324 lines
43 KiB
XML
324 lines
43 KiB
XML
<document id="2CC346F1AD62A1FA93E5E563BB75F0F8" ID-CLB-Dataset="217329" ID-DOI="10.37520/aemnp.2022.017" ID-GBIF-Dataset="795cf556-a4a9-477b-b55d-cb80ed8a0f2d" ID-ISSN="1804-6487" ID-Zenodo-Dep="7399702" ID-ZooBank="28D55112-98B1-49A5-B382-58B1B068570B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670240514228" checkinUser="felipe" docAuthor="Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas & Solodovnikov, Alexey" docDate="2022" docId="038987A0FFF54B09B384A65C1056FA0A" docLanguage="en" docName="ActaEntMusNatPra.62.1.225-299.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Quedius afrofuliginosus Gusarov 1991" docType="treatment" docVersion="5" lastPageNumber="241" masterDocId="FFB0FFD8FFFA4B19B016A005151AFF9F" masterDocTitle="Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)" masterLastPageNumber="299" masterPageNumber="225" pageNumber="240" updateTime="1699412962807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="BE23A24DCEEEAB5A6E4D695CFF003DB5">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="0D72633CBA8C5AF2D03C1627F2C2ED09">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="3C4076B159C26F9E07D9AD35F2055A61">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark & Department of Bioscience, University of Aarhus, Ny Munkegade 116, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:affiliation id="5B70843129E800A8513BBC6BD567A9DB">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Zoological Institute, Russian Academy of Science, Universitetskaja nab.</mods:affiliation>
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<taxonomicName id="4C204D35FFF54B16B384A65C105CF9EE" ID-CoL="4R2Y8" authority="Gusarov, 1991" authorityName="Gusarov" authorityYear="1991" box="[914,1350,1625,1649]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="240" phylum="Arthropoda" rank="species" species="afrofuliginosus">
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<emphasis id="B954EAA4FFF54B16B384A65C1187F9EE" bold="true" box="[914,1181,1625,1649]" italics="true" pageId="15" pageNumber="240">Quedius afrofuliginosus</emphasis>
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<bibRefCitation id="EFB14B47FFF54B16B4B2A65F105CF9EE" author="GUSAROV V. I." box="[1188,1350,1626,1649]" pageId="15" pageNumber="240" pagination="3 - 12" refId="ref83976" refString="GUSAROV V. I. 1991: Novye i maloizvestnye palearkticheskie stafilinidy (Coleoptera: Staphylinidae) [New and little-known Palearctic rove beetles (Coleoptera: Staphylinidae)] Soobshchenie 3: 3 - 12 (in Russian)." type="journal article" year="1991">Gusarov, 1991</bibRefCitation>
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<paragraph id="8B9F36B6FFF54B16B3E3A67D11F9F914" blockId="15.[828,1390,1625,1710]" box="[1013,1251,1656,1675]" pageId="15" pageNumber="240">
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(
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<figureCitation id="131B2A33FFF54B16B3EDA67D112BF914" box="[1019,1073,1656,1675]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="15" pageNumber="240">Figs 1</figureCitation>
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,
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<figureCitation id="131B2A33FFF54B16B42DA67D115DF914" box="[1083,1095,1656,1675]" captionStart="Fig" captionStartId="11.[151,182,2008,2027]" captionTargetBox="[151,1437,191,1879]" captionTargetId="figure-13@11.[151,1437,191,1968]" captionTargetPageId="11" captionText="Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80." figureDoi="http://doi.org/10.5281/zenodo.7399708" httpUri="https://zenodo.org/record/7399708/files/figure.png" pageId="15" pageNumber="240">3</figureCitation>
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,
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<figureCitation id="131B2A33FFF54B16B447A67D1171F914" box="[1105,1131,1656,1675]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="15" pageNumber="240">6E</figureCitation>
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,
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<figureCitation id="131B2A33FFF54B16B465A67D1195F914" box="[1139,1167,1656,1675]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="15" pageNumber="240">7A</figureCitation>
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,
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<figureCitation id="131B2A33FFF54B16B48EA67D11A4F914" box="[1176,1214,1656,1675]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="15" pageNumber="240">11A</figureCitation>
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,
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<figureCitation id="131B2A33FFF54B16B4D1A67D11C7F914" box="[1223,1245,1656,1675]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="15" pageNumber="240">18</figureCitation>
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)
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</paragraph>
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<treatmentCitation id="0A8110A7FFF54B16B32AA69E11BAF931" author="GUSAROV V. I." box="[828,1184,1691,1710]" page="9" pageId="15" pageNumber="240" year="1991">
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<emphasis id="B954EAA4FFF54B16B32AA69E1112F931" box="[828,1032,1691,1710]" italics="true" pageId="15" pageNumber="240">Quedius afrofuliginosus</emphasis>
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<bibRefCitation id="EFB14B47FFF54B16B418A69E11BAF931" author="GUSAROV V. I." box="[1038,1184,1691,1710]" pageId="15" pageNumber="240" pagination="3 - 12" refId="ref83976" refString="GUSAROV V. I. 1991: Novye i maloizvestnye palearkticheskie stafilinidy (Coleoptera: Staphylinidae) [New and little-known Palearctic rove beetles (Coleoptera: Staphylinidae)] Soobshchenie 3: 3 - 12 (in Russian)." type="journal article" year="1991">Gusarov, 1991: 9</bibRefCitation>
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[
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<typeStatus id="549B8814FFF54B16B4BCA69E11CDF931" box="[1194,1239,1691,1710]" pageId="15" pageNumber="240">Type</typeStatus>
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locality:
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<collectingCountry id="F3377626FFF54B16B530A69E1072F931" box="[1318,1384,1691,1710]" name="Algeria" pageId="15" pageNumber="240">Algeria</collectingCountry>
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]
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Hඈඅඈඍඒඉൾ: “
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<collectingCountry id="F3377626FFF54B16B43AA6C11175F948" box="[1068,1135,1732,1751]" name="Algeria" pageId="15" pageNumber="240">Algerie</collectingCountry>
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, Raffray / [black circle] / c.A.Yakovleva [in Russian] /
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<typeStatus id="549B8814FFF54B16B3F4A6DA1121F96D" box="[994,1083,1759,1778]" pageId="15" pageNumber="240" type="holotype">Holotypus</typeStatus>
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<taxonomicName id="4C204D35FFF54B16B429A6DA1087F96D" authority="V. Gusarov 1990" authorityName="V. Gusarov" authorityYear="1990" box="[1087,1437,1759,1778]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="240" phylum="Arthropoda" rank="species" species="afrofuliginosus">Quedius afrofuliginosus V.Gusarov 1990</taxonomicName>
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[red label] / Zoological Institute St. Petersburg INS_COL_0002505” (J
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<collectionCode id="ED31AE73FFF54B16B32AA7111679F8B8" box="[828,867,1812,1831]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="15" pageNumber="240" type="Museum">ZIN</collectionCode>
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). Pൺඋൺඍඒඉൾඌ: same as
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<typeStatus id="549B8814FFF54B16B40EA711117FF8B8" box="[1048,1125,1812,1831]" pageId="15" pageNumber="240" type="holotype">holotype</typeStatus>
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, except “
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<typeStatus id="549B8814FFF54B16B4A6A711101CF8B8" box="[1200,1286,1812,1831]" pageId="15" pageNumber="240" type="allotype">Allotypus</typeStatus>
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<taxonomicName id="4C204D35FFF54B16B51CA7111115F8DD" authority="V. Gusarov 1990" authorityName="V. Gusarov" authorityYear="1990" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="240" phylum="Arthropoda" rank="species" species="afrofuliginosus">Quedius afrofuliginosus V. Gusarov 1990</taxonomicName>
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[red label] / Zoological Institute St. Petersburg INS_COL_0002506” (
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|
||
<collectionCode id="ED31AE73FFF54B16B43EA74F1155F8C2" box="[1064,1103,1866,1885]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="15" pageNumber="240" type="Museum">ZIN</collectionCode>
|
||
); same as
|
||
<typeStatus id="549B8814FFF54B16B4BCA74F11EDF8C2" box="[1194,1271,1866,1885]" pageId="15" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
, except “
|
||
<typeStatus id="549B8814FFF54B16B550A74F1086F8C2" box="[1350,1436,1866,1885]" pageId="15" pageNumber="240" type="paratype">Paratypus</typeStatus>
|
||
<taxonomicName id="4C204D35FFF54B16B32AA7611185F8E8" authority="V. Gusarov 1990" authorityName="V. Gusarov" authorityYear="1990" box="[828,1183,1892,1911]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="240" phylum="Arthropoda" rank="species" species="afrofuliginosus">Quedius afrofuliginosus V. Gusarov 1990</taxonomicName>
|
||
[red label] / Zoological Institute St. Petersburg INS_COL_0002507-000250710” (3 JJ
|
||
<specimenCount id="9D26FD3FFFF54B16B52AA77A1046F80D" box="[1340,1372,1913,1938]" pageId="15" pageNumber="240" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFF54B16B577A77A1092F80D" box="[1377,1416,1919,1938]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="15" pageNumber="240" type="Museum">ZIN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFF54B09B32AA79F15F8FA15" blockId="15.[828,1437,1732,2071]" lastBlockId="16.[151,760,1367,1418]" lastPageId="16" lastPageNumber="241" pageId="15" pageNumber="240">
|
||
<emphasis id="B954EAA4FFF54B16B32AA79F101CF832" bold="true" box="[828,1286,1946,1965]" pageId="15" pageNumber="240">
|
||
Additional material examined.
|
||
<collectingCountry id="F3377626FFF54B16B44AA79F11A2F832" box="[1116,1208,1946,1965]" name="France" pageId="15" pageNumber="240">FRANCE</collectingCountry>
|
||
: Cඈඋඌൾ:
|
||
</emphasis>
|
||
Col de Bravella, [41.83, 9.26],
|
||
<quantity id="4CD89B53FFF54B16B3AAA7B116EAF858" box="[956,1008,1972,1991]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="15" pageNumber="240" unit="m" value="490.0">490m</quantity>
|
||
,
|
||
<date id="FF9E1076FFF54B16B3EDA7B1117DF858" box="[1019,1127,1972,1991]" pageId="15" pageNumber="240" value="1994-07-25">25.VII.1994</date>
|
||
, leg. Zerche (1 J
|
||
<collectionCode id="ED31AE73FFF54B16B507A7B1105FF858" box="[1297,1349,1972,1991]" pageId="15" pageNumber="240">SDEI</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFF54B16B540A7B1168FF87D" bold="true" pageId="15" pageNumber="240">
|
||
<collectingCountry id="F3377626FFF54B16B540A7B11082F858" box="[1366,1432,1972,1991]" name="Italy" pageId="15" pageNumber="240">ITALY</collectingCountry>
|
||
: Sൺඋൽංඇංൺ:
|
||
</emphasis>
|
||
Aritzo, Mte.Gennargentu,[39.94, 9.17], leg.C. Krüger (2JJ
|
||
<collectionCode id="ED31AE73FFF54B16B32AA7EF166CF862" box="[828,886,2026,2045]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="240">NMW</collectionCode>
|
||
); Ottana, [40.23, 9.04],
|
||
<date id="FF9E1076FFF54B16B42AA7EF118FF862" box="[1084,1173,2026,2045]" pageId="15" pageNumber="240" value="1974-10-11">11.X.1974</date>
|
||
(
|
||
<specimenCount id="9D26FD3FFFF54B16B4B7A7EF11A7F863" box="[1185,1213,2019,2045]" pageId="15" pageNumber="240" type="female">1 ♀</specimenCount>
|
||
cBor);
|
||
<emphasis id="B954EAA4FFF54B16B4EBA7EF1062F862" bold="true" box="[1277,1400,2026,2045]" pageId="15" pageNumber="240">
|
||
<collectingCountry id="F3377626FFF54B16B4EBA7EF106EF862" box="[1277,1396,2026,2045]" name="Morocco" pageId="15" pageNumber="240">MOROCCO</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Ain Leuh, [33.28, -5.33],
|
||
<date id="FF9E1076FFF54B16B3F9A8011152F788" box="[1007,1096,2052,2071]" pageId="15" pageNumber="240" value="1925-05-17">17.V.1925</date>
|
||
, leg. M. Escalera (
|
||
<specimenCount id="9D26FD3FFFF54B16B4F3A801101BF788" box="[1253,1281,2046,2071]" pageId="15" pageNumber="240" type="female">1 ♀</specimenCount>
|
||
<collectionCode id="ED31AE73FFF54B16B510A8011050F788" box="[1286,1354,2052,2071]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="15" pageNumber="240" type="Museum">MNCN</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFF54B16B541A8011086F788" bold="true" box="[1367,1436,2052,2071]" pageId="15" pageNumber="240">
|
||
<collectingCountry id="F3377626FFF54B16B541A8011083F788" box="[1367,1433,2052,2071]" name="Spain" pageId="15" pageNumber="240">SPAIN</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
Cadiz,
|
||
<collectingRegion id="49E4F854FFEA4B09B0C2A559140EFAF0" box="[212,276,1372,1391]" country="Philippines" name="Laguna" pageId="16" pageNumber="241">Laguna</collectingRegion>
|
||
de la Janda, [36.24,-5.84],
|
||
<date id="FF9E1076FFEA4B09B1EAA559174CFAF0" box="[508,598,1372,1391]" pageId="16" pageNumber="241" value="1966-05-30">30.V.1966</date>
|
||
, leg. Besuchet (1J
|
||
<collectionCode id="ED31AE73FFEA4B09B081A57215CDFA15" box="[151,215,1399,1418]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="16" pageNumber="241">NMPC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F663EFFEA4B09B081A4481488FA85" ID-DOI="http://doi.org/10.5281/zenodo.7399716" ID-Zenodo-Dep="7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="16" pageNumber="241" startId="16.[151,181,1101,1120]" targetBox="[171,736,194,1069]" targetPageId="16">
|
||
<paragraph id="8B9F36B6FFEA4B09B081A4481488FA85" blockId="16.[151,761,1101,1306]" pageId="16" pageNumber="241">
|
||
Fig.6. Head and pronotal chaetotaxy of
|
||
<taxonomicName id="4C204D35FFEA4B09B1CBA4481739FBFF" authorityName="Stephens" authorityYear="1829" box="[477,547,1101,1120]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B954EAA4FFEA4B09B1CBA4481739FBFF" box="[477,547,1101,1120]" italics="true" pageId="16" pageNumber="241">Quedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26757DFFFEA4B09B230A448174BFBFF" box="[550,593,1101,1120]" pageId="16" pageNumber="241" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
A – general scheme; B –
|
||
<taxonomicName id="4C204D35FFEA4B09B0A8A46214CBFBE5" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[190,465,1127,1146]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="curtipennis">
|
||
<emphasis id="B954EAA4FFEA4B09B0A8A4621423FBE5" box="[190,313,1127,1146]" italics="true" pageId="16" pageNumber="241">Q. curtipennis</emphasis>
|
||
<bibRefCitation id="EFB14B47FFEA4B09B156A46214CBFBE5" author="BERNHAUER M." box="[320,465,1127,1146]" pageId="16" pageNumber="241" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
, C –
|
||
<taxonomicName id="4C204D35FFEA4B09B214A46215D1FB0A" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="fuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B214A4621761FBE5" box="[514,635,1127,1146]" italics="true" pageId="16" pageNumber="241">Q. fuliginosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB14B47FFEA4B09B29EA46215DFFB0A" author="GRAVENHORST J. L. C." pageId="16" pageNumber="241" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; D –
|
||
<taxonomicName id="4C204D35FFEA4B09B0EBA48714C0FB0A" authority="Brulle, 1832" authorityName="Brulle" authorityYear="1832" box="[253,474,1154,1173]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="levicollis">
|
||
<emphasis id="B954EAA4FFEA4B09B0EBA487147FFB0A" box="[253,357,1154,1173]" italics="true" pageId="16" pageNumber="241">Q. levicollis</emphasis>
|
||
<bibRefCitation id="EFB14B47FFEA4B09B17AA48714C0FB0A" author="BRULLE G. A." box="[364,474,1154,1173]" pageId="16" pageNumber="241" pagination="289 - 400" refId="ref80797" refString="BRULLE G. A. 1832 - 1833: Expedition scientifique de Moree. Section des Sciences Physiques. Tome III. 1. re Partie. Zoologie. Deuxieme Section. Des animales articules. F. G. Levrault, Paris, 400 pp. [Pp. 1 - 288 published in 1832, pp. 289 - 400 in 1833.]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
|
||
</taxonomicName>
|
||
; E –
|
||
<taxonomicName id="4C204D35FFEA4B09B21FA48715DEFB2F" authority="Gusarov, 1991" authorityName="Gusarov" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="afrofuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B21FA48717BEFB0A" box="[521,676,1154,1173]" italics="true" pageId="16" pageNumber="241">Q. afrofuliginosus</emphasis>
|
||
<bibRefCitation id="EFB14B47FFEA4B09B2BDA48715DEFB2F" author="GUSAROV V. I." pageId="16" pageNumber="241" pagination="3 - 12" refId="ref83976" refString="GUSAROV V. I. 1991: Novye i maloizvestnye palearkticheskie stafilinidy (Coleoptera: Staphylinidae) [New and little-known Palearctic rove beetles (Coleoptera: Staphylinidae)] Soobshchenie 3: 3 - 12 (in Russian)." type="journal article" year="1991">Gusarov, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
; F –
|
||
<taxonomicName id="4C204D35FFEA4B09B0E7A4981474FB2F" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[241,366,1181,1200]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="molochinus">
|
||
<emphasis id="B954EAA4FFEA4B09B0E7A4981474FB2F" box="[241,366,1181,1200]" italics="true" pageId="16" pageNumber="241">Q. molochinus</emphasis>
|
||
</taxonomicName>
|
||
- and
|
||
<taxonomicName id="4C204D35FFEA4B09B1B7A4981712FB2F" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[417,520,1181,1200]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFEA4B09B1B7A4981712FB2F" box="[417,520,1181,1200]" italics="true" pageId="16" pageNumber="241">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
-groups; G –
|
||
<taxonomicName id="4C204D35FFEA4B09B26DA49815A3FB55" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="molochinoides">
|
||
<emphasis id="B954EAA4FFEA4B09B26DA49815A3FB55" italics="true" pageId="16" pageNumber="241">Q. molochinoides</emphasis>
|
||
</taxonomicName>
|
||
-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A653DFFEA4B09B081A5A71681FC8A" pageId="16" pageNumber="241" type="description">
|
||
<paragraph id="8B9F36B6FFEA4B09B081A5A715FAF8A6" blockId="16.[151,760,1437,2073]" pageId="16" pageNumber="241">
|
||
<emphasis id="B954EAA4FFEA4B09B081A5A7145AFA26" bold="true" box="[151,320,1442,1465]" pageId="16" pageNumber="241">Redescription.</emphasis>
|
||
Measurements JJ (n = 6): HW = 1.78– 1.89 (1.84); HL = 1.54–1.58 (1.56); HL/HW 0.82–0.89 (0.85); PW = 2.24–2.44 (2.36); PL = 2.11–2.22 (2.16); PL/PW 0.89–0.96 (0.92); EW = 2.33–2.68 (2.59); EL = 2.40–2.51 (2.47); EL/EW 0.90–1.03 (0.95); EL/PL 1.10–1.19 (1.14); PW/HW 1.44–1.57 (1.51); forebody length 6.10–6.29 (6.19).
|
||
<specimenCount id="9D26FD3FFFEA4B09B1BBA65E14C9F9E5" box="[429,467,1627,1658]" pageId="16" pageNumber="241" type="female">♀♀</specimenCount>
|
||
(n = 3): HW = 1.84–1.89 (1.87); HL = 1.56–1.62 (1.59); HL/HW 0.84–0.86 (0.85); PW = 2.38–2.40 (2.39); PL = 2.04–2.24 (2.17); PL/PW 0.86–0.94 (0.91); EW = 2.69–2.73 (2.72); EL = 2.36–2.58 (2.48); EL/EW 0.88–0.94 (0.91); EL/PL 1.06–1.26 (1.15); PW/HW 1.47–1.53 (1.50); forebody length 6.18–6.36 (6.24).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B0A3A7471726F8C6" blockId="16.[151,760,1437,2073]" box="[181,572,1858,1881]" pageId="16" pageNumber="241">
|
||
Large species; body black (
|
||
<figureCitation id="131B2A33FFEA4B09B1CFA7471735F8C6" box="[473,559,1858,1881]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="16" pageNumber="241">Fig. 7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B0A3A7671797F786" blockId="16.[151,760,1437,2073]" pageId="16" pageNumber="241">
|
||
Head black, distinctly transverse, with microsculpture of head transverse waves, four punctures between anterior frontal punctures, one additional unique puncture on either side posterior of these (
|
||
<figureCitation id="131B2A33FFEA4B09B199A7C714FBF846" box="[399,481,1986,2009]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="16" pageNumber="241">Fig. 6E</figureCitation>
|
||
), eyes very large (EyL/TL = 3.17–3.91 (3.68)) and clearly protruding; antennae and palpi pale reddish, all anntennomeres elongate.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B34CA0BB16B6FE4A" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row, two punctures in sublateral row with posteriormost puncture reaching level of middle puncture of dorsal row; scutellum smooth and glabrous; elytra black, uniformly pubescent, quadrate or slightly wider than long, slightly longer than pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B34CA1DB16B1FD8A" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">Abdomen black, tergites uniformly punctured, slightly iridescent.</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B34CA2181681FC8A" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">
|
||
<emphasis id="B954EAA4FFEA4B09B34CA2181680FDAA" bold="true" box="[858,922,541,565]" italics="true" pageId="16" pageNumber="241">Male.</emphasis>
|
||
Aedeagus (
|
||
<figureCitation id="131B2A33FFEA4B09B436A21B1192FDAA" box="[1056,1160,542,565]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="16" pageNumber="241">Fig. 11A</figureCitation>
|
||
): paramere broad, rather parallel-sided, reaching apex of median lobe, with sensory peg setae forming two single rows along parameral lateral margins for 2/3 of their length; median lobe on parameral side with clear apical protrusion and two subapical small teeth, the latter positioned near end of peg setae rows of paramere; internal sac with four distal elongate medial sclerites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFEA4B09B32AA31B1686FB2A" pageId="16" pageNumber="241" type="diagnosis">
|
||
<paragraph id="8B9F36B6FFEA4B09B32AA31B11ADFBAA" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">
|
||
<emphasis id="B954EAA4FFEA4B09B32AA31B1120FCAA" bold="true" box="[828,1082,798,821]" pageId="16" pageNumber="241">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C204D35FFEA4B09B455A31B1053FCAA" authorityName="Gusarov" authorityYear="1991" box="[1091,1353,798,821]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="afrofuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B455A31B1053FCAA" box="[1091,1353,798,821]" italics="true" pageId="16" pageNumber="241">Quedius afrofuliginosus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="4C204D35FFEA4B09B47DA33B11DEFCCA" authorityName="Stephens" authorityYear="1829" box="[1131,1220,830,853]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B954EAA4FFEA4B09B47DA33B11DEFCCA" box="[1131,1220,830,853]" italics="true" pageId="16" pageNumber="241">Quedius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26757DFFFEA4B09B4DDA33B101EFCCA" box="[1227,1284,830,853]" pageId="16" pageNumber="241" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
by the unique head chaetotaxy, with a puncture on either side posterior of the anterior frontal puncture (
|
||
<figureCitation id="131B2A33FFEA4B09B48AA37B11EAFC0A" box="[1180,1264,894,917]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="16" pageNumber="241">Fig. 6E</figureCitation>
|
||
). Within the
|
||
<taxonomicName id="4C204D35FFEA4B09B569A37B1686FC2A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="fuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B569A37B1686FC2A" italics="true" pageId="16" pageNumber="241">fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
-group it is additionally distinguished from
|
||
<taxonomicName id="4C204D35FFEA4B09B594A39B16ABFC4A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="fuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B594A39B16ABFC4A" italics="true" pageId="16" pageNumber="241">Q. fuliginosus</emphasis>
|
||
</taxonomicName>
|
||
by the pale antennae, from
|
||
<taxonomicName id="4C204D35FFEA4B09B4C3A3BB104FFC4A" authorityName="Brulle" authorityYear="1832" box="[1237,1365,958,981]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="levicollis">
|
||
<emphasis id="B954EAA4FFEA4B09B4C3A3BB104FFC4A" box="[1237,1365,958,981]" italics="true" pageId="16" pageNumber="241">Q. levicollis</emphasis>
|
||
</taxonomicName>
|
||
by the impunctate and glabrous scutellum, and from
|
||
<taxonomicName id="4C204D35FFEA4B09B50AA3DB1641FB8A" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="curtipennis">
|
||
<emphasis id="B954EAA4FFEA4B09B50AA3DB1641FB8A" italics="true" pageId="16" pageNumber="241">Q. curtipennis</emphasis>
|
||
</taxonomicName>
|
||
by wider head (HW/HL = 0.74 vs 0.85) and aedeagal median lobe with apical protrusion.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFEA4B09B32AA43B1686FB2A" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">
|
||
<emphasis id="B954EAA4FFEA4B09B32AA43B16A0FBCA" bold="true" box="[828,954,1086,1109]" pageId="16" pageNumber="241">Bionomics.</emphasis>
|
||
Nothing is known about the biology of
|
||
<taxonomicName id="4C204D35FFEA4B09B594A43B16C7FBEA" authorityName="Gusarov" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="afrofuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B594A43B16C7FBEA" italics="true" pageId="16" pageNumber="241">Q. afrofuliginosus</emphasis>
|
||
</taxonomicName>
|
||
. Based on collecting sites it is most likely occurring in forested areas of medium elevation (
|
||
<quantity id="4CD89B53FFEA4B09B543A47B168AFB2A" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="14.0" metricValueMin="4.5" pageId="16" pageNumber="241" unit="m" value="925.0" valueMax="1400.0" valueMin="450.0">450 to 1400 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFEA4B09B32AA4BB1056FA0A" pageId="16" pageNumber="241" type="distribution">
|
||
<paragraph id="8B9F36B6FFEA4B09B32AA4BB1056FA0A" blockId="16.[828,1437,190,1429]" pageId="16" pageNumber="241">
|
||
<emphasis id="B954EAA4FFEA4B09B32AA4BB16CFFB4A" bold="true" box="[828,981,1214,1237]" pageId="16" pageNumber="241">Distribution.</emphasis>
|
||
<taxonomicName id="4C204D35FFEA4B09B3C9A4BB11F5FB4A" authorityName="Gusarov" authorityYear="1991" box="[991,1263,1214,1237]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="afrofuliginosus">
|
||
<emphasis id="B954EAA4FFEA4B09B3C9A4BB11F5FB4A" box="[991,1263,1214,1237]" italics="true" pageId="16" pageNumber="241">Quedius afrofuliginosus</emphasis>
|
||
</taxonomicName>
|
||
was originally described from an unknown locality in
|
||
<collectingCountry id="F3377626FFEA4B09B4F4A4DB102FFB6A" box="[1250,1333,1246,1269]" name="Algeria" pageId="16" pageNumber="241">Algeria</collectingCountry>
|
||
. The new material studied in this paper brought to light additional specimens from
|
||
<collectingCountry id="F3377626FFEA4B09B3FCA51B1123FAAA" box="[1002,1081,1310,1333]" name="Tunisia" pageId="16" pageNumber="241">Tunisia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3377626FFEA4B09B47AA51B11D6FAAA" box="[1132,1228,1310,1333]" name="Morocco" pageId="16" pageNumber="241">Morocco</collectingCountry>
|
||
in Northern Africa, as well as the first records of this species from the Mediterranean region in southern
|
||
<collectingCountry id="F3377626FFEA4B09B46DA55B11A2FAEA" box="[1147,1208,1374,1397]" name="Spain" pageId="16" pageNumber="241">Spain</collectingCountry>
|
||
and on the southern European islands of
|
||
<collectingCountry id="F3377626FFEA4B09B400A57B114EFA0A" box="[1046,1108,1406,1429]" name="France" pageId="16" pageNumber="241">Corse</collectingCountry>
|
||
and Sardinia (
|
||
<figureCitation id="131B2A33FFEA4B09B4E6A57B1025FA0A" box="[1264,1343,1406,1429]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="16" pageNumber="241">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |