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<document id="B98E017828D6EEC732B42F98128FF601" ID-DOI="10.11646/zootaxa.4088.4.7" ID-GBIF-Dataset="32a23ac3-40ec-4578-868d-8716d5bb9bfe" ID-ISSN="1175-5326" ID-Zenodo-Dep="263904" ID-ZooBank="59D52D3C-3FD7-42A0-BDE9-8FC58DA38787" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457683834578" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="038987906669FFADAFF0F892BCA9FA44" docLanguage="en" docName="15607-5127-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocephennium auriculatum Jałoszyński, 2016, sp. n." docType="treatment" docVersion="10" lastPageNumber="567" masterDocId="FFB0FFE8666AFFA9AF67FFF5B921FFA1" masterDocTitle="A new species and new records of Pseudocephennium Reitter in Venezuela, with notes on movable endophallic structures (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="570" masterPageNumber="563" pageNumber="566" updateTime="1698177983155" updateUser="plazi">
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<mods:title id="4A87F2D33D10ACDCCD5A5B814CB9ADB1">A new species and new records of Pseudocephennium Reitter in Venezuela, with notes on movable endophallic structures (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="82D796287FA57D1858B7AA14AFC7A397">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="24D332FE7853526FDE1EB43A9F14D3F1">2016</mods:date>
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<mods:identifier id="C788A7C92E7B6D5AE0D03E6E6BA205B1" type="DOI">10.11646/zootaxa.4088.4.7</mods:identifier>
<mods:identifier id="B4A5E3025F5EE5EA07334EF2FB812481" type="GBIF-Dataset">32a23ac3-40ec-4578-868d-8716d5bb9bfe</mods:identifier>
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<treatment id="038987906669FFADAFF0F892BCA9FA44" ID-DOI="http://doi.org/10.5281/zenodo.6068506" ID-GBIF-Taxon="125783126" ID-Zenodo-Dep="6068506" LSID="urn:lsid:plazi:treatment:038987906669FFADAFF0F892BCA9FA44" httpUri="http://treatment.plazi.org/id/038987906669FFADAFF0F892BCA9FA44" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566">
<subSubSection id="C33A650D6669FFAAAFF0F892B871F803" pageId="3" pageNumber="566" type="nomenclature">
<paragraph id="8B9F36866669FFAAAFF0F892BB4AF820" blockId="3.[151,619,1895,1954]" box="[151,619,1895,1921]" pageId="3" pageNumber="566">
<heading id="D0D781EA6669FFAAAFF0F892BB4AF820" bold="true" box="[151,619,1895,1921]" fontSize="11" level="1" pageId="3" pageNumber="566" reason="1">
<emphasis id="B954EA946669FFAAAFF0F892BB4AF820" bold="true" box="[151,619,1895,1921]" pageId="3" pageNumber="566">
<taxonomicName id="4C204D056669FFAAAFF0F892BB03F820" ID-CoL="4NR35" box="[151,546,1895,1921]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="auriculatum" status="sp. nov.">
<emphasis id="B954EA946669FFAAAFF0F892BB03F820" bold="true" box="[151,546,1895,1921]" italics="true" pageId="3" pageNumber="566">Pseudocephennium auriculatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757EF6669FFAAAD4EF892BB4AF820" box="[553,619,1895,1921]" pageId="3" pageNumber="566" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F36866669FFAAAFF0F87CB871F803" blockId="3.[151,619,1895,1954]" box="[151,336,1929,1954]" pageId="3" pageNumber="566">(Figs 1314, 22)</paragraph>
</subSubSection>
<subSubSection id="C33A650D6669FFADAFF0F827BB8FFF59" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566" type="materials_examined">
<paragraph id="8B9F36866669FFADAFF0F827BB8FFF59" blockId="3.[151,1436,2001,2026]" lastBlockId="4.[151,1437,151,1509]" lastPageId="4" lastPageNumber="567" pageId="3" pageNumber="566">
<emphasis id="B954EA946669FFAAAFF0F827BAC2F84B" bold="true" box="[151,995,2001,2026]" pageId="3" pageNumber="566">
<typeStatus id="549B88246669FFAAAFF0F827B9F3F848" box="[151,210,2002,2025]" pageId="3" pageNumber="566">Type</typeStatus>
material studied:
<typeStatus id="549B88246669FFAAAEDEF824BB7DF84B" box="[441,604,2001,2026]" pageId="3" pageNumber="566" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F33776166669FFAAAD0BF824BA3BF84B" box="[620,794,2001,2026]" name="Venezuela" pageId="3" pageNumber="566">VENEZUELA</collectingCountry>
(Aragua State):
</emphasis>
♂, two labels: &quot;Rancho Grande [i.e., Henri Pittier National Park] / b. [bei] Maracay, Vene- / zuela, leg. Franz&quot; with &quot;SA258&quot; on the reverse side [white, printed, reverse handwritten in green ball pen]. &quot;
<taxonomicName id="4C204D05666EFFADADA4FF49BAC5FF75" box="[707,996,188,212]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="B954EA94666EFFADADA4FF49BAC5FF75" box="[707,996,188,212]" italics="true" pageId="4" pageNumber="567">PSEUDOCEPHENNIUM</emphasis>
</taxonomicName>
/
<taxonomicName id="4C204D05666EFFADAC99FF49BDA6FF75" box="[1022,1159,188,212]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="auriculatum">
<emphasis id="B954EA94666EFFADAC99FF49BDA6FF75" box="[1022,1159,188,212]" italics="true" pageId="4" pageNumber="567">auriculatum</emphasis>
</taxonomicName>
m. / det. P. Jałoszyński, 2015 /
<typeStatus id="549B8824666EFFADAF83FF15B8AFFF59" box="[228,398,224,248]" pageId="4" pageNumber="567" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (NHMW).
</paragraph>
</subSubSection>
<subSubSection id="C33A650D666EFFADAFA0FEF1BAC7FEC4" pageId="4" pageNumber="567" type="diagnosis">
<paragraph id="8B9F3686666EFFADAFA0FEF1BAC7FEC4" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
<emphasis id="B954EA94666EFFADAFA0FEF1B860FEBC" bold="true" box="[199,321,260,285]" pageId="4" pageNumber="567">Diagnosis.</emphasis>
Aedeagus with long apical lateral projections oblique in relation to the long axis of median lobe, with broad bases and subrectangular apical portions; median lobe in ventral view narrowly and deeply constricted in subapical region and broadly, shallowly constricted in submedian region.
</paragraph>
</subSubSection>
<subSubSection id="C33A650D666EFFADAFA0FE9AB9F7FAAD" pageId="4" pageNumber="567" type="description">
<paragraph id="8B9F3686666EFFADAFA0FE9AB80CFE6E" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
<emphasis id="B954EA94666EFFADAFA0FE9AB878FE29" bold="true" box="[199,345,367,392]" pageId="4" pageNumber="567">Description.</emphasis>
Externally indistinguishable from
<taxonomicName id="4C204D05666EFFADAD97FE9ABD76FE26" box="[752,1111,367,391]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="araguanum">
<emphasis id="B954EA94666EFFADAD97FE9ABD76FE26" box="[752,1111,367,391]" italics="true" pageId="4" pageNumber="567">Pseudocephennium araguanum</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A03666EFFADAB0CFE9ABD92FE29" box="[1131,1203,367,392]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1403,1424]" captionTargetBox="[176,1413,220,1366]" captionTargetId="figure@2.[151,1436,195,1381]" captionTargetPageId="2" captionText="FIGURES 1 2. Dorsal habitus of Pseudocephennium maximum (1) and P. araguanum (2)." httpUri="https://zenodo.org/record/263905/files/figure.png" pageId="4" pageNumber="567">Fig. 2</figureCitation>
); body suboval and strongly convex, light reddish brown, constriction between pronotum and elytra barely marked, setae yellowish; BL 1.23 mm.
</paragraph>
<paragraph id="8B9F3686666EFFADAFA0FE29BDB3FD25" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">Head broadest at small, moderately convex and coarsely faceted eyes, HL 0.13 mm, HW 0.30 mm; vertex and frons confluent, weakly convex; supraantennal tubercles weakly raised. Punctures on vertex and frons fine and inconspicuous, setae sparse, short, suberect. Antennae short, strongly thickened distally, AnL 0.50 mm, antennomeres III distinctly elongated, IIIVII indistinctly elongated, VIIIX each distinctly transverse, XI about as long as IXX together, about 1.6× as long as broad, broader than X and with blunt apex.</paragraph>
<paragraph id="8B9F3686666EFFADAFA0FD65BB7CFCFD" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">Pronotum semioval, strongly convex, broadest at base; PL 0.38 mm, PW 0.55 mm. Anterior and lateral margins in anterior half strongly rounded, sides nearly parallel in posterior fourth; posterior pronotal corners nearly right-angled, with acute tips; posterior margin shallowly bisinuate; lateral marginal carinae narrow and sharply demarcated from disc; lateral antebasal pits very shallow but distinct, each located slightly closer to posterior than lateral pronotal margin. Punctures as fine and inconspicuous as those on frons and vertex; setae moderately dense, short, nearly recumbent to slightly erect.</paragraph>
<paragraph id="8B9F3686666EFFADAFA0FC92B8B9FC4C" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
Elytra about as convex as pronotum, oval, broadest in front of middle; EL 0.73 mm, EW 0.58 mm, EI 1.26; subhumeral lines very short and indistinct, posteriorly not extending beyond distinct humeral calli; humeral carinae distinct, sharply marked, slightly shorter than 0.2 EL. Punctures and setae similar to those on pronotum.
<collectingCountry id="F3377616666EFFADAA03FC5ABCBDFC69" box="[1380,1436,943,968]" name="India" pageId="4" pageNumber="567">Hind</collectingCountry>
wings fully developed.
</paragraph>
<paragraph id="8B9F3686666EFFADAFA0FC0DBBF9FBB1" blockId="4.[151,1437,151,1509]" box="[199,728,1015,1040]" pageId="4" pageNumber="567">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="8B9F3686666EFFADAFA0FBE9B9F7FAAD" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
Aedeagus (Figs 1314) stout and symmetrical, AeL 0.35 mm; capsular part of median lobe in ventral view broadest near base, shallowly and broadly constricted in submedian region and deeply and narrowly constricted (nearly notched) in subapical region. Distal region of median lobe with one pair of large lateral projections (
<emphasis id="B954EA94666EFFADAA08FB91BCB2FBDD" box="[1391,1427,1124,1148]" italics="true" pageId="4" pageNumber="567">lap</emphasis>
) broadly separated at middle (where apex of short median projection (
<emphasis id="B954EA94666EFFADACD9FB7DBACCFB3E" box="[958,1005,1160,1183]" italics="true" pageId="4" pageNumber="567">mep</emphasis>
) can be seen), lateral projections are broad at bases and narrowing distally up to truncated, subrectangular apices. In lateral view apical assemblage of projections is very large and bent dorsally. Parameres very slender, with broadened apices, each with three apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C33A650D666EFFADAFA0FAE2BBD2FA91" box="[199,755,1303,1328]" pageId="4" pageNumber="567" type="distribution">
<paragraph id="8B9F3686666EFFADAFA0FAE2BBD2FA91" blockId="4.[151,1437,151,1509]" box="[199,755,1303,1328]" pageId="4" pageNumber="567">
<emphasis id="B954EA94666EFFADAFA0FAE2B841FA91" bold="true" box="[199,352,1303,1328]" pageId="4" pageNumber="567">Distribution.</emphasis>
North-western
<collectingCountry id="F3377616666EFFADAD77FAE2BBA2FA91" box="[528,643,1303,1328]" name="Venezuela" pageId="4" pageNumber="567">Venezuela</collectingCountry>
(
<figureCitation id="131B2A03666EFFADADF5FAE2BBC7FA91" box="[658,742,1303,1328]" captionStart="FIGURE 22" captionStartId="6.[151,250,1189,1210]" captionTargetBox="[151,1413,193,1167]" captionTargetId="figure@6.[151,1436,193,1167]" captionTargetPageId="6" captionText="FIGURE 22. Distribution of Pseudocephennium (except P. integricolle Reitter, recorded generally from Venezuela without precise locality)." httpUri="https://zenodo.org/record/263907/files/figure.png" pageId="4" pageNumber="567">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A650D666EFFADAFA0FAC8B858FAD9" pageId="4" pageNumber="567" type="etymology">
<paragraph id="8B9F3686666EFFADAFA0FAC8B858FAD9" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
<emphasis id="B954EA94666EFFADAFA0FAC8B86FFAF5" bold="true" box="[199,334,1341,1364]" pageId="4" pageNumber="567">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C204D05666EFFADAD22FAC9BBEFFAF5" box="[581,718,1340,1364]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="auriculatum">
<emphasis id="B954EA94666EFFADAD22FAC9BBEFFAF5" box="[581,718,1340,1364]" italics="true" pageId="4" pageNumber="567">auriculatum</emphasis>
</taxonomicName>
refers to the ear-like apical projections of the aedeagus (after Latin
<emphasis id="B954EA94666EFFADAFBEFAAAB819FAD6" box="[217,312,1375,1399]" italics="true" pageId="4" pageNumber="567">auricula</emphasis>
, ear).
</paragraph>
</subSubSection>
<subSubSection id="C33A650D666EFFADAFA0FA70BCA9FA44" pageId="4" pageNumber="567" type="discussion">
<paragraph id="8B9F3686666EFFADAFA0FA70BCA9FA44" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="567">
<emphasis id="B954EA94666EFFADAFA0FA70B81AFA3D" bold="true" box="[199,315,1413,1436]" pageId="4" pageNumber="567">Remarks.</emphasis>
The
<typeStatus id="549B8824666EFFADAE10FA71B8F8FA3C" box="[375,473,1412,1437]" pageId="4" pageNumber="567" type="holotype">holotype</typeStatus>
is slightly damaged; the club of left antenna and right elytron are missing. Although this species is externally indistinguishable from
<taxonomicName id="4C204D05666EFFADADA9FA5DBA4EFA1E" box="[718,879,1448,1471]" class="Insecta" family="Staphylinidae" genus="Pseudocephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="araguanum">
<emphasis id="B954EA94666EFFADADA9FA5DBA4EFA1E" box="[718,879,1448,1471]" italics="true" pageId="4" pageNumber="567">P. araguanum</emphasis>
</taxonomicName>
, aedeagi of these sympatric species are clearly different, not only in the shape and arrangement of the apical structures, but also in the shape of the median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>