="3" captionText="FIGURE 2. Discochiton bambusae Choi &amp; Lee sp. n. A, slide-mounted adult female; B, stigmatic cleft and spines; C, anal plates; D, marginal setae; E, antenna; F, prothoracic leg; G, mesothoracic leg; H, metathoracic leg; I, long dorsal seta; J, short dorsal seta; K, Clear areas (CA14); L, clear areas and associated preopercular pores; M, sclerotized plates forming ventral band; N, submarginal sclerotized ridges. Scale lines for A = 5 mm; K= 1 mm; M = 0.5 mm; B, C, E, F, G, H, L = 100 μm; D = 50 μm; I, J, N = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.10010333" httpUri="https://zenodo.org/record/10010333/files/figure.png" pageId="1" pageNumber="480">Figs 2</figureCitation>
and
<figureCitation id="13302A04FF82FFF7FD7CF869FD04F874" box="[673,689,1945,1971]" captionStart="FIGURE 3" captionStartId="4.[152,255,1856,1880]" captionTargetBox="[210,1378,181,1832]" captionTargetId="figure-21@4.[210,1378,181,1832]" captionTargetPageId="4" captionText="FIGURE 3. Discochiton bambusae Choi &amp; Lee sp. n., adult female.A, reticulations; B, marginal radial lines indicated by pores and setae; C, marginal seta; D, stigmatic cleft and spines, and protrusions on outer margins of stigmatic cleft; E, submarginal sclerotized ridges; F, dorsal setae; G, preopercular pore; H, clear areas surrounded by preopercular pores; I, dorsal pores with a ductule and an unsclerotized pale area; J, anal plates; K, multilocular disc-pore; L, sclerotized plates forming a band; M, metathoracic leg; N, mesothoracic leg; O, prothoracic leg; P, spiracular disc-pore; Q, antenna." figureDoi="http://doi.org/10.5281/zenodo.10010335" httpUri="https://zenodo.org/record/10010335/files/figure.png" pageId="1" pageNumber="480">3</figureCitation>
). Body circular to oval (broadest across abdomen),
<quantity id="4CF39B64FF82FFF7FB24F869FA28F873" box="[1273,1437,1945,1972]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" metricValueMax="1.45" metricValueMin="1.01" pageId="1" pageNumber="480" unit="mm" value="12.3" valueMax="14.5" valueMin="10.1">10.114.5 mm</quantity>
long and
<quantity id="4CF39B64FF82FFF7FF22F84DFE26F81F" box="[255,403,1981,2008]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" metricValueMax="11.9" metricValueMin="7.9" pageId="1" pageNumber="480" unit="mm" value="9.9" valueMax="11.9" valueMin="7.9">7.911.9 mm</quantity>
wide.
</paragraph>
<caption id="DF746609FF81FFF4FF45FA18FA80F98F" ID-DOI="http://doi.org/10.5281/zenodo.10010330" ID-Zenodo-Dep="10010330" httpUri="https://zenodo.org/record/10010330/files/figure.png" pageId="2" pageNumber="481" startId="2.[152,255,1512,1536]" targetBox="[164,1423,181,1488]" targetPageId="2" targetType="figure">
<paragraph id="8BB43681FF81FFF4FF45FA18FA80F98F" blockId="2.[151,1436,1512,1609]" pageId="2" pageNumber="481">
<emphasis id="B97FEA93FF81FFF4FF45FA18FEACF9C7" bold="true" box="[152,281,1512,1536]" pageId="2" pageNumber="481">FIGURE 1.</emphasis>
<taxonomicName id="4C0B4D02FF81FFF4FEFCFA18FD3FF9C7" authority="Choi &amp; Lee" authorityName="Choi &amp; Lee" authorityYear="2023" box="[289,650,1512,1537]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="bambusae" status="sp. n.">
<emphasis id="B97FEA93FF81FFF4FEFCFA18FDBCF9C7" box="[289,521,1512,1536]" italics="true" pageId="2" pageNumber="481">Discochiton bambusae</emphasis>
Choi &amp; Lee
</taxonomicName>
<emphasis id="B97FEA93FF81FFF4FD4CFA19FD7EF9C6" bold="true" box="[657,715,1513,1537]" pageId="2" pageNumber="481">
<taxonomicNameLabel id="A24C57E8FF81FFF4FD4CFA19FD7EF9C6" box="[657,715,1513,1537]" pageId="2" pageNumber="481" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, the host plant,
<taxonomicName id="4C0B4D02FF81FFF4FC57FA18FB91F9C7" box="[906,1060,1512,1536]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="2" pageNumber="481" phylum="Tracheophyta" rank="species" species="tulda">
<emphasis id="B97FEA93FF81FFF4FC57FA18FB91F9C7" box="[906,1060,1512,1536]" italics="true" pageId="2" pageNumber="481">Bambusa tulda</emphasis>
</taxonomicName>
; B, population of adult females on a culm of
<taxonomicName id="4C0B4D02FF81FFF4FF36F9FDFE8EF9E3" box="[235,315,1548,1572]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="2" pageNumber="481" phylum="Tracheophyta" rank="species" species="tulda">
<emphasis id="B97FEA93FF81FFF4FF36F9FDFE8EF9E3" box="[235,315,1548,1572]" italics="true" pageId="2" pageNumber="481">B. tulda</emphasis>
</taxonomicName>
; C, dorsal view of adult female in life; D, ventral view of adult female in life; E, first-instar nymph; F, group of first-instar nymphs with plant debris; G, mass of first-instar nymphs left on the host after detaching the adult female.
</paragraph>
</caption>
<paragraph id="8BB43681FF81FFF5FF1AF96BFD98FEDA" blockId="2.[151,1437,1691,2041]" lastBlockId="3.[151,1437,150,285]" lastPageId="3" lastPageNumber="482" pageId="2" pageNumber="481">
<emphasis id="B97FEA93FF81FFF4FF1AF96BFE94F973" box="[199,289,1691,1716]" italics="true" pageId="2" pageNumber="481">Dorsum</emphasis>
. Derm uniformly lightly to heavily sclerotized, with abundant areolations and small pale spots, some forming a polygonal pattern medially. Stigmatic rays broad, extending medially from each stigmatic cleft. Marginal radial lines indistinct, indicated by distribution of pores and setae; with about 22 on head between anterior stigmatic clefts, and on each side with about 6 between stigmatic clefts and about 18 on abdomen. Dorsal setae mostly flagellate but also rarely spinose, variable in length, each 657 μm long, with a broad basal socket 710 μm wide; sparsely present. Abdomen with 4 pairs of grouped clear areas (CA) submedially, numbered from anterior to posterior (
<figureCitation id="13302A04FF81FFF4FAB2F8BFFF08F84A" captionStart="FIGURE 2" captionStartId="3.[152,255,1513,1537]" captionTargetBox="[164,1423,310,1489]" captionTargetId="figure-79@3.[164,1423,309,1490]" captionTargetPageId="3" captionText="FIGURE 2. Discochiton bambusae Choi &amp; Lee sp. n. A, slide-mounted adult female; B, stigmatic cleft and spines; C, anal plates; D, marginal setae; E, antenna; F, prothoracic leg; G, mesothoracic leg; H, metathoracic leg; I, long dorsal seta; J, short dorsal seta; K, Clear areas (CA14); L, clear areas and associated preopercular pores; M, sclerotized plates forming ventral band; N, submarginal sclerotized ridges. Scale lines for A = 5 mm; K= 1 mm; M = 0.5 mm; B, C, E, F, G, H, L = 100 μm; D = 50 μm; I, J, N = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.10010333" httpUri="https://zenodo.org/record/10010333/files/figure.png" pageId="2" pageNumber="481">Fig. 2K</figureCitation>
), each group surrounded by preopercular pores, each pore roundly convex, 36 μm wide, present as follows (number of clear areas in each group in brackets): CA1 with 2240 (14), CA2 2548 (13), CA3 3143 (24) and CA4 with 2940 pores (13). Other dorsal pores each 23 μm wide, each consisting of a ductule associated with an unsclerotized pale area, these pores forming a polygonal pattern medially and present in marginal radial lines with setae. Anal plates rather elongate, together quadrate, length of plates 251293 μm, combined width 216290 μm; each plate with anterior margin usually shorter than posterior margin, with 4 minute setae near posterior apex and 5 or 6 small pores medially. Anogenital fold with 3 or 4 minute setae at each corner with anterior margin, and each lateral margin with 1 seta anteriorly.
</paragraph>
<caption id="DF746609FF80FFF5FF45FA19FE33F955" ID-DOI="http://doi.org/10.5281/zenodo.10010333" ID-Zenodo-Dep="10010333" httpUri="https://zenodo.org/record/10010333/files/figure.png" pageId="3" pageNumber="482" startId="3.[152,255,1513,1537]" targetBox="[164,1423,310,1489]" targetPageId="3" targetType="figure">
<paragraph id="8BB43681FF80FFF5FF45FA19FE33F955" blockId="3.[151,1437,1513,1682]" pageId="3" pageNumber="482">
<emphasis id="B97FEA93FF80FFF5FF45FA19FEAFF9C6" bold="true" box="[152,282,1513,1537]" pageId="3" pageNumber="482">FIGURE 2.</emphasis>
<taxonomicName id="4C0B4D02FF80FFF5FEFFFA19FD38F9C6" authority="Choi &amp; Lee" authorityName="Choi &amp; Lee" authorityYear="2023" box="[290,653,1513,1538]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="bambusae" status="sp. n.">
<emphasis id="B97FEA93FF80FFF5FEFFFA19FDBEF9C6" box="[290,523,1513,1537]" italics="true" pageId="3" pageNumber="482">Discochiton bambusae</emphasis>
Choi &amp; Lee
</taxonomicName>
<emphasis id="B97FEA93FF80FFF5FD48FA1AFD65F9C5" bold="true" box="[661,720,1514,1538]" pageId="3" pageNumber="482">
<taxonomicNameLabel id="A24C57E8FF80FFF5FD48FA1AFD65F9C5" box="[661,720,1514,1538]" pageId="3" pageNumber="482" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, slide-mounted adult female; B, stigmatic cleft and spines; C, anal plates; D, marginal setae; E, antenna; F, prothoracic leg; G, mesothoracic leg; H, metathoracic leg; I, long dorsal seta; J, short dorsal seta; K, Clear areas (CA14); L, clear areas and associated preopercular pores; M, sclerotized plates forming ventral band; N, submarginal sclerotized ridges. Scale lines for A = 5 mm; K= 1 mm; M = 0.5 mm; B, C, E, F, G, H, L = 100 μm; D = 50 μm; I, J, N = 10 μm.
</paragraph>
</caption>
<paragraph id="8BB43681FF80FFF5FF1AF914FB81F811" blockId="3.[151,1436,1764,2006]" pageId="3" pageNumber="482">
<emphasis id="B97FEA93FF80FFF5FF1AF914FEAFF93A" box="[199,282,1764,1789]" italics="true" pageId="3" pageNumber="482">Margin</emphasis>
. Outer margins of stigmatic clefts with distinct irregular protrusions. Sclerotized ridges present submarginally, shallow, parallel with margin or extending inwards. Marginal setae broadly fan-shaped, each fan-shaped seta wider than long, about 2853 μm wide and 1839 μm long; numbering about 189283 setae anteriorly on head between anterior stigmatic clefts, and each side with 87107 between stigmatic clefts and 177230 along abdominal margin. Stigmatic clefts quite deep, each with a narrow entrance, a highly sclerotized inner margin and 1016 (usually 1114) elongate, parallel-sided, blunt stigmatic spines. Eyespots displaced some distance from margin, each situated in a socket 4168 μm wide, each lens 2137 μm in diameter.
</paragraph>
<caption id="DF746609FF87FFF2FF45F8B0FBF8F82F" ID-DOI="http://doi.org/10.5281/zenodo.10010335" ID-Zenodo-Dep="10010335" httpUri="https://zenodo.org/record/10010335/files/figure.png" pageId="4" pageNumber="483" startId="4.[152,255,1856,1880]" targetBox="[210,1378,181,1832]" targetPageId="4" targetType="figure">
<paragraph id="8BB43681FF87FFF2FF45F8B0FBF8F82F" blockId="4.[151,1436,1856,2024]" pageId="4" pageNumber="483">
<emphasis id="B97FEA93FF87FFF2FF45F8B0FEA2F89F" bold="true" box="[152,279,1856,1880]" pageId="4" pageNumber="483">FIGURE 3.</emphasis>
<taxonomicName id="4C0B4D02FF87FFF2FEC0F8B0FDCBF89F" authority="Choi &amp; Lee" authorityName="Choi &amp; Lee" authorityYear="2023" box="[285,638,1856,1880]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="483" phylum="Arthropoda" rank="species" species="bambusae" status="sp. n.">
<emphasis id="B97FEA93FF87FFF2FEC0F8B0FDB6F89F" box="[285,515,1856,1880]" italics="true" pageId="4" pageNumber="483">Discochiton bambusae</emphasis>
Choi &amp; Lee
</taxonomicName>
<emphasis id="B97FEA93FF87FFF2FD5EF8B0FD0EF89F" bold="true" box="[643,699,1856,1880]" pageId="4" pageNumber="483">
<taxonomicNameLabel id="A24C57E8FF87FFF2FD5EF8B0FD0EF89F" box="[643,699,1856,1880]" pageId="4" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female. A, reticulations; B, marginal radial lines indicated by pores and setae; C, marginal seta; D, stigmatic cleft and spines, and protrusions on outer margins of stigmatic cleft; E, submarginal sclerotized ridges; F, dorsal setae; G, preopercular pore; H, clear areas surrounded by preopercular pores; I, dorsal pores with a ductule and an unsclerotized pale area; J, anal plates; K, multilocular disc-pore; L, sclerotized plates forming a band; M, metathoracic leg; N, mesothoracic leg; O, prothoracic leg; P, spiracular disc-pore; Q, antenna.
</paragraph>
</caption>
<paragraph id="8BB43681FF86FFF3FF1AFF67FB74FD0A" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="484">
<emphasis id="B97FEA93FF86FFF3FF1AFF67FEBAFF77" box="[199,271,151,176]" italics="true" pageId="5" pageNumber="484">Venter</emphasis>
. Derm membranous only in medial area; rest of derm lightly to heavily sclerotized, with sclerotized plates forming a band submedially; with a cluster of plates just laterad to each antenna, a similar-sized cluster of plates just laterad to each mesothoracic leg, and a much longer band of plates on abdomen, extending from area laterad to each metacoxa to anal cleft (these plates larger and more distinct on older specimens)); sclerotized band on each side of thorax about
<quantity id="4CF39B64FF86FFF3FE00FED7FD8FFE86" box="[477,570,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="5" pageNumber="484" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
wide. Multilocular disc-pores abundant on either side of genital opening and on preceding segment only, each side with 61104 on segment VII and 74150 on segment VI. Spiracular disc-pores present between margin and each spiracle in a narrow band 15 pores wide, with 109150 pores in each anterior band and 98168 pores in each posterior band. Ventral microducts each 23 μm wide, densely present just posterior to and on either side of labium, along with several setae; also, sparsely present elsewhere. Ventral setae: a pair of long setae present medially on segment VII, each seta 6890 μm long; and short to long setae frequently present on medial areas of abdominal segments VIV, each 871 μm long; otherwise, setae short and very sparse; interantennal setae absent. Antennae reduced, each with 57 segments but occasionally with a pseudo-articulation in segment III; total length 231350 μm. Clypeolabral shield 291397 μm long. Spiracles: width of each anterior peritreme 99112 μm and posterior peritreme 96119 μm. Legs present but very small, each with 5 segments much reduced in length, each segment unclearly divided, claw denticles absent; claw digitules and tarsal digitules narrow; total length of prothoracic leg 167206 μm, mesothoracic leg 115243 μm and metathoracic leg 137227 μm.
</paragraph>
</subSubSection>
<subSubSection id="C311650AFF86FFF3FF1AFD26FC71FBF2" pageId="5" pageNumber="484" type="discussion">
<paragraph id="8BB43681FF86FFF3FF1AFD26FC71FBF2" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="484">
<emphasis id="B97FEA93FF86FFF3FF1AFD26FEF1FD37" bold="true" box="[199,324,726,752]" pageId="5" pageNumber="484">Comment.</emphasis>
The adult female of
<taxonomicName id="4C0B4D02FF86FFF3FDF7FD27FD0BFD36" authority="Choi &amp; Lee, 2023" authorityName="Choi &amp; Lee" authorityYear="2023" box="[554,702,727,753]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="bambusae" status="sp. n.">
<emphasis id="B97FEA93FF86FFF3FDF7FD27FD0BFD36" box="[554,702,727,753]" italics="true" pageId="5" pageNumber="484">D. bambusae</emphasis>
</taxonomicName>
<emphasis id="B97FEA93FF86FFF3FD19FD27FCB7FD36" bold="true" box="[708,770,727,753]" pageId="5" pageNumber="484">
<taxonomicNameLabel id="A24C57E8FF86FFF3FD19FD27FCB7FD36" box="[708,770,727,753]" pageId="5" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from all other known
<taxonomicName id="4C0B4D02FF86FFF3FB95FD27FA93FD36" box="[1096,1318,727,753]" pageId="5" pageNumber="484">
<emphasis id="B97FEA93FF86FFF3FB95FD27FB7AFD36" box="[1096,1231,727,753]" italics="true" pageId="5" pageNumber="484">Discochiton</emphasis>
species
</taxonomicName>
in having: (i) a much larger body; (ii) a ventral band of sclerotisations in submedial areas of body; (iii) many stigmatic setae in each stigmatic cleft; and (iv) many marginal setae. While the morphology of the new species agrees with most character-states of
<taxonomicName id="4C0B4D02FF86FFF3FEB8FCB3FE59FC9A" authorityName="Hodgson &amp; Williams" authorityYear="2018" box="[357,492,835,861]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEA93FF86FFF3FEB8FCB3FE59FC9A" box="[357,492,835,861]" italics="true" pageId="5" pageNumber="484">Discochiton</emphasis>
</taxonomicName>
, the adult female displays a greater number of marginal radial lines and more stigmatic spines than are found on other
<taxonomicName id="4C0B4D02FF86FFF3FE2FFC97FD6CFC46" box="[498,729,871,897]" class="Insecta" family="Coccidae" genus="Discochiton" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97FEA93FF86FFF3FE2FFC97FDCCFC46" box="[498,633,871,897]" italics="true" pageId="5" pageNumber="484">Discochiton</emphasis>
species.
</taxonomicName>
This may be due to the large body size of
<taxonomicName id="4C0B4D02FF86FFF3FB6BFC97FAFFFC46" authorityName="Choi &amp; Lee" authorityYear="2023" box="[1206,1354,871,897]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="bambusae">
<emphasis id="B97FEA93FF86FFF3FB6BFC97FAFFFC46" box="[1206,1354,871,897]" italics="true" pageId="5" pageNumber="484">D. bambusae</emphasis>
</taxonomicName>
, which is the largest species in the genus, reaching approximately
<quantity id="4CF39B64FF86FFF3FCEAFC7AFC32FC62" box="[823,903,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="484" unit="mm" value="15.0">15 mm</quantity>
long. In addition, the legs of
<taxonomicName id="4C0B4D02FF86FFF3FB01FC7BFAC7FC62" authorityName="Choi &amp; Lee" authorityYear="2023" box="[1244,1394,907,933]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="bambusae">
<emphasis id="B97FEA93FF86FFF3FB01FC7BFAC7FC62" box="[1244,1394,907,933]" italics="true" pageId="5" pageNumber="484">D. bambusae</emphasis>
</taxonomicName>
are much reduced (similar to those of
<taxonomicName id="4C0B4D02FF86FFF3FDCBFC5FFC89FC0E" authority="(Hodgson))" baseAuthorityName="Hodgson" box="[534,828,943,969]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="paucipedis">
<emphasis id="B97FEA93FF86FFF3FDCBFC5FFD04FC0E" box="[534,689,943,969]" italics="true" pageId="5" pageNumber="484">D. paucipedis</emphasis>
(Hodgson))
</taxonomicName>
, whereas other species of
<taxonomicName id="4C0B4D02FF86FFF3FB82FC5FFB53FC0E" authorityName="Hodgson &amp; Williams" authorityYear="2018" box="[1119,1254,943,969]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEA93FF86FFF3FB82FC5FFB53FC0E" box="[1119,1254,943,969]" italics="true" pageId="5" pageNumber="484">Discochiton</emphasis>
</taxonomicName>
show only fully developed or completely reduced legs. Moreover, the sclerotized plate-like areas on the venter of
<taxonomicName id="4C0B4D02FF86FFF3FB22FC23FA23FC2A" authorityName="Choi &amp; Lee" authorityYear="2023" box="[1279,1430,979,1005]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="bambusae">
<emphasis id="B97FEA93FF86FFF3FB22FC23FA23FC2A" box="[1279,1430,979,1005]" italics="true" pageId="5" pageNumber="484">D. bambusae</emphasis>
</taxonomicName>
, which extend in a submedial band from the antennae to the anal cleft, are unique and have not been recorded previously in other species of
<taxonomicName id="4C0B4D02FF86FFF3FE35FBEBFC75FBF2" authority="(Hodgson &amp; Williams, 2018)" baseAuthorityName="Hodgson &amp; Williams" baseAuthorityYear="2018" box="[488,960,1051,1077]" class="Insecta" family="Coccidae" genus="Discochiton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEA93FF86FFF3FE35FBEBFDDAFBF2" box="[488,623,1051,1077]" italics="true" pageId="5" pageNumber="484">Discochiton</emphasis>
(
<bibRefCitation id="EF9A4B70FF86FFF3FDA3FBEBFC0DFBF2" author="Hodgson, C. J. &amp; Williams, D. J." box="[638,952,1051,1077]" pageId="5" pageNumber="484" pagination="1 - 162" refId="ref4181" refString="Hodgson, C. J. &amp; Williams, D. J. (2018) Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species. Zootaxa, 4443 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 4443.1.1" type="journal article" year="2018">Hodgson &amp; Williams, 2018</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>