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<document id="70029706E5B59B5DD303D130F56D8D16" ID-DOI="10.11646/zootaxa.3860.1.2" ID-GBIF-Dataset="a26eaf5f-4eba-45ad-8806-dd7288d78493" ID-ISSN="1175-5326" ID-Zenodo-Dep="287060" ID-ZooBank="B9713C6D-BC3B-4EF2-8495-8BDCD535A80D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460903601137" checkinUser="plazi" docAuthor="Nakthong, La-Au, Vitheepradit, Akekawat &amp; Sites, Robert W." docDate="2014" docId="03DF879BFFC8FFD209C99FB549E83566" docLanguage="en" docName="zt03860p063.pdf" docOrigin="Zootaxa 3860 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amemboa Esaki" docType="treatment" docVersion="5" lastPageNumber="49" masterDocId="FFE6FFE3FFCAFFD0095E9B004B3B3256" masterDocTitle="Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun Mountain Range" masterLastPageNumber="63" masterPageNumber="47" pageNumber="49" updateTime="1698360479435" updateUser="plazi">
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<mods:title id="E4273039F7C63B5A960EDAAF6DD31C81">Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun Mountain Range</mods:title>
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<mods:namePart id="AB553325D0F6E637FDC42409E0B08051">Nakthong, La-Au</mods:namePart>
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<mods:namePart id="66648747CB71E47D915A5DAD29E2F01A">Sites, Robert W.</mods:namePart>
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<treatment id="03DF879BFFC8FFD209C99FB549E83566" ID-DOI="http://doi.org/10.5281/zenodo.6137507" ID-GBIF-Taxon="127669781" ID-Zenodo-Dep="6137507" LSID="urn:lsid:plazi:treatment:03DF879BFFC8FFD209C99FB549E83566" httpUri="http://treatment.plazi.org/id/03DF879BFFC8FFD209C99FB549E83566" lastPageNumber="49" pageId="2" pageNumber="49">
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<paragraph id="8BC9368DFFC8FFD209C99FB54A8E3699" blockId="2.[151,437,1205,1264]" box="[151,437,1205,1231]" pageId="2" pageNumber="49">
<heading id="D08181E1FFC8FFD209C99FB54A8E3699" bold="true" box="[151,437,1205,1231]" fontSize="11" level="1" pageId="2" pageNumber="49" reason="1">
<emphasis id="B902EA9FFFC8FFD209C99FB54A8E3699" bold="true" box="[151,437,1205,1231]" pageId="2" pageNumber="49">
Genus
<taxonomicName id="4C764D0EFFC8FFD209B09FB54A8E3699" ID-CoL="TNV" authority="Esaki" authorityName="Esaki" box="[238,437,1205,1231]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA9FFFC8FFD209B09FB54A533699" bold="true" box="[238,360,1205,1231]" italics="true" pageId="2" pageNumber="49">Amemboa</emphasis>
Esaki
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<paragraph id="8BC9368DFFC8FFD209C99FD74A1236A6" blockId="2.[151,437,1205,1264]" box="[151,297,1239,1264]" pageId="2" pageNumber="49">
<figureCitation id="134D2A08FFC8FFD209C99FD74BDF36A6" box="[151,228,1239,1264]" captionStart="FIGURES 3 5" captionStartId="3.[151,264,1892,1915]" captionTargetBox="[168,1434,452,1871]" captionTargetId="figure@3.[168,1453,452,1872]" captionTargetPageId="3" captionText="FIGURES 3 5. Genera of male Eotrechinae. (3) Amemboa, (4) Eotrechus, (5) Onychotrechus." httpUri="https://zenodo.org/record/287062/files/figure.png" pageId="2" pageNumber="49">Figs. 3</figureCitation>
,
<figureCitation id="134D2A08FFC8FFD209AF9FD74A1236A6" box="[241,297,1239,1264]" captionStart-0="FIGURES 8 21" captionStart-1="FIGURES 22 35" captionStartId-0="6.[151,264,1668,1691]" captionStartId-1="8.[151,264,1910,1933]" captionTargetBox-0="[151,1436,229,1647]" captionTargetBox-1="[152,1434,193,1888]" captionTargetId-0="figure@6.[151,1436,229,1647]" captionTargetId-1="figure@8.[152,1435,193,1889]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="FIGURES 8 21. Left forelegs of male Amemboa. (8) A. aquafrigida, (9) A. armata, (10) A. brevifasciata, (11) A. cristata, (12) A. heissi, (13) A. incurvata, (14) A. javanica, (15) A. laotica, (16) A. prostata, (17) A. riparia, (18) A. schwendingeri, (19) A. speciosa, (20) A. perlata, (21) A. velaris." captionText-1="FIGURES 22 35. Terminal abdominal segments of male Amemboa. (22) A. aquafrigida, (23) A. armata, (24) A. brevifasciata, (25) A. cristata, (26) A. heissi, (27) A. incurvata, (28) A. javanica, (29) A. laotica, (30) A. prostata, (31) A. riparia, (32) A. schwendingeri, (33) A. speciosa, (34) A. perlata, (35) A. velaris." httpUri-0="https://zenodo.org/record/287064/files/figure.png" httpUri-1="https://zenodo.org/record/287065/files/figure.png" pageId="2" pageNumber="49">835</figureCitation>
</paragraph>
<paragraph id="8BC9368DFFC8FFD209C99E1F49E83566" blockId="2.[151,1436,1311,1840]" pageId="2" pageNumber="49">
This genus is distributed in Asia from
<collectingCountry id="F361761DFFC8FFD20B1B9E1F4945376E" box="[581,638,1311,1336]" name="India" pageId="2" pageNumber="49">India</collectingCountry>
to the
<collectingCountry id="F361761DFFC8FFD20B929E1F4871376E" box="[716,842,1311,1336]" name="Philippines" pageId="2" pageNumber="49">Philippines</collectingCountry>
(
<bibRefCitation id="EFE74B7CFFC8FFD20A079E1F4FA1376E" author="Polhemus" box="[857,1178,1311,1336]" pageId="2" pageNumber="49" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
, Zettel et al. 2007) and was revised by
<bibRefCitation id="EFE74B7CFFC8FFD2081A9E44499D370A" author="Polhemus" box="[324,678,1348,1373]" pageId="2" pageNumber="49" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus and Andersen (1984)</bibRefCitation>
with a key for 23 species requiring male specimens. Later, 11 more species were described from Southeast Asia (
<bibRefCitation id="EFE74B7CFFC8FFD20BF89E67481437D6" author="Zettel" box="[678,815,1383,1408]" pageId="2" pageNumber="49" refString="Zettel, H. (1995) Two new Amemboa Esaki, 1925 from Borneo and Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici, 87, 63 - 66." type="journal article" year="1995">Zettel 1995</bibRefCitation>
, 1998, 2002;
<bibRefCitation id="EFE74B7CFFC8FFD20A8F9E674F8537D6" author="Zettel" box="[977,1214,1383,1408]" pageId="2" pageNumber="49" refString="Zettel, H. &amp; Chen, P. - P. (1996) Beitrag zur Taxonomie und Faunistik der Gerridae Vietnams mit Neubeschreibungen der Gattung Andersenius gen. nov. aus der Unteramilie Ptilomerinae und weiterer Arten (Insecta: Heteroptera: Gerridae). Entomologische Abhandlungen des Museums fur Tierkunde Dresden, 57, 149 - 182." type="journal article" year="1996">Zettel &amp; Chen 1996</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC8FFD20D929E684E3C37D6" author="Zettel" box="[1228,1287,1384,1408]" pageId="2" pageNumber="49" refString="Zettel, H. &amp; Chen, P. - P. (1997) Three new taxa of Amemboa ESAKI, 1925, from Thailand and Vietnam. Annales Historico- Naturales Musei Nationalis Hungarici, 89, 93 - 101." type="journal article" year="1997">1997</bibRefCitation>
; Zettel et al. 2007). Including two species reported here from
<collectingCountry id="F361761DFFC8FFD20B8F9E8C480E37F3" box="[721,821,1420,1445]" name="Thailand" pageId="2" pageNumber="49">Thailand</collectingCountry>
for the first time, the
<collectingCountry id="F361761DFFC8FFD20D629E8C4F5437F3" box="[1084,1135,1420,1445]" name="Thailand" pageId="2" pageNumber="49">Thai</collectingCountry>
fauna contains 15 known species of
<emphasis id="B902EA9FFFC8FFD208519EAF4ABD379E" box="[271,390,1455,1480]" italics="true" pageId="2" pageNumber="49">
<taxonomicName id="4C764D0EFFC8FFD208519EAF4AB9379E" box="[271,386,1455,1480]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">Amemboa</taxonomicName>
,
</emphasis>
more than that of any other country in Indochina (Zettel 1996,
<bibRefCitation id="EFE74B7CFFC8FFD20D029EAF4E75379E" author="Chen" box="[1116,1358,1455,1480]" pageId="2" pageNumber="49" refString="Chen, P. - P. &amp; Zettel, H. (1998) Key to genera and subgenera of Gerridae (Gerromorpha) of Thailand and adjacent countries, with a check-list of species known from Thailand. Amemboa, 2, 24 - 41." type="journal article" year="1998" yearSuffix="a">Chen &amp; Zettel 1998a</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC8FFD20C059EAF4BE937BA" author="Zettel" pageId="2" pageNumber="49" refString="Zettel, H. (2002) Description of Amemboa heissi sp. n. (Heteroptera: Gerridae) from south Thailand. Linzer Biologische Beitraege, 34, 1531 - 1534." type="journal article" year="2002">Zettel 2002</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC8FFD209829ED44A9137BA" author="Vitheepradit" box="[220,426,1492,1517]" pageId="2" pageNumber="49" refString="Vitheepradit, A. (2008) The semiaquatic Heteroptera (Gerromorpha) of Thailand: Faunistics, Biogeography, and Phylogeography. Ph. D. Dissertation, University of Missouri, Columbia, Missouri, USA. [total page number not cited]" type="book" year="2008">Vitheepradit 2008</bibRefCitation>
). Mostly, members of this genus in
<collectingCountry id="F361761DFFC8FFD20A6D9ED448AC37BB" box="[819,919,1492,1517]" name="Thailand" pageId="2" pageNumber="49">Thailand</collectingCountry>
are found resting at the margins of rock pools, puddles, and streams. Moreover, they are one of few gerromorphans that are found in temporary mud puddles in
<collectingCountry id="F361761DFFC8FFD209C99D1C4BC53463" box="[151,254,1564,1589]" name="Thailand" pageId="2" pageNumber="49">Thailand</collectingCountry>
. When they are pursued, they usually jump further onto the water surface to escape (
<bibRefCitation id="EFE74B7CFFC8FFD20C579D1C4A7D340E" author="Polhemus" pageId="2" pageNumber="49" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
). The genus is represented by two subgenera,
<taxonomicName id="4C764D0EFFC8FFD20A0D9D3F48F8340E" box="[851,963,1599,1624]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA9FFFC8FFD20A0D9D3F48F8340E" box="[851,963,1599,1624]" italics="true" pageId="2" pageNumber="49">Amemboa</emphasis>
</taxonomicName>
and
<emphasis id="B902EA9FFFC8FFD20AA29D3F4FB0340E" box="[1020,1163,1599,1624]" italics="true" pageId="2" pageNumber="49">Amemboides</emphasis>
(
<bibRefCitation id="EFE74B7CFFC8FFD20DC49D3F4BE8342A" author="Polhemus" pageId="2" pageNumber="49" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
). Members of the subgenus
<taxonomicName id="4C764D0EFFC8FFD20B4D9D6449B8342B" box="[531,643,1636,1661]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA9FFFC8FFD20B4D9D6449B8342B" box="[531,643,1636,1661]" italics="true" pageId="2" pageNumber="49">Amemboa</emphasis>
</taxonomicName>
have the first segment of the middle tarsus about twice as long as the second segment, whereas members of subgenus
<emphasis id="B902EA9FFFC8FFD20B9C9D87486A34F6" box="[706,849,1671,1696]" italics="true" pageId="2" pageNumber="49">Amemboides</emphasis>
have the first segment distinctly less than twice as long as the second segment (
<bibRefCitation id="EFE74B7CFFC8FFD208859DAC48243492" author="Polhemus" box="[475,799,1708,1733]" pageId="2" pageNumber="49" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
). Six species of the subgenus
<taxonomicName id="4C764D0EFFC8FFD20D2C9DAC4FD93493" box="[1138,1250,1708,1733]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA9FFFC8FFD20D2C9DAC4FD93493" box="[1138,1250,1708,1733]" italics="true" pageId="2" pageNumber="49">Amemboa</emphasis>
</taxonomicName>
and two species of the subgenus
<emphasis id="B902EA9FFFC8FFD208339DCF4AC734BE" box="[365,508,1743,1768]" italics="true" pageId="2" pageNumber="49">Amemboides</emphasis>
were collected in the Phetchabun Mountains. Diagnoses are provided for comparisons among males only within each subgenus.
<taxonomicName id="4C764D0EFFC8FFD20A569DF44FCF355B" authority="Polhemus &amp; Andersen" authorityName="Polhemus &amp; Andersen" box="[776,1268,1780,1805]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="incurvata">
<emphasis id="B902EA9FFFC8FFD20A569DF448D7355A" box="[776,1004,1780,1805]" italics="true" pageId="2" pageNumber="49">Amemboa incurvata</emphasis>
Polhemus &amp; Andersen
</taxonomicName>
and
<taxonomicName id="4C764D0EFFC8FFD20C739DF44BE33566" authority="Zettel" authorityName="Zettel" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="laotica">
<emphasis id="B902EA9FFFC8FFD20C739DF44EA7355B" box="[1325,1436,1780,1805]" italics="true" pageId="2" pageNumber="49">A. laotica</emphasis>
Zettel
</taxonomicName>
are reported from
<collectingCountry id="F361761DFFC8FFD208F49C1749353566" box="[426,526,1815,1840]" name="Thailand" pageId="2" pageNumber="49">Thailand</collectingCountry>
for the first time.
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