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<document id="EDD36E7AC337954F1CB387A213064211" ID-DOI="10.5281/zenodo.282079" ID-GBIF-Dataset="13186f0a-7e29-45d1-9004-d7fd4851960b" ID-ISSN="1175-5326" ID-Zenodo-Dep="282079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460512734246" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2012" docId="03898793FFC4E767FF4E45DEFA8D7C2B" docLanguage="en" docName="zt03435p039.pdf" docOrigin="Zootaxa 3435" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Retrozetes" docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="FFB0FFEBFFD6E775FFD94549FFF87913" masterDocTitle="New eremaeozetid mites (Acari: Oribatida: Eremaeozetoidea) from the south-western Pacific region and the taxonomic status of the Eremaeozetidae and Idiozetidae" masterLastPageNumber="39" masterPageNumber="1" pageNumber="19" updateTime="1698313531070" updateUser="plazi">
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<mods:title id="1152721A6BFEA4B287C4710422B18E67">New eremaeozetid mites (Acari: Oribatida: Eremaeozetoidea) from the south-western Pacific region and the taxonomic status of the Eremaeozetidae and Idiozetidae</mods:title>
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<mods:namePart id="55C52573F9667705FAFB6F6F16984804">Colloff, Matthew J.</mods:namePart>
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<mods:title id="DF7508D27E6CC61CD941F48EE470F1B5">Zootaxa</mods:title>
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<mods:date id="614A6BD47A23464320CF26694E1FA9BC">2012</mods:date>
<mods:detail id="A61D0CCBAFF161F7A575CA420D308078" type="volume">
<mods:number id="4DFA89AC63E2DE9AC54807BCA5369F14">3435</mods:number>
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<mods:classification id="603E3315C9B0E44CD7DF2AD8C7AD3F04">journal article</mods:classification>
<mods:identifier id="818541246E20C1F61E263A03B6148B04" type="DOI">10.5281/zenodo.282079</mods:identifier>
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<mods:identifier id="DD201354F1FD2F10383946F4C427A4B6" type="Zenodo-Dep">282079</mods:identifier>
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<treatment id="03898793FFC4E767FF4E45DEFA8D7C2B" ID-DOI="http://doi.org/10.5281/zenodo.6179442" ID-GBIF-Taxon="119523530" ID-Zenodo-Dep="6179442" LSID="urn:lsid:plazi:treatment:03898793FFC4E767FF4E45DEFA8D7C2B" httpUri="http://treatment.plazi.org/id/03898793FFC4E767FF4E45DEFA8D7C2B" lastPageNumber="19" pageId="18" pageNumber="19">
<subSubSection id="C33A650EFFC4E767FF4E45DEFE7879A2" box="[151,384,151,177]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B9F3685FFC4E767FF4E45DEFE7879A2" blockId="18.[151,384,151,177]" box="[151,384,151,177]" pageId="18" pageNumber="19">
<heading id="D0D781E9FFC4E767FF4E45DEFE7879A2" bold="true" box="[151,384,151,177]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B954EA97FFC4E767FF4E45DEFE7879A2" bold="true" box="[151,384,151,177]" pageId="18" pageNumber="19">
<taxonomicName id="4C204D06FFC4E767FF4E45DEFEEB79A2" ID-CoL="77XL" box="[151,275,151,177]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EA97FFC4E767FF4E45DEFEEB79A2" bold="true" box="[151,275,151,177]" italics="true" pageId="18" pageNumber="19">Retrozetes</emphasis>
</taxonomicName>
gen nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A650EFFC4E767FF4E4597FD057BC0" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8B9F3685FFC4E767FF4E4597FD057BC0" blockId="18.[151,1437,222,1338]" pageId="18" pageNumber="19">
<emphasis id="B954EA97FFC4E767FF4E4597FEE979E4" bold="true" box="[151,273,222,247]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Cerotegument thick, reticulate-alveolate. Lamellae extremely long, elevated above prodorsum; separated and recurved apically, extending well beyond rostrum and prominent projecting tutorium; with narrow slit or broad separation anteriorly. Lamellar setae on medial margins of lamellar cusps, sub-apical. Lamellar cuspides folded ventrally in the transverse plane. Interlamellar setae present. Bothridial seta with long stalk, broadening and barbed apically. Bothridium small, cup-shaped. Dorsosejugal suture transverse; bounded laterally by conical apophyses of setae
<emphasis id="B954EA97FFC4E767FE3E44DDFE0B78B8" box="[487,499,404,427]" italics="true" pageId="18" pageNumber="19">c</emphasis>
2 positioned on strongly reduced dorsosejugal lobes. Lenticulus strongly protuberant on conical projection, positioned close to dorsosejugal suture. Notogaster in dorsal view broader than long; in lateral view wedge-shaped with posterior margin elevated; centrodorsal region convex, elevated. Notogaster with ten pairs of setae,
<emphasis id="B954EA97FFC4E767FEBF4748FE7A7B0A" box="[358,386,513,537]" italics="true" pageId="18" pageNumber="19">lm</emphasis>
in centrodorsal position; at least one pair of sacculi. Mentum elongated, anterior margin not reflexed. Epimeral setal formulae 2-1-2-2 or 2-1-2-1. Pre-genital sclerite absent. Genital and anal plates separated by distance of about half length of genital setae. Aggenital setae present. Six pairs of genital setae, all in alignment,
<emphasis id="B954EA97FFC4E767FF4E4727FF5D7B96" box="[151,165,622,645]" italics="true" pageId="18" pageNumber="19">g</emphasis>
1 not displaced laterally of others. Anal setae two pairs, with or without thick, bacilliform cerotegument. Setae
<emphasis id="B954EA97FFC4E767FAAA4724FA777B96" box="[1395,1423,621,645]" italics="true" pageId="18" pageNumber="19">ad</emphasis>
2- 3 positioned close together near lateral margin of ventral plate;
<emphasis id="B954EA97FFC4E767FCA547DDFC607BBF" box="[892,920,660,684]" italics="true" pageId="18" pageNumber="19">ad</emphasis>
<quantity id="4CD89B60FFC4E767FC4147EAFC397BBF" box="[920,961,660,689]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="18" pageNumber="19" unit="in" value="1.0">1 in</quantity>
post-anal position. Legs monodactylous. Solenidion σ on genu I greatly elongated, flagelliform.
</paragraph>
</subSubSection>
<subSubSection id="C33A650EFFC4E767FF104796FD3F7BE4" box="[201,711,734,759]" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8B9F3685FFC4E767FF104796FD3F7BE4" blockId="18.[151,1437,222,1338]" box="[201,711,734,759]" pageId="18" pageNumber="19">
<emphasis id="B954EA97FFC4E767FF104796FE9C7BE4" bold="true" box="[201,356,734,759]" pageId="18" pageNumber="19">
<typeStatus id="549B8827FFC4E767FF104796FEFC7BE4" box="[201,260,735,759]" pageId="18" pageNumber="19">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C204D06FFC4E767FEB247A9FD9F7BE4" box="[363,615,735,759]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="koghisensis" status="sp. nov.">
<emphasis id="B954EA97FFC4E767FEB247A9FD9F7BE4" box="[363,615,735,759]" italics="true" pageId="18" pageNumber="19">Retrozetes koghisensis</emphasis>
</taxonomicName>
<emphasis id="B954EA97FFC4E767FDB44796FD3F7BE4" bold="true" box="[621,711,735,759]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A26757ECFFC4E767FDB44796FD3F7BE4" box="[621,711,735,759]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33A650EFFC4E767FF10464AFE727ADC" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8B9F3685FFC4E767FF10464AFE727ADC" blockId="18.[151,1437,222,1338]" pageId="18" pageNumber="19">
<emphasis id="B954EA97FFC4E767FF10464AFEC57A08" bold="true" box="[201,317,771,795]" pageId="18" pageNumber="19">Remarks.</emphasis>
<taxonomicName id="4C204D06FFC4E767FE9F464DFE427A08" box="[326,442,772,795]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EA97FFC4E767FE9F464DFE427A08" box="[326,442,772,795]" italics="true" pageId="18" pageNumber="19">Retrozetes</emphasis>
</taxonomicName>
shares character states with
<taxonomicName id="4C204D06FFC4E767FCD1464AFC947A08" box="[776,876,771,795]" class="Arachnida" family="Idiozetidae" genus="Idiozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA97FFC4E767FCD1464AFC947A08" box="[776,876,771,795]" italics="true" pageId="18" pageNumber="19">Idiozetes</emphasis>
</taxonomicName>
including the presence of sacculi; the very long lamellar cuspides; the conical, protuberant lenticulus; a strongly projecting tutorial apex, the enclosed bothridia with small openings and a rim without an incision and the long, typically thin, blade-like pteromorphs.
<taxonomicName id="4C204D06FFC4E767FAF14605FA647A70" box="[1320,1436,844,867]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EA97FFC4E767FAF14605FA647A70" box="[1320,1436,844,867]" italics="true" pageId="18" pageNumber="19">Retrozetes</emphasis>
</taxonomicName>
also shares character states with
<emphasis id="B954EA97FFC4E767FDDA4639FD867A94" box="[515,638,880,903]" italics="true" pageId="18" pageNumber="19">Rogerzetes</emphasis>
, particularly the sub-apical lamellar setae; the lamellar cuspides folded ventrally in the transverse plane; the transverse dorsosejugal suture and with setae
<emphasis id="B954EA97FFC4E767FBA746DAFB627AB8" box="[1150,1178,915,939]" italics="true" pageId="18" pageNumber="19">lm</emphasis>
close together in the centrodorsal position.
</paragraph>
</subSubSection>
<subSubSection id="C33A650EFFC4E767FF104695FA8D7C2B" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8B9F3685FFC4E767FF104695FA8D7C2B" blockId="18.[151,1437,222,1338]" pageId="18" pageNumber="19">
<taxonomicName id="4C204D06FFC4E767FF104695FD597AE0" ID-CoL="8SXR4" authority="Mahunka 1999" authorityName="Mahunka" authorityYear="1999" box="[201,673,987,1011]" class="Arachnida" family="Eremaeozetidae" genus="Eremaeozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hanswursti">
<emphasis id="B954EA97FFC4E767FF104695FE117AE0" box="[201,489,987,1011]" italics="true" pageId="18" pageNumber="19">Eremaeozetes hanswursti</emphasis>
<bibRefCitation id="EFB14B74FFC4E767FE2D4692FD597AE0" author="Mahunka" box="[500,673,987,1011]" pageId="18" pageNumber="19" refString="Mahunka, S. (1999) Oribatids from Singapore III (Acari: Oribatida) (Acarologica Genavensia XCII). Archives des Sciences, Geneve, 52, 1 - 8." type="journal article" year="1999">Mahunka 1999</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3377615FFC4E767FD374692FCA77AE0" box="[750,863,987,1011]" name="Singapore" pageId="18" pageNumber="19">Singapore</collectingCountry>
is a
<taxonomicName id="4C204D06FFC4E767FC464695FBEB7AE0" ID-CoL="77XL" box="[927,1043,988,1011]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EA97FFC4E767FC464695FBEB7AE0" box="[927,1043,988,1011]" italics="true" pageId="18" pageNumber="19">Retrozetes</emphasis>
</taxonomicName>
based on the following character states and is hereby recombined: 1) in lateral view the notogaster is wedge-shaped and deepest posteriorly and the circumgastric scissure is clearly visible; 2) the centrodosal region with an oval area; 3) the humeral region and cerotegument of humeral regions strongly triangular (cf.
<figureCitation id="131B2A00FFC4E767FCF2410EFC867D4C" box="[811,894,1095,1119]" captionStart="FIGURE 12" captionStartId="20.[151,250,1234,1257]" captionTargetBox="[253,1327,220,1201]" captionTargetId="figure@20.[244,1345,193,1216]" captionTargetPageId="20" captionText="FIGURE 12. Retrozetes koghisensis sp. nov., a) left palp, antiaxial view; b) anal plates with cerotegument removed showing anal setae; c) ovipositor." httpUri="https://zenodo.org/record/282091/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
of
<bibRefCitation id="EFB14B74FFC4E767FC73410EFBA57D4C" author="Mahunka" box="[938,1117,1095,1119]" pageId="18" pageNumber="19" refString="Mahunka, S. (1999) Oribatids from Singapore III (Acari: Oribatida) (Acarologica Genavensia XCII). Archives des Sciences, Geneve, 52, 1 - 8." type="journal article" year="1999">Mahunka, 1999</bibRefCitation>
and
<figureCitation id="131B2A00FFC4E767FB43410EFB147D4C" box="[1178,1260,1095,1119]" captionStart="FIGURE 15" captionStartId="24.[151,250,1873,1896]" captionTargetBox="[168,1422,752,1829]" captionTargetId="figure@24.[151,1436,735,1851]" captionTargetPageId="24" captionText="FIGURE 15. Retrozetes novaecaledoniae sp. nov., holotype female; a) dorsal, with cerotegument; b) dorsal, with cerotegument removed." httpUri="https://zenodo.org/record/282094/files/figure.png" pageId="18" pageNumber="19">Fig. 15</figureCitation>
herein); 3) the marginal arrangement of the notogastral setae except the centrodorsal
<emphasis id="B954EA97FFC4E767FC6F4122FC2A7D90" box="[950,978,1131,1155]" italics="true" pageId="18" pageNumber="19">lm</emphasis>
and the presence of a saccule medial to seta
<emphasis id="B954EA97FFC4E767FF1441C6FF1B7DB4" box="[205,227,1167,1191]" italics="true" pageId="18" pageNumber="19">lp</emphasis>
; 4) the genital setae are aligned longitudinally:
<emphasis id="B954EA97FFC4E767FCC541D9FCD27DB4" box="[796,810,1168,1191]" italics="true" pageId="18" pageNumber="19">g</emphasis>
1 is not displaced laterally as in
<taxonomicName id="4C204D06FFC4E767FB7341D9FABD7DB4" ID-CoL="4DPZ" box="[1194,1349,1168,1191]" class="Arachnida" family="Eremaeozetidae" genus="Eremaeozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA97FFC4E767FB7341D9FABD7DB4" box="[1194,1349,1168,1191]" italics="true" pageId="18" pageNumber="19">Eremaeozetes</emphasis>
</taxonomicName>
; 5) the configuration of the lamellae with a central, expanded notch and medially-opposed lamellar setae in dorsal view and reflexed (though covered in cerotegument) in lateral view; 6) the large anal setae each at the far ends of the plate, curved and covered in thick cerotegument; 7) its small size, 152-158 μm long, fits closer with the length of
<taxonomicName id="4C204D06FFC4E767FF4E406AFEF37C29" ID-CoL="77XL" box="[151,267,1315,1338]" class="Arachnida" family="Eremaeozetidae" genus="Retrozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EA97FFC4E767FF4E406AFEF37C29" box="[151,267,1315,1338]" italics="true" pageId="18" pageNumber="19">Retrozetes</emphasis>
</taxonomicName>
spp. (mean 308 μm, range 252–356 μm) than
<taxonomicName id="4C204D06FFC4E767FCC8406AFC537C29" ID-CoL="4DPZ" box="[785,939,1315,1338]" class="Arachnida" family="Eremaeozetidae" genus="Eremaeozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA97FFC4E767FCC8406AFC537C29" box="[785,939,1315,1338]" italics="true" pageId="18" pageNumber="19">Eremaeozetes</emphasis>
</taxonomicName>
spp. (mean 426 μm range 288–614 μm).
</paragraph>
</subSubSection>
</treatment>
</document>
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