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<document id="75156CD0E213A6C19803CFB9FBCF3B2F" ID-DOI="http://dx.doi.org/10.3897/zookeys.635.9915" ID-GBIF-Dataset="3dd9eb3f-9dd4-43f8-bf05-e576d7f99cc7" ID-PMC="PMC5126513" ID-Pensoft-Pub="1313-2970-635-123" ID-PubMed="27917061" ID-ZooBank="39F78A7F204142E3BB9C7C4A6B87CD9B" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-635-123" ModsDocOrigin="ZooKeys 635" ModsDocTitle="A taxonomic revision of Neoserica (sensu lato): the species groups N.lubrica, N.obscura, and N.silvestris (Coleoptera, Scarabaeidae, Sericini)" checkinTime="1480088907804" checkinUser="pensoft" docAuthor="Liu, Wan-Gang, Fabrizi, Silvia, Bai, Ming &amp; Ahrens, Dirk" docDate="2016" docId="94D7E45E81F481382CD3A68E20109550" docLanguage="en" docName="ZooKeys 635: 123-160" docOrigin="ZooKeys 635" docSource="http://dx.doi.org/10.3897/zookeys.635.9915" docTitle="Neoserica hainana Brenske 1898, comb. n." docType="treatment" docVersion="5" lastPageNumber="140" masterDocId="FFA75D54FF87FFBDFFF7FFFDFF88FFC7" masterDocTitle="A taxonomic revision of Neoserica (sensu lato): the species groups N. lubrica, N. obscura, and N. silvestris (Coleoptera, Scarabaeidae, Sericini)" masterLastPageNumber="160" masterPageNumber="123" pageNumber="139" updateTime="1701317176665" updateUser="plazi">
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<mods:title id="E3AF988E3BA12AB0FD2683B69AD94D53">A taxonomic revision of Neoserica (sensu lato): the species groups N. lubrica, N. obscura, and N. silvestris (Coleoptera, Scarabaeidae, Sericini)</mods:title>
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<mods:namePart id="F0A4C17A6D2F951F8F49586A93E8D301">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="65E6D8498F340C0D4373BCD2857BED05">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="EFA061BB6BADD3EA386C1D0116EAE33D">Bai, Ming</mods:namePart>
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<mods:namePart id="AEBDABDFA27B7218C8666D7BD2F5046E">Ahrens, Dirk</mods:namePart>
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<treatment id="94D7E45E81F481382CD3A68E20109550" ID-GBIF-Taxon="127886818" LSID="urn:lsid:plazi:treatment:94D7E45E81F481382CD3A68E20109550" httpUri="http://treatment.plazi.org/id/94D7E45E81F481382CD3A68E20109550" lastPageId="17" lastPageNumber="140" pageId="16" pageNumber="139">
<subSubSection id="4087ACDB9B014C4F0F36E91BA240B9CA" pageId="16" pageNumber="139" type="nomenclature">
<paragraph id="A2939792222D8B1535F990EE8CFA6BC6" pageId="16" pageNumber="139">
<taxonomicName id="D7C805042D5166895B86BC061A7CA6B9" ID-CoL="46TDS" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" class="Insecta" family="Scarabaeidae" genus="Neoserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoserica hainana" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" sensu="(s.l.)" species="hainana">Neoserica (s.l.) hainana (Brenske, 1898)</taxonomicName>
<taxonomicNameLabel id="29987AE5C5B58D52DAAA5CBA60421EF5" pageId="16" pageNumber="139">comb. n.</taxonomicNameLabel>
Figs 3
<normalizedToken id="AB18870C64D30A2977AD5801D0661ECD" originalValue="A–D">A-D</normalizedToken>
, 6
</paragraph>
</subSubSection>
<subSubSection id="E51700F8CFFED21B260A94BF35C6141E" pageId="16" pageNumber="139" type="reference_group">
<paragraph id="E68C44D02C171236573C8582D6BD56C1" pageId="16" pageNumber="139">
<taxonomicName id="F375D48E3AEA36E4882D6C76E58DFAE0" class="Insecta" family="Scarabaeidae" genus="Microserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microserica hainana" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="hainana">Microserica hainana</taxonomicName>
Brenske, 1898: 216.
</paragraph>
</subSubSection>
<subSubSection id="18CBB1CD076442E86E4B72E989BB97D1" pageId="16" pageNumber="139" type="type material">
<paragraph id="0B28492520DAD322F9F8CB0A74C6A538" pageId="16" pageNumber="139">Type material examined.</paragraph>
<paragraph id="CA4B2580207CEEBDE7C86ED1D3C63425" pageId="16" pageNumber="139">
Lectotype (here designated): ♂ &quot;Hainan Schmack/
<taxonomicName id="75F41F79F62654EADBE9AF8B2FCA67FD" class="Insecta" family="Scarabaeidae" genus="Serica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serica hainana" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="hainana">Serica hainana</taxonomicName>
var. type Brsk./ Coll. v.
<normalizedToken id="DC7D2C04D47B619545EFDF3876DBA7EB" originalValue="Schönfeldt”">Schoenfeldt&quot;</normalizedToken>
(SMFD). Paralectotypes: 4 ♂♂, 3 ♀♀ &quot;Hainan Schmack/ Coll. v.
<normalizedToken id="79CE3A220F9A05E1B334DC26F84824FF" originalValue="Schönfeldt”">Schoenfeldt&quot;</normalizedToken>
(SMFD), 1 ♂ &quot;
<taxonomicName id="73846FC73204882B24F5538C27749FE6" lsidName="hainana" pageId="16" pageNumber="139" rank="species" species="hainana">hainana</taxonomicName>
var. type/ Coll. v.
<normalizedToken id="45C8720DF1B95081618CC02EF429B388" originalValue="Schönfeldt”">Schoenfeldt&quot;</normalizedToken>
(SMFD), 1 ♂ &quot;Hainan Schmack/
<taxonomicName id="8F75684C4FEB7070FF30A95820E26859" class="Insecta" family="Scarabaeidae" genus="Serica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serica" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="genus">Serica</taxonomicName>
<taxonomicName id="F6EE00CF4735D49FF9617A6FE716D178" lsidName="hainana" pageId="16" pageNumber="139" rank="species" species="hainana">hainana</taxonomicName>
type Brsk./ Coll. v.
<normalizedToken id="EC6F8BF3BC4AFD1B7663087704E2E093" originalValue="Schönfeldt">Schoenfeldt</normalizedToken>
/
<taxonomicName id="845F7ADDAB1C1B92DB0ACAA4549EBFF1" lsidName="hainana" pageId="16" pageNumber="139" rank="species" species="hainana">hainana</taxonomicName>
Brske&quot; (SMFD), 1 ♀ &quot;Hainan v.
<normalizedToken id="775165D9F5EC8B5CEFF8A1DCF7D64ECB" originalValue="Schönfeldt">Schoenfeldt</normalizedToken>
/
<taxonomicName id="BFA81E5FB27D9931AF5361821ACD2E04" class="Insecta" family="Scarabaeidae" genus="Serica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serica" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="genus">Serica</taxonomicName>
<taxonomicName id="20D4DBE3AC2FCB5CE504ABD1A74E5BA7" lsidName="hainana" pageId="16" pageNumber="139" rank="species" species="hainana">hainana</taxonomicName>
type Brsk.&quot; (ZMHB), 1 ♂ &quot;19./ Hainan Schmack/
<taxonomicName id="37D2DBE4A86053D5F3802A20DEBF5BB8" class="Insecta" family="Scarabaeidae" genus="Serica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Serica" order="Coleoptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="genus">Serica</taxonomicName>
<taxonomicName id="39307B17D6EE2F2A04C0EC552C4D7E22" lsidName="hainana" pageId="16" pageNumber="139" rank="species" species="hainana">hainana</taxonomicName>
var. type Brsk./ Coll. v.
<normalizedToken id="ED81ED003995EF6B84A223AE6DA9BA95" originalValue="Schönfeldt”">Schoenfeldt&quot;</normalizedToken>
(SMFD).
</paragraph>
</subSubSection>
<subSubSection id="6FD433B31ABFA89C244B9A30664E1708" pageId="16" pageNumber="139" type="materials_examined">
<paragraph id="9ADA72FBD6C55E24BDD4146735602D69" pageId="16" pageNumber="139">Additional material examined.</paragraph>
<paragraph id="13B6F3BA521E8B0F733188D907DA22EE" pageId="16" pageNumber="139">1 ♂ &quot;Qiongzhong, Hainan, Guangdong, 17.VII.1960, 400m, leg. Zhang Xuezhong&quot; (IZAS), 1 ♂, 1 ♀ &quot;Bawangzhen, Changjiang, Hainan, 5-7.VI.2008, leg. Ba Yibin, Lang Juntong&quot; (HBUM), 3 ♂♂ &quot;Bawangzhen, Changjiang, Hainan, 5-7.VI.2008, leg. Ba Yibin, Lang Juntong&quot; (HBUM).</paragraph>
</subSubSection>
<subSubSection id="C05F5CF41589F9D2F559B20DB861C5C7" lastPageId="17" lastPageNumber="140" pageId="16" pageNumber="139" type="redescription">
<paragraph id="059B6047D4110EEFE1035D683F6347E6" pageId="16" pageNumber="139">Redescription.</paragraph>
<paragraph id="F85FE1FD7CD83FBD2AB246F9C02AAB1F" pageId="16" pageNumber="139">Body length: 6.8 mm, length of elytra: 4.8 mm, body width: 4.2 mm. Body short-oval, black, elytra reddish brown, dorsal surface except anterior labroclypeus dull, pronotum and elytra glabrous.</paragraph>
<paragraph id="EFCD85D3A2BA051DE9D3BC735252B3B8" pageId="16" pageNumber="139">Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, convergent anteriorly; anterior angles strongly rounded; anterior margin shallowly sinuate medially, margins moderately reflexed; surface weakly convex, shiny, base dull, coarsely and densely punctate, with numerous erect setae; frontoclypeal suture indistinctly incised, nearly vanishing under dull toment; smooth area in front of eye convex, nearly as long as wide; ocular canthus short and triangular (1/3 of ocular diameter), sparsely punctate, terminal setae in lectotype lacking. Frons with fine and moderately dense punctures, without erect setae. Eyes small, ratio diameter/ interocular width: 0.4. Antenna with ten antennomeres, yellowish, club (♂) with four antennomeres and straight, as long as remaining antennomeres combined. Mentum convexly elevated and flattened anteriorly.</paragraph>
<paragraph id="C4BB356D56CE2F296FFA3C614489E674" pageId="16" pageNumber="139">
Pronotum transverse, widest shortly before base, lateral margins in basal half nearly straight and moderately convergent to middle, evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin straight, with a fine complete marginal line; surface densely and finely punctate, glabrous, with minute setae in punctures (100
<normalizedToken id="ADD02E4C51F2A431013FAA43C74E84E7" originalValue="×">x</normalizedToken>
magnification); lateral border densely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum triangular, with fine, dense punctures, glabrous.
</paragraph>
<paragraph id="4BD77583D2A9BED6D9268690D5A2570B" pageId="16" pageNumber="139">
Elytra short-oval, widest shortly behind middle, striae finely impressed, finely and densely punctate, intervals weakly convex, with sparse, fine punctures concentrated along striae, glabrous except a few single, short setae on odd intervals; epipleural edge robust, ending at nearly blunt external apical angle of elytra, epipleura densely setose; apical border without a fine fringe of microtrichomes (visible at 100
<normalizedToken id="8E365F7E9E0DEC05CCEA9F3356A68095" originalValue="×">x</normalizedToken>
magnification).
</paragraph>
<paragraph id="D7569EC78F87AA898221D2A4A1094C1D" pageId="16" pageNumber="139">
Ventral surface dull, finely and densely punctate; metasternum nearly glabrous except a few long robust setae on disc, punctures with minute setae (100
<normalizedToken id="E94955669F684693B9DE7E43577E420F" originalValue="×">x</normalizedToken>
magnification); metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short robust seta, last sternite half as long as penultimate one. Mesosternum between mesocoxae as wide as the mesofemur, with a semi-circular ridge bearing long setae. Ratio of length of metepisternum/ metacoxa: 1/ 2.0. Pygidium dull, moderately convex, finely and densely punctate, without smooth midline, with a few long setae along apical margin.
</paragraph>
<paragraph id="CD404F71A73B329F11F321E93B77FD92" pageId="16" pageNumber="139">Legs short; femora moderately shiny, with two rudimentary longitudinal rows of setae, finely and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, in apical half only weakly widened, posterior margin smooth dorsally. Metatibia wide and short, widest at middle, ratio of width/ length: 1/ 2.8; dorsal margin sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate and slightly concavely sinuate. Tarsomeres ventrally with sparse, short setae, smooth, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, glabrous; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate, distal tooth sharply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex.</paragraph>
<paragraph id="9F48AA8C9BF8AAEE0365E155BD0DF273" pageId="17" pageNumber="140">
<pageBreakToken id="B14139F3E3F1DE31DB9C50BE35B7C333" pageId="17" pageNumber="140" start="start">Aedeagus</pageBreakToken>
: Fig. 3
<normalizedToken id="E8F4E4B102235EF1963B3BC2D9F0C1C6" originalValue="A–C">A-C</normalizedToken>
. Habitus: Fig. 3D.
</paragraph>
<caption id="FB1B87FED5C770717ACA644448E84574" pageId="17" pageNumber="140">
<paragraph id="7382C5E9D2CEE55D33E914F16DA86632" pageId="17" pageNumber="140">
Figure 3.
<normalizedToken id="845E5429155609481B5985EB369E9383" originalValue="A–D">A-D</normalizedToken>
<taxonomicName id="900413ECCC282F999897558A8FF93773" class="Insecta" family="Scarabaeidae" genus="Neoserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoserica hainana" order="Coleoptera" pageId="17" pageNumber="140" phylum="Arthropoda" rank="species" species="hainana">Neoserica hainana</taxonomicName>
(Brenske) (lectotype)
<normalizedToken id="256B65E373098CC031FB85D555F169AB" originalValue="E–H">E-H</normalizedToken>
<taxonomicName id="B8E96F19A89E38D8265B5370B129612F" class="Insecta" family="Scarabaeidae" genus="Neoserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoserica shoyungi" order="Coleoptera" pageId="17" pageNumber="140" phylum="Arthropoda" rank="species" species="shoyungi">Neoserica shoyungi</taxonomicName>
Ahrens, Fabrizi &amp; Liu, sp. n. (holotype)
<normalizedToken id="F6D74D28C5F67152DD8DEF8A7BA526ED" originalValue="I–L">I-L</normalizedToken>
<taxonomicName id="F11F689E43D38C975017B96B57E8477E" class="Insecta" family="Scarabaeidae" genus="Neoserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoserica sakoliana" order="Coleoptera" pageId="17" pageNumber="140" phylum="Arthropoda" rank="species" species="sakoliana">Neoserica sakoliana</taxonomicName>
Ahrens, Fabrizi &amp; Liu, sp. n. (holotype) A, E aedeagus, left side lateral view C, G aedeagus, right side lateral view B, F parameres, dorsal view D, H habitus. Scale bars: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0C40ADE46CD2F9E24DC5FB2202D5B17B" pageId="17" pageNumber="140" type="variation">
<paragraph id="027DE768A65475C29D7CB7054135DC2F" pageId="17" pageNumber="140">Variation.</paragraph>
<paragraph id="307DDB3852F2B038FA4E1CCEB609A193" pageId="17" pageNumber="140">The colour varies from an entirely black body, or a blackish anterior body (head and pronotum) with reddish elytra, to a nearly entirely reddish body with dark head and anterior pronotum, dorsal surface sometimes with greenish shine. Female: pygidium moderately convex, at middle strongly shiny and finely punctate; antennal club slightly shorter than the remaining antennomeres combined, composed of 4 antennomeres.</paragraph>
</subSubSection>
</treatment>
</document>
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