135 lines
14 KiB
XML
135 lines
14 KiB
XML
<document id="80CFDCE46DCBA1CF5D91D75CF243EDBF" ID-DOI="10.11646/zootaxa.4088.4.7" ID-GBIF-Dataset="32a23ac3-40ec-4578-868d-8716d5bb9bfe" ID-ISSN="1175-5326" ID-Zenodo-Dep="263904" ID-ZooBank="59D52D3C-3FD7-42A0-BDE9-8FC58DA38787" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457683834578" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="03898790666BFFAAAFF0F8A4B824F880" docLanguage="en" docName="15607-5127-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocephennium araguanum Jaloszynski" docType="treatment" docVersion="10" lastPageNumber="566" masterDocId="FFB0FFE8666AFFA9AF67FFF5B921FFA1" masterDocTitle="A new species and new records of Pseudocephennium Reitter in Venezuela, with notes on movable endophallic structures (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="570" masterPageNumber="563" pageNumber="564" updateTime="1698177983155" updateUser="plazi">
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<mods:title id="9EBEAB1F798B9C9E1AD7141712F4BEB7">A new species and new records of Pseudocephennium Reitter in Venezuela, with notes on movable endophallic structures (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="6AFA7CC3C187F57303B906C6D42CECF8">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="A9B744FF6B517CD504A98E774B62F6F5">Zootaxa</mods:title>
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<mods:part id="BDF39B2071F63079604C897C04A11290">
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<mods:date id="D7D433EE46BFC3E3270899F01279B6AC">2016</mods:date>
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<mods:number id="EFA5D4D5FB49D1D5460271A6F564EC3A">4088</mods:number>
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<mods:detail id="3CCE572BE68A6C791E67CCBFA151BECA" type="issue">
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<mods:number id="D16C27BAAAD715B4BF203985FAF14776">4</mods:number>
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<mods:classification id="4C3A3F8A0A2802152DD21DFCECE21487">journal article</mods:classification>
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<mods:identifier id="DF222D1125664A6A58E3D241C25D6256" type="DOI">10.11646/zootaxa.4088.4.7</mods:identifier>
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<mods:identifier id="980240C68C7FD39F32728CE09286269A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A811D0755106EBFC8A36267F71BAC025" type="Zenodo-Dep">263904</mods:identifier>
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<treatment id="03898790666BFFAAAFF0F8A4B824F880" ID-DOI="http://doi.org/10.5281/zenodo.6068504" ID-GBIF-Taxon="125783124" ID-Zenodo-Dep="6068504" LSID="urn:lsid:plazi:treatment:03898790666BFFAAAFF0F8A4B824F880" httpUri="http://treatment.plazi.org/id/03898790666BFFAAAFF0F8A4B824F880" lastPageId="3" lastPageNumber="566" pageId="1" pageNumber="564">
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<subSubSection id="C33A650D666BFFA8AFF0F8A4BABAF873" pageId="1" pageNumber="564" type="nomenclature">
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<paragraph id="8B9F3686666BFFA8AFF0F8A4BB91F8CA" blockId="1.[151,688,1873,1933]" box="[151,688,1873,1899]" pageId="1" pageNumber="564">
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<heading id="D0D781EA666BFFA8AFF0F8A4BB91F8CA" bold="true" box="[151,688,1873,1899]" fontSize="11" level="1" pageId="1" pageNumber="564" reason="1">
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<taxonomicName id="4C204D05666BFFA8AFF0F8A4BB91F8CA" ID-CoL="4NR34" authority="Jaloszynski" authorityName="Jaloszynski" box="[151,688,1873,1899]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="araguanum">
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<emphasis id="B954EA94666BFFA8AFF0F8A4BB91F8CA" bold="true" box="[151,688,1873,1899]" pageId="1" pageNumber="564">
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<emphasis id="B954EA94666BFFA8AFF0F8A4BB3BF8CA" bold="true" box="[151,538,1873,1899]" italics="true" pageId="1" pageNumber="564">Pseudocephennium araguanum</emphasis>
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Jałoszyński
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9F3686666BFFA8AFF0F881B8A9F82C" blockId="1.[151,688,1873,1933]" box="[151,392,1908,1933]" pageId="1" pageNumber="564">
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(
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<figureCitation id="131B2A03666BFFA8AFF8F881B9C6F82D" box="[159,231,1908,1933]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1403,1424]" captionTargetBox="[176,1413,220,1366]" captionTargetId="figure@2.[151,1436,195,1381]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Dorsal habitus of Pseudocephennium maximum (1) and P. araguanum (2)." httpUri="https://zenodo.org/record/263905/files/figure.png" pageId="1" pageNumber="564">Figs 2</figureCitation>
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, 7–12, 19–22)
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</paragraph>
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<paragraph id="8B9F3686666BFFA8AFF0F849BABAF873" blockId="1.[151,923,1980,2002]" box="[151,923,1980,2002]" pageId="1" pageNumber="564">
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<taxonomicName id="4C204D05666BFFA8AFF0F849BBE7F873" authority="Jaloszynski, 2012: 466" authorityName="Jaloszynski" authorityPageNumber="466" authorityYear="2012" box="[151,710,1980,2002]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="araguanum">
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<emphasis id="B954EA94666BFFA8AFF0F849B8F6F873" box="[151,471,1980,2002]" italics="true" pageId="1" pageNumber="564">Pseudocephennium araguanum</emphasis>
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Jałoszyński, 2012: 466
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</taxonomicName>
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, figs 10A–D, 11H–I.
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</paragraph>
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</subSubSection>
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<caption id="DF5F660E6668FFABAFF0FA8EBD0EFA30" box="[151,1071,1403,1425]" httpUri="https://zenodo.org/record/263905/files/figure.png" pageId="2" pageNumber="565" targetBox="[176,1413,220,1366]" targetPageId="2">
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<paragraph id="8B9F36866668FFABAFF0FA8EBD0EFA30" blockId="2.[151,1071,1403,1425]" box="[151,1071,1403,1425]" pageId="2" pageNumber="565">
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<emphasis id="B954EA946668FFABAFF0FA8EB81BFA31" bold="true" box="[151,314,1403,1424]" pageId="2" pageNumber="565">FIGURES 1–2.</emphasis>
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Dorsal habitus of
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<taxonomicName id="4C204D056668FFABAE92FA8EBA02FA30" box="[501,803,1403,1425]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="maximum">
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<emphasis id="B954EA946668FFABAE92FA8EBA02FA30" box="[501,803,1403,1425]" italics="true" pageId="2" pageNumber="565">Pseudocephennium maximum</emphasis>
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</taxonomicName>
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(1) and
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<taxonomicName id="4C204D056668FFABAC10FA89BD24FA30" box="[887,1029,1404,1425]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="araguanum">
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<emphasis id="B954EA946668FFABAC10FA89BD24FA30" box="[887,1029,1404,1425]" italics="true" pageId="2" pageNumber="565">P. araguanum</emphasis>
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</taxonomicName>
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(2).
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</paragraph>
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</caption>
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<subSubSection id="C33A650D6668FFABAFF0FA4DBD87FA54" pageId="2" pageNumber="565" type="materials_examined">
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<paragraph id="8B9F36866668FFABAFF0FA4DBD87FA54" blockId="2.[151,1436,1464,1921]" pageId="2" pageNumber="565">
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<emphasis id="B954EA946668FFABAFF0FA4DBBFCFA70" bold="true" box="[151,733,1464,1489]" pageId="2" pageNumber="565">
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Material studied:
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<collectingCountry id="F33776166668FFABAE08FA4DBB3CFA70" box="[367,541,1464,1489]" name="Venezuela" pageId="2" pageNumber="565">VENEZUELA</collectingCountry>
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(Aragua State):
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</emphasis>
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2 ♂♂, Rancho Grande (i.e., Henri Pittier National Park), near Maracay, presumably spring 1982 (see Materials and methods), leg. H. Franz (NHMW, cPJ).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A650D6668FFAAAFA0F9F4B824F880" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565" type="discussion">
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<paragraph id="8B9F36866668FFABAFA0F9F4BB92F820" blockId="2.[151,1436,1464,1921]" pageId="2" pageNumber="565">
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<emphasis id="B954EA946668FFABAFA0F9F4B81AF9B9" bold="true" box="[199,315,1537,1560]" pageId="2" pageNumber="565">Remarks.</emphasis>
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This species has been known so far from a single
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<typeStatus id="549B88246668FFABACD7F9F5BD33F9B8" box="[944,1042,1536,1561]" pageId="2" pageNumber="565" type="holotype">holotype</typeStatus>
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male collected in the same area (Jałoszyński 2012). The aedeagus of the
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<typeStatus id="549B88246668FFABAD1BF9D1BBFFF99C" box="[636,734,1572,1597]" pageId="2" pageNumber="565" type="holotype">holotype</typeStatus>
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is in repose (Figs 7–8), whereas those of the newly found specimens are erected, one representing a possible intermediary stage (Figs 9–10) and the other one with fully extruded endophallic sclerites and membranes. In the
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<typeStatus id="549B88246668FFABAC62F999BA46F924" box="[773,871,1644,1669]" pageId="2" pageNumber="565" type="holotype">holotype</typeStatus>
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(Figs 7–8), only apices of the lateral projections (
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<emphasis id="B954EA946668FFABAFC7F965B9E5F909" box="[160,196,1680,1704]" italics="true" pageId="2" pageNumber="565">lap</emphasis>
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) extend beyond the apex of the median lobe, and the membranous projections (
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<emphasis id="B954EA946668FFABABEFF964BD9FF909" box="[1160,1214,1681,1704]" italics="true" pageId="2" pageNumber="565">mmp</emphasis>
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) are folded in the subapical area, inside the median lobe. In the intermediary stage (Figs 9–10) the membranous projections are still folded inside the median lobe, but shifted more distally, while the lateral projections are largely extruded and pointing apically. When all movable structures are fully extended (Figs 11–12), the membranous projections (
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<emphasis id="B954EA946668FFABAA3BF908BCB3F8B5" box="[1372,1426,1789,1812]" italics="true" pageId="2" pageNumber="565">mmp</emphasis>
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) are inflated, bent at a nearly right angle and extending laterally outside the median lobe; the lateral projections (
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<emphasis id="B954EA946668FFABAA08F8D5BCB2F899" box="[1391,1427,1824,1848]" italics="true" pageId="2" pageNumber="565">lap</emphasis>
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) are also fully extruded and rotated laterally, so that they are perpendicular to the long axis of the aedeagus; and the median projection (
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<emphasis id="B954EA946668FFABAE15F89CB880F821" box="[370,417,1897,1920]" italics="true" pageId="2" pageNumber="565">mep</emphasis>
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) is rotated dorsobasally.
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</paragraph>
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<paragraph id="8B9F36866669FFAAAFF0F965B824F880" blockId="3.[151,1436,1680,1825]" pageId="3" pageNumber="566">
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<emphasis id="B954EA946669FFAAAFF0F965B866F904" bold="true" box="[151,327,1680,1701]" pageId="3" pageNumber="566">FIGURES 3–14.</emphasis>
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Aedeagus in ventral (3, 5, 7, 9, 11, 13) and lateral (4, 6, 8, 10, 12, 14) views.
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<taxonomicName id="4C204D056669FFAAAB00F965BCB4F907" box="[1127,1429,1680,1702]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="maximum">
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<emphasis id="B954EA946669FFAAAB00F965BCB4F907" box="[1127,1429,1680,1702]" italics="true" pageId="3" pageNumber="566">Pseudocephennium maximum</emphasis>
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</taxonomicName>
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, non-type specimen, non-erected (3−4) and
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<typeStatus id="549B88246669FFAAAD01F95ABBE0F964" box="[614,705,1711,1733]" pageId="3" pageNumber="566" type="holotype">holotype</typeStatus>
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, fully erected (5−6);
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<taxonomicName id="4C204D056669FFAAACCBF95ABDD0F964" box="[940,1265,1711,1733]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="araguanum">
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<emphasis id="B954EA946669FFAAACCBF95ABDD0F964" box="[940,1265,1711,1733]" italics="true" pageId="3" pageNumber="566">Pseudocephennium araguanum</emphasis>
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</taxonomicName>
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,
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<typeStatus id="549B88246669FFAAAA64F95ABC7FF964" box="[1283,1374,1711,1733]" pageId="3" pageNumber="566" type="holotype">holotype</typeStatus>
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, nonerected (7−8), and two non-type specimens, partly erected (9−10) and fully erected (11−12);
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<taxonomicName id="4C204D056669FFAAAB33F938BCBDF942" box="[1108,1436,1741,1763]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="auriculatum" status="sp. nov.">
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<emphasis id="B954EA946669FFAAAB33F938BCBDF942" box="[1108,1436,1741,1763]" italics="true" pageId="3" pageNumber="566">Pseudocephennium auriculatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26757EF6669FFAAAFF0F918B9F0F8A3" box="[151,209,1773,1794]" pageId="3" pageNumber="566" rank="species">
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<emphasis id="B954EA946669FFAAAFF0F918B9F0F8A3" bold="true" box="[151,209,1773,1794]" pageId="3" pageNumber="566">sp. n.</emphasis>
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</taxonomicNameLabel>
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,
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<typeStatus id="549B88246669FFAAAFB8F919B819F8A3" box="[223,312,1772,1794]" pageId="3" pageNumber="566" type="holotype">holotype</typeStatus>
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(13−14). Abbreviations: dph, diaphragm; lap, lateral projection; mep, median projection; mmp, membranous projection.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |