317 lines
42 KiB
XML
317 lines
42 KiB
XML
<document id="B522E202001EE99A60C6671076E9348D" ID-DOI="10.11646/zootaxa.4755.1.4" ID-GBIF-Dataset="70aae18d-e6ec-4bb1-abe0-27d1f24fbab8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724261" ID-ZooBank="53E41147-52AA-4A16-BB69-3A9279A303AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968507449" checkinUser="plazi" docAuthor="Barr, Cheryl B. & Shepard, William D." docDate="2020" docId="039CB74BFFD63A5BF9DFFCD1FBE49061" docLanguage="en" docName="zootaxa.4755.1.4.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parygrus angustatus Grouvelle 1896" docType="treatment" docVersion="5" lastPageNumber="108" masterDocId="FFA5CF33FFDE3A52F948FFB8FF87956E" masterDocTitle="Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)" masterLastPageNumber="128" masterPageNumber="99" pageNumber="107" updateTime="1698730760387" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0B2B725AB68EC47AB8B7B635984CC912">Hiding in plain sight: rediscovery and review of Parygrus Erichson, 1847, with description of five new species from the Neotropics (Coleoptera: Byrrhoidea Dryopidae)</mods:title>
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<mods:namePart id="ACCFF639FE927BE2C4C51645B5394D1C">Barr, Cheryl B.</mods:namePart>
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<mods:namePart id="E4F40A477A80996B8D1F3C1BA59F9D0B">Shepard, William D.</mods:namePart>
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<mods:title id="DF10470250A2C7899554038FAEAC9A50">Zootaxa</mods:title>
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<mods:date id="AD67B6DA11B26BD6A48ACB034EB199AA">2020</mods:date>
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<mods:number id="936CDF0423F7C6723DB6CEAF367B3756">4755</mods:number>
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<treatment id="039CB74BFFD63A5BF9DFFCD1FBE49061" ID-DOI="http://doi.org/10.5281/zenodo.3730221" ID-GBIF-Taxon="162696912" ID-Zenodo-Dep="3730221" LSID="urn:lsid:plazi:treatment:039CB74BFFD63A5BF9DFFCD1FBE49061" httpUri="http://treatment.plazi.org/id/039CB74BFFD63A5BF9DFFCD1FBE49061" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
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<subSubSection id="C32F55D6FFD63A5AF9DFFCD1FEA096C9" pageId="8" pageNumber="107" type="nomenclature">
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<paragraph id="8B8A065DFFD63A5AF9DFFCD1FDD996ED" blockId="8.[151,606,873,935]" box="[151,606,873,899]" pageId="8" pageNumber="107">
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<heading id="D0C2B131FFD63A5AF9DFFCD1FDD996ED" bold="true" box="[151,606,873,899]" fontSize="11" level="1" pageId="8" pageNumber="107" reason="1">
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<taxonomicName id="4C357DDEFFD63A5AF9DFFCD1FDD996ED" authority="Grouvelle, 1896" authorityName="Grouvelle" authorityYear="1896" box="[151,606,873,899]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="B941DA4FFFD63A5AF9DFFCD1FDD996ED" bold="true" box="[151,606,873,899]" pageId="8" pageNumber="107">
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<emphasis id="B941DA4FFFD63A5AF9DFFCD1FE1696ED" bold="true" box="[151,401,873,899]" italics="true" pageId="8" pageNumber="107">Parygrus angustatus</emphasis>
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Grouvelle, 1896
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8A065DFFD63A5AF9DFFC35FEA096C9" blockId="8.[151,606,873,935]" box="[151,295,909,935]" pageId="8" pageNumber="107">(Figs 2, 8, 9)</paragraph>
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</subSubSection>
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<subSubSection id="C32F55D6FFD63A5BF9DFFC6DFD139471" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107" type="materials_examined">
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<paragraph id="8B8A065DFFD63A5AF9DFFC6DFAA59135" blockId="8.[151,1437,980,2015]" pageId="8" pageNumber="107">
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<materialsCitation id="3B5D0C00FFD63A5AF9DFFC6DFAA59135" ID-GBIF-Occurrence="2579576872" collectionCode="MNHN" collectorName="R. de Jan & Coll. Grouvelle" country="Brazil" location="Rio de Janeiro" pageId="8" pageNumber="107" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
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<emphasis id="B941DA4FFFD63A5AF9DFFC6DFE7A9681" bold="true" box="[151,509,980,1007]" pageId="8" pageNumber="107">
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<typeStatus id="548EB8FFFFD63A5AF9DFFC6DFE839681" box="[151,260,981,1007]" pageId="8" pageNumber="107" type="holotype">Holotype</typeStatus>
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specimen.
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<collectingCountry id="F32246CDFFD63A5AF8C0FC6CFE7F9681" box="[392,504,980,1007]" name="Brazil" pageId="8" pageNumber="107">BRAZIL</collectingCountry>
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:
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</emphasis>
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“Bresil /
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<collectorName id="26C0638BFFD63A5AFB22FC6DFD569681" box="[618,721,981,1007]" pageId="8" pageNumber="107">R. de Jan</collectorName>
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[blue-green, handwritten] // Museum Paris / 1917 /
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<collectorName id="26C0638BFFD63A5AFC53FC6DFF4A917D" pageId="8" pageNumber="107">Coll. Grouvelle</collectorName>
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[white, printed] // TYPE [red, printed] //
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<taxonomicName id="4C357DDEFFD63A5AFBEDFC41FD70917D" authorityName="Olivier" authorityYear="1791" box="[677,759,1017,1043]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="126" phylum="Arthropoda" rank="genus">Dryops</taxonomicName>
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/angustatus / ty. Grouv [off-white, handwritten]”. In his description, Grouvelle stated “
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<collectingRegion id="49F1C8BFFFD63A5AF89AFBA5FDFC9159" box="[466,635,1053,1079]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="107">Rio de Janeiro</collectingRegion>
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. Collection Grouvelle” (
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<bibRefCitation id="EFA47BACFFD63A5AFAC4FBA5FBCA9159" author="Grouvelle, A." box="[908,1101,1053,1079]" pageId="8" pageNumber="107" pagination="177 - 216" refId="ref21501" refString="Grouvelle, A. (1896 a) Nitidulides, Colydiides, Cucujides et Parnides recoltes par M. E. Gounelle au Bresil et autres Clavicornes nouveaux d'Amerique. Annales de la Societe Entomologique de France, 65, 177 - 216." type="journal article" year="1896">Grouvelle 1896a</bibRefCitation>
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, p. 215). Depository,
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<collectionCode id="ED249E98FFD63A5AFC08FBA5FA1E9159" box="[1344,1433,1053,1079]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="107">MNHN</collectionCode>
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. The type specimen itself was not examined, only the photographic image and label data from the photo.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8A065DFFD63A5AF98FFBDCFC129185" blockId="8.[151,1437,980,2015]" pageId="8" pageNumber="107">
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<emphasis id="B941DA4FFFD63A5AF98FFBDCFC819111" bold="true" box="[199,774,1124,1151]" pageId="8" pageNumber="107">Additional specimens, photographic images (2).</emphasis>
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<materialsCitation id="3B5D0C00FFD63A5AFA47FBDCFC159185" ID-GBIF-Occurrence="2579576874" collectingDate="1897-09-11" collectionCode="MNHN" collectorName="Coll. A. Grouvelle & F. Wiengreen" country="Brazil" location="BRAZIL" pageId="8" pageNumber="107" specimenCount="1">
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<emphasis id="B941DA4FFFD63A5AFA47FBDCFC039111" bold="true" box="[783,900,1124,1151]" pageId="8" pageNumber="107">
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<collectingCountry id="F32246CDFFD63A5AFA47FBDCFCF89111" box="[783,895,1124,1151]" name="Brazil" pageId="8" pageNumber="107">BRAZIL</collectingCountry>
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:
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</emphasis>
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“Bresil / Theresop [handwritten] // MUSEUM PARIS [light blue, printed] /
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<collectorName id="26C0638BFFD63A5AF897FB31FD3791CD" box="[479,688,1161,1187]" pageId="8" pageNumber="107">Coll. A. Grouvelle</collectorName>
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1915 //
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<taxonomicName id="4C357DDEFFD63A5AFA47FB31FCF491CD" authorityName=", Erichson" authorityYear="1847" box="[783,883,1161,1187]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="genus">Parygrus</taxonomicName>
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/ angustatus / gr. [handwritten]”; “Neu-Freiburg, / Süd-Brasilien. /
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<collectorName id="26C0638BFFD63A5AF828FB15FE7191A9" box="[352,502,1197,1223]" pageId="8" pageNumber="107">F. Wiengreen</collectorName>
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leg. / ded.
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<date id="FF8B209DFFD63A5AFB3EFB15FD7791A9" box="[630,752,1197,1223]" pageId="8" pageNumber="107" value="1897-09-11">
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<collectingDate id="EFCFD975FFD63A5AFB3EFB15FD7791A9" box="[630,752,1197,1223]" pageId="8" pageNumber="107" value="1897-09-11">9.11. 1897</collectingDate>
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</date>
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. //
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<taxonomicName id="4C357DDEFFD63A5AFA5CFB15FC7191A9" authorityName="Grouvelle" authorityYear="1896" box="[788,1014,1197,1223]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="angustatus">Parygrus angustatus</taxonomicName>
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/ Grouv.”. Depository,
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<collectionCode id="ED249E98FFD63A5AFC48FB15FADE91A9" box="[1280,1369,1197,1223]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="107">MNHN</collectionCode>
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. Only photographic images and label data from the photos were examined
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8A065DFFD63A5AF98FFB4CFE519341" blockId="8.[151,1437,980,2015]" pageId="8" pageNumber="107">
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<materialsCitation id="3B5D0C00FFD63A5AF98FFB4CFA1090AD" ID-GBIF-Occurrence="2579576863" country="Brazil" location="Grouv" pageId="8" pageNumber="107" specimenCount="2">
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<emphasis id="B941DA4FFFD63A5AF98FFB4CFD6E9061" bold="true" box="[199,745,1268,1295]" pageId="8" pageNumber="107">
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Additional specimens examined (7).
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<collectingCountry id="F32246CDFFD63A5AFB3CFB4CFD639061" box="[628,740,1268,1295]" name="Brazil" pageId="8" pageNumber="107">BRAZIL</collectingCountry>
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||
:
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</emphasis>
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“
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<typeStatus id="548EB8FFFFD63A5AFBB0FB4DFCB49061" box="[760,819,1269,1295]" pageId="8" pageNumber="107">Type</typeStatus>
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[circular label with red border, printed] // 20295 [handwritten] // Fry / Rio Jan°. [handwritten] // Fry Coll. / 1905-100.” [printed] // “
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<taxonomicName id="4C357DDEFFD63A5AFD63FAA1FA95905D" authorityName="Grouvelle" authorityYear="1896" box="[1067,1298,1305,1331]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="angustatus">Parygrus angustatus</taxonomicName>
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/ ty.
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<location id="8EEA5086FFD63A5AFC1DFAA1FA1A905D" LSID="urn:lsid:plazi:treatment:039CB74BFFD63A5BF9DFFCD1FBE49061:8EEA5086FFD63A5AFC1DFAA1FA1A905D" box="[1365,1437,1305,1331]" country="Brazil" name="Grouv" pageId="8" pageNumber="107">Grouv</location>
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[handwritten]” (1; head missing); “Fry / Rio Jan°. [handwritten] // Fry Coll. / 1905-100. [printed]” (2); “Fry / Rio Jan°. [handwritten] // Fry Coll. / 1905-100. [printed] //
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<taxonomicName id="4C357DDEFFD63A5AFA4AFAD9FCE19015" authorityName=", Erichson" authorityYear="1847" box="[770,870,1377,1403]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="genus">Parygrus</taxonomicName>
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/ angustatus [handwritten]” (1; right elytron missing); “9354 // Fry / Rio Jan°. [handwritten] // Fry Coll. / 1905-100. [printed] //
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<taxonomicName id="4C357DDEFFD63A5AFD44FA3DFBF790F1" authorityName=", Erichson" authorityYear="1847" box="[1036,1136,1413,1439]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="genus">Parygrus</taxonomicName>
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/ angustatus / var. [?-illegible word] Grouv [handwritten]” (1); “Rio [pink oval label, handwritten] // Pascoe / Coll. / 93–60.” [printed] (1)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B5D0C00FFD63A5AF9DFFA74FE559341" ID-GBIF-Occurrence="2579576869" collectingDate="1924-12" collectionCode="NHMUK" collectorName="K. J. Hayward." country="Argentina" location="Discussion" pageId="8" pageNumber="107" specimenCount="1">
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<emphasis id="B941DA4FFFD63A5AF9DFFA74FECB9089" bold="true" box="[151,332,1484,1511]" pageId="8" pageNumber="107">
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||
<collectingCountry id="F32246CDFFD63A5AF9DFFA74FEC09089" box="[151,327,1484,1511]" name="Argentina" pageId="8" pageNumber="107">ARGENTINA</collectingCountry>
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||
:
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</emphasis>
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“Argentine Republic: / Villa Ana, F. C. S. Fe. /
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<date id="FF8B209DFFD63A5AFA3EFA75FBAA9089" box="[886,1069,1485,1511]" pageId="8" pageNumber="107" value="1924-12">
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||
<collectingDate id="EFCFD975FFD63A5AFA3EFA75FBAA9089" box="[886,1069,1485,1511]" pageId="8" pageNumber="107" value="1924-12">December 1924</collectingDate>
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</date>
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. /
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<collectorName id="26C0638BFFD63A5AFD02FA75FB709089" box="[1098,1271,1485,1511]" pageId="8" pageNumber="107">K. J. Hayward.</collectorName>
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// Brit. Mus. / 1925-217” [printed] (1). Depository,
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<collectionCode id="ED249E98FFD63A5AFB7AFA49FD279365" box="[562,672,1521,1547]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="107" type="Museum">NHMUK</collectionCode>
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||
. These specimens may not be conspecific with the
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<typeStatus id="548EB8FFFFD63A5AFD98FA49FA879365" box="[1232,1280,1521,1547]" pageId="8" pageNumber="107">type</typeStatus>
|
||
nor with each other; see
|
||
<location id="8EEA5086FFD63A5AF842F9ADFE029341" LSID="urn:lsid:plazi:treatment:039CB74BFFD63A5BF9DFFCD1FBE49061:8EEA5086FFD63A5AF842F9ADFE029341" box="[266,389,1557,1583]" country="Argentina" name="Discussion" pageId="8" pageNumber="107">Discussion</location>
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||
below
|
||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B8A065DFFD63A5AF98FF980FDDD9242" blockId="8.[151,1437,980,2015]" pageId="8" pageNumber="107">
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||
<emphasis id="B941DA4FFFD63A5AF98FF980FE4D933D" bold="true" box="[199,458,1592,1619]" pageId="8" pageNumber="107">Provisional diagnosis.</emphasis>
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||
The
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<typeStatus id="548EB8FFFFD63A5AFB49F981FDB6933D" box="[513,561,1593,1619]" pageId="8" pageNumber="107">type</typeStatus>
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||
specimen of
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<taxonomicName id="4C357DDEFFD63A5AFB8CF982FCD0933D" authorityName="Grouvelle" authorityYear="1896" box="[708,855,1594,1619]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="angustatus">
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||
<emphasis id="B941DA4FFFD63A5AFB8CF982FCD0933D" box="[708,855,1594,1619]" italics="true" pageId="8" pageNumber="107">P. angustatus</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="130E1AD8FFD63A5AFA2EF981FC20933D" box="[870,935,1593,1619]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="8" pageNumber="107">Fig. 8</figureCitation>
|
||
) is much longer (
|
||
<quantity id="4CCDABB8FFD63A5AFD21F981FB40933D" box="[1129,1223,1593,1619]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="8" pageNumber="107" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||
) than specimens of any of the other species except for
|
||
<taxonomicName id="4C357DDEFFD63A5AFB69F9E6FD289319" baseAuthorityName="Grouvelle" baseAuthorityYear="1890" box="[545,687,1629,1655]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B941DA4FFFD63A5AFB69F9E6FD289319" box="[545,687,1629,1655]" italics="true" pageId="8" pageNumber="107">P. parallelus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130E1AD8FFD63A5AFBF6F9E5FC979319" box="[702,784,1629,1655]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="8" pageNumber="107">Fig. 11</figureCitation>
|
||
). Keeping in mind that the images of
|
||
<taxonomicName id="4C357DDEFFD63A5AFDF2F9E6FACF9319" baseAuthorityName="Grouvelle" baseAuthorityYear="1890" box="[1210,1352,1629,1655]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B941DA4FFFD63A5AFDF2F9E6FACF9319" box="[1210,1352,1629,1655]" italics="true" pageId="8" pageNumber="107">P. parallelus</emphasis>
|
||
</taxonomicName>
|
||
are not of the
|
||
<typeStatus id="548EB8FFFFD63A5AF995F939FE9793F5" box="[221,272,1665,1691]" pageId="8" pageNumber="107">type</typeStatus>
|
||
,
|
||
<taxonomicName id="4C357DDEFFD63A5AF852F93AFE2993F5" authorityName="Grouvelle" authorityYear="1896" box="[282,430,1666,1691]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD63A5AF852F93AFE2993F5" box="[282,430,1666,1691]" italics="true" pageId="8" pageNumber="107">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
appears to differ from that species by the following: elytra parallel-sided (vs. sinuate laterally, not parallel-sided); pronotum widest in about a third of the distance from the base (vs. widest at base), lateral margins not raised nor widely explanate (vs. raised, widely explanate), with a weak longitudinal carina (not visible in
|
||
<figureCitation id="130E1AD8FFD63A5AF9FCF955FF719269" box="[180,246,1773,1799]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="8" pageNumber="107">Fig. 8</figureCitation>
|
||
) (vs. without a carina); elytral striae not distinct, punctures small (vs. striae distinct, punctures larger), effaced at posterior third (vs. not effaced).
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFD63A5BF98FF88CFD139471" blockId="8.[151,1437,980,2015]" lastBlockId="9.[151,1437,153,1295]" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
|
||
<emphasis id="B941DA4FFFD63A5AF98FF88CFE399221" bold="true" box="[199,446,1844,1871]" pageId="8" pageNumber="107">General description.</emphasis>
|
||
Since we were unable to examine the
|
||
<typeStatus id="548EB8FFFFD63A5AFAC9F88DFC619221" box="[897,998,1845,1871]" pageId="8" pageNumber="107" type="holotype">holotype</typeStatus>
|
||
, only a limited description is possible based on the original descriptions (
|
||
<bibRefCitation id="EFA47BACFFD63A5AFB65F8E1FD68921A" author="Grouvelle, A." box="[557,751,1881,1908]" pageId="8" pageNumber="107" pagination="177 - 216" refId="ref21501" refString="Grouvelle, A. (1896 a) Nitidulides, Colydiides, Cucujides et Parnides recoltes par M. E. Gounelle au Bresil et autres Clavicornes nouveaux d'Amerique. Annales de la Societe Entomologique de France, 65, 177 - 216." type="journal article" year="1896">Grouvelle 1896a</bibRefCitation>
|
||
) and images of the type (
|
||
<figureCitation id="130E1AD8FFD63A5AFD54F8E1FBE4921D" box="[1052,1123,1881,1907]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="8" pageNumber="107">Fig. 8</figureCitation>
|
||
). From a translation of the original Latin and French descriptions:
|
||
<quantity id="4CCDABB8FFD63A5AFB07F8C5FD2B92F9" box="[591,684,1917,1943]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="8" pageNumber="107" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||
long, elongate-oval; convex; pronotum widest at basal third, with a weak longitudinal carina (subcarinate), anterior margin not rimmed in the middle, lateral borders arcuate;
|
||
<taxonomicName id="4C357DDEFFD63A5AFC67F819FA1B92D5" box="[1327,1436,1953,1979]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="8" pageNumber="123" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
curvilinear triangular; elytra a little wider than pronotum at base, punctate-striate, striae effaced at posterior third. Additional observations from the image of the type: body is parallel-sided and tapered posteriorly; pronotal posterolateral angles are depressed, if not explanate; median longitudinal pronotal carina is not discernable in the
|
||
<figureCitation id="130E1AD8FFD73A5BFC38FF05FF209595" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 8</figureCitation>
|
||
; elytra punctate but not obviously striate, punctures small and shallow; protibiae deflexed and enlarged at apical third, indicating that the type is likely a male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFD73A5BF98FFE90FB0E942D" box="[199,1161,296,324]" pageId="9" pageNumber="108" type="distribution">
|
||
<paragraph id="8B8A065DFFD73A5BF98FFE90FB0E942D" blockId="9.[151,1437,153,1295]" box="[199,1161,296,324]" pageId="9" pageNumber="108">
|
||
<emphasis id="B941DA4FFFD73A5BF98FFE90FEE6942D" bold="true" box="[199,353,296,323]" pageId="9" pageNumber="108">Distribution.</emphasis>
|
||
<collectingCountry id="F32246CDFFD73A5BF82FFE91FE2C942D" box="[359,427,297,323]" name="Brazil" pageId="9" pageNumber="108">Brazil</collectingCountry>
|
||
(described from
|
||
<collectingRegion id="49F1C8BFFFD73A5BFB24FE91FC95942A" box="[620,786,297,324]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="108">Rio de Janeiro</collectingRegion>
|
||
), and possibly
|
||
<collectingCountry id="F32246CDFFD73A5BFAFFFE91FBA1942D" box="[951,1062,297,323]" name="Argentina" pageId="9" pageNumber="108">Argentina</collectingCountry>
|
||
(Fig. 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFD73A5BF98FFEF5FB8994E5" pageId="9" pageNumber="108" type="etymology">
|
||
<paragraph id="8B8A065DFFD73A5BF98FFEF5FB8994E5" blockId="9.[151,1437,153,1295]" pageId="9" pageNumber="108">
|
||
<emphasis id="B941DA4FFFD73A5BF98FFEF5FEC99409" bold="true" box="[199,334,333,359]" pageId="9" pageNumber="108">Etymology.</emphasis>
|
||
Grouvelle did not cite an etymology. However, the specific epithet
|
||
<taxonomicName id="4C357DDEFFD73A5BFD0DFEF6FB399409" authorityName="Grouvelle" authorityYear="1896" box="[1093,1214,334,359]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BFD0DFEF6FB399409" box="[1093,1214,334,359]" italics="true" pageId="9" pageNumber="108">angustatus</emphasis>
|
||
</taxonomicName>
|
||
is composed of
|
||
<emphasis id="B941DA4FFFD73A5BFC3FFEF6FF5894E5" italics="true" pageId="9" pageNumber="108">angustus</emphasis>
|
||
(L.), meaning “narrow,” and
|
||
<emphasis id="B941DA4FFFD73A5BFB62FECAFDE094E5" box="[554,615,370,395]" italics="true" pageId="9" pageNumber="108">–atus</emphasis>
|
||
(L.), meaning “having the nature of.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F55D6FFD73A5BF98FFE2CFBE49061" pageId="9" pageNumber="108" type="discussion">
|
||
<paragraph id="8B8A065DFFD73A5BF98FFE2CFC179751" blockId="9.[151,1437,153,1295]" pageId="9" pageNumber="108">
|
||
<emphasis id="B941DA4FFFD73A5BF98FFE2CFEC894C1" bold="true" box="[199,335,404,431]" pageId="9" pageNumber="108">Comments.</emphasis>
|
||
<taxonomicName id="4C357DDEFFD73A5BF811FE2EFDC494C1" authorityName="Grouvelle" authorityYear="1896" box="[345,579,406,431]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BF811FE2EFDC494C1" box="[345,579,406,431]" italics="true" pageId="9" pageNumber="108">Parygrus angustatus</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFA47BACFFD73A5BFA53FE2DFC6B94DE" author="Grouvelle, A." box="[795,1004,405,432]" pageId="9" pageNumber="108" pagination="177 - 216" refId="ref21501" refString="Grouvelle, A. (1896 a) Nitidulides, Colydiides, Cucujides et Parnides recoltes par M. E. Gounelle au Bresil et autres Clavicornes nouveaux d'Amerique. Annales de la Societe Entomologique de France, 65, 177 - 216." type="journal article" year="1896">Grouvelle (1896a)</bibRefCitation>
|
||
in both Latin and French, the French description being the more detailed. There are two specimens labeled as the
|
||
<typeStatus id="548EB8FFFFD73A5BFAA8FE01FB9794BD" box="[992,1040,441,467]" pageId="9" pageNumber="108">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C357DDEFFD73A5BFD7CFE02FB4094BD" authorityName="Grouvelle" authorityYear="1896" box="[1076,1223,442,467]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BFD7CFE02FB4094BD" box="[1076,1223,442,467]" italics="true" pageId="9" pageNumber="108">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
, one in the MNHN (
|
||
<figureCitation id="130E1AD8FFD73A5BF9E8FE65FF669499" box="[160,225,477,503]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 8</figureCitation>
|
||
) and the other in the NHMUK (
|
||
<figureCitation id="130E1AD8FFD73A5BFB0DFE65FD0E9499" box="[581,649,477,503]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 9</figureCitation>
|
||
). The true
|
||
<typeStatus id="548EB8FFFFD73A5BFA49FE65FCB39499" box="[769,820,477,503]" pageId="9" pageNumber="108">type</typeStatus>
|
||
, with data closest in agreement with that in Grouvelle’s description, is the MNHN specimen. Nonetheless, it is puzzling that it bears the determination label “
|
||
<taxonomicName id="4C357DDEFFD73A5BFDBEFDB9FF499751" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">Dryops angustatus</taxonomicName>
|
||
” (
|
||
<figureCitation id="130E1AD8FFD73A5BF9A0FD9DFEAD9751" box="[232,298,549,575]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 8</figureCitation>
|
||
) because Grouvelle described the species in
|
||
<taxonomicName id="4C357DDEFFD73A5BFA69FD9EFC0E9751" authorityName=", Erichson" authorityYear="1847" box="[801,905,550,575]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941DA4FFFD73A5BFA69FD9EFC0E9751" box="[801,905,550,575]" italics="true" pageId="9" pageNumber="108">Parygrus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFD73A5BF98FFDF1FBFF979D" blockId="9.[151,1437,153,1295]" pageId="9" pageNumber="108">
|
||
It is interesting that the determination label of the NHMUK specimen (
|
||
<figureCitation id="130E1AD8FFD73A5BFA9FFDF1FB9C970D" box="[983,1051,585,611]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 9</figureCitation>
|
||
), also from Rio de Janeiero,
|
||
<collectingCountry id="F32246CDFFD73A5BFC10FDF1FA1B970D" box="[1368,1436,585,611]" name="Brazil" pageId="9" pageNumber="108">Brazil</collectingCountry>
|
||
and labeled as the type of
|
||
<taxonomicName id="4C357DDEFFD73A5BF88FFDD6FDDA97E9" box="[455,605,622,647]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BF88FFDD6FDDA97E9" box="[455,605,622,647]" italics="true" pageId="9" pageNumber="108">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
, is written in the same hand as the determination label of Grouvelle’s
|
||
<taxonomicName id="4C357DDEFFD73A5BFCC0FDD6FE9597C5" authorityName=", Grouvelle" authorityYear="1896" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="elateroides">
|
||
<emphasis id="B941DA4FFFD73A5BFCC0FDD6FE9597C5" italics="true" pageId="9" pageNumber="108">P. elateroides</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="548EB8FFFFD73A5BF851FD29FEFC97C5" box="[281,379,657,683]" pageId="9" pageNumber="108" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130E1AD8FFD73A5BF8C2FD29FE5B97C5" box="[394,476,657,683]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 10</figureCitation>
|
||
). Yet the circular format of the type label of the NHMUK specimen, from the Fry Collection, is identical to the type label of
|
||
<taxonomicName id="4C357DDEFFD73A5BFB0AFD0EFD4297A1" box="[578,709,693,719]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="erichsoni">
|
||
<emphasis id="B941DA4FFFD73A5BFB0AFD0EFD4297A1" box="[578,709,693,719]" italics="true" pageId="9" pageNumber="108">P. erichsoni</emphasis>
|
||
</taxonomicName>
|
||
described by Waterhouse 20 years earlier. These are of a different format than the type labels from the MNHN Grouvelle collection, which are rectangular.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFD73A5BF98FFD45FE579159" blockId="9.[151,1437,153,1295]" pageId="9" pageNumber="108">
|
||
The determination labels on two other images sent from the MNHN (Additional specimens, photographic images) do not match the handwriting of the label on the
|
||
<typeStatus id="548EB8FFFFD73A5BFBB0FC99FCAC9655" box="[760,811,801,827]" pageId="9" pageNumber="108">type</typeStatus>
|
||
, but one of them bears a Grouvelle Collection label. We suspect that they may not be
|
||
<taxonomicName id="4C357DDEFFD73A5BF890FCFEFDEB9631" box="[472,620,838,863]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BF890FCFEFDEB9631" box="[472,620,838,863]" italics="true" pageId="9" pageNumber="108">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
for the following reasons: In comparison with the image of the
|
||
<typeStatus id="548EB8FFFFD73A5BFC67FCFDFAE59631" box="[1327,1378,837,863]" pageId="9" pageNumber="108">type</typeStatus>
|
||
, they both have pronota with widely explanate lateral margins that are widest near the base. The image of the
|
||
<typeStatus id="548EB8FFFFD73A5BFC79FCD1FAE696ED" box="[1329,1377,873,899]" pageId="9" pageNumber="108">type</typeStatus>
|
||
does not appear to have these pronotal characters, but the angle at which it was taken may have influenced this. Nonetheless, explanate pronotal margins are not mentioned in the description, and the pronotum was stated to be widest at the first third from the base. In addition, the
|
||
<typeStatus id="548EB8FFFFD73A5BFBD8FC6DFD479681" box="[656,704,981,1007]" pageId="9" pageNumber="108">type</typeStatus>
|
||
has the elytra parallel-sided, whereas the others are sinuate with the margin explanate at the basal third. They more closely resemble the non-type images of
|
||
<taxonomicName id="4C357DDEFFD73A5BFDEDFC42FAB4917D" baseAuthorityName="Grouvelle" baseAuthorityYear="1890" box="[1189,1331,1017,1043]" class="Insecta" family="Dryopidae" genus="Parygrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B941DA4FFFD73A5BFDEDFC42FAB4917D" box="[1189,1331,1017,1043]" italics="true" pageId="9" pageNumber="108">P. parallelus</emphasis>
|
||
</taxonomicName>
|
||
than that of the
|
||
<typeStatus id="548EB8FFFFD73A5BF997FBA5FE889159" box="[223,271,1053,1079]" pageId="9" pageNumber="108">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C357DDEFFD73A5BF87CFBA6FE4E9159" box="[308,457,1054,1079]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BF87CFBA6FE4E9159" box="[308,457,1054,1079]" italics="true" pageId="9" pageNumber="108">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8A065DFFD73A5BF98FFBF9FBE49061" blockId="9.[151,1437,153,1295]" pageId="9" pageNumber="108">
|
||
Among the seven specimens examined from the NHMUK (Additional specimens examined), five were labeled as being from the Fry Collection, including the erroneously-labeled “type.” The “type” and another of these specimens have similarly handwritten determination labels, which are not in the same hand as the actual type. There is a good deal of morphological variation among the additional specimens (i.e., pronota with or without a longitudinal carina and/or explanate lateral margins) so they are likely not all
|
||
<taxonomicName id="4C357DDEFFD73A5BFA2EFB6AFC7D9185" box="[870,1018,1234,1259]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B941DA4FFFD73A5BFA2EFB6AFC7D9185" box="[870,1018,1234,1259]" italics="true" pageId="9" pageNumber="108">P. angustatus</emphasis>
|
||
</taxonomicName>
|
||
or all conspecific. However, the false type, a female (
|
||
<figureCitation id="130E1AD8FFD73A5BF80CFB4DFE0E9061" box="[324,393,1269,1295]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 9</figureCitation>
|
||
), does resemble the image of the
|
||
<typeStatus id="548EB8FFFFD73A5BFA49FB4DFCE19061" box="[769,870,1269,1295]" pageId="9" pageNumber="108" type="holotype">holotype</typeStatus>
|
||
, likely a male (
|
||
<figureCitation id="130E1AD8FFD73A5BFD59FB4DFBD29061" box="[1041,1109,1269,1295]" captionStart="FIGURES 8–11" captionStartId="10.[151,264,1948,1973]" captionTargetBox="[262,1376,247,1919]" captionTargetId="figure@10.[211,1376,237,1923]" captionTargetPageId="10" captionText="FIGURES 8–11. Parygrus species: 8) Parygrus angustatus, holotype: dorsal habitus; specimen labels; 9) Parygrus angustatus, false holotype: dorsal habitus; specimen labels; 10) Parygrus elateroides, holotype: dorsal habitus; specimen labels; 11) Parygrus parallelus, non-type: dorsal habitus; specimen labels (images 8, 10, 11 provided by the MNHN collection-Paris)." figureDoi="http://doi.org/10.5281/zenodo.3724269" httpUri="https://zenodo.org/record/3724269/files/figure.png" pageId="9" pageNumber="108">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |