treatments-xml/data/86/32/84/8632842AFF9E1D1FA624FE82FB7BFB43.xml

<document id="0D7133FC5D4AC751B4B1CD3805DA5EAB" ID-CLB-Dataset="51374" ID-DOI="10.5281/zenodo.156789" ID-GBIF-Dataset="49eb0094-f3f4-4741-9a8d-88af88ca478a" ID-ISSN="1175­5326" ID-Zenodo-Dep="156789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459583253783" checkinUser="plazi" docAuthor="Solís, Ángel &amp; Kohlmann, Bert" docDate="2003" docId="8632842AFF9E1D1FA624FE82FB7BFB43" docLanguage="en" docName="zt00139.pdf" docOrigin="Zootaxa 139" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Onthophagus notiodes Solis &amp; Kohlmann, sp. nov." docType="treatment" docVersion="10" lastPageNumber="4" masterDocId="7A0BFC52FF9C1D1CA72CFF9EFFF7FFE5" masterDocTitle="New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1698202942371" updateUser="ExternalLinkService">
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<mods:title id="1F461B29F3A78E1BBDDA81A544F977F2">New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama</mods:title>
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<mods:namePart id="F498D713DECAEF799449DD61A8011408">Solís, Ángel</mods:namePart>
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<mods:namePart id="85440CEA2677D4D28DF6569E40779A6D">Kohlmann, Bert</mods:namePart>
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<treatment id="8632842AFF9E1D1FA624FE82FB7BFB43" ID-DOI="http://doi.org/10.5281/zenodo.6274458" ID-GBIF-Taxon="119330982" ID-Zenodo-Dep="6274458" LSID="urn:lsid:plazi:treatment:8632842AFF9E1D1FA624FE82FB7BFB43" httpUri="http://treatment.plazi.org/id/8632842AFF9E1D1FA624FE82FB7BFB43" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="468166B7FF9E1D1EA624FE82FC2BFED0" box="[264,988,283,310]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="0E24353CFF9E1D1EA624FE82FC2BFED0" blockId="2.[264,988,283,310]" box="[264,988,283,310]" pageId="2" pageNumber="3">
<heading id="556C8250FF9E1D1EA624FE82FC2BFED0" bold="true" box="[264,988,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="3CEFE92EFF9E1D1EA624FE82FC2BFED0" bold="true" box="[264,988,283,310]" pageId="2" pageNumber="3">
<taxonomicName id="C99B4EBFFF9E1D1EA624FE82FCFEFED0" ID-CoL="7XNPH" authority="Solis &amp; Kohlmann" authorityName="Solis &amp; Kohlmann" box="[264,777,283,310]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="notiodes" status="sp. nov.">
<emphasis id="3CEFE92EFF9E1D1EA624FE82FDEEFED3" bold="true" box="[264,537,284,310]" italics="true" pageId="2" pageNumber="3">Onthophagus notiodes</emphasis>
Solís &amp; Kohlmann
</taxonomicName>
, 
<taxonomicNameLabel id="27DC5455FF9E1D1EA43AFE85FC83FED0" box="[790,884,283,309]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="96A029B9FF9E1D1EA4AFFE85FC23FED0" box="[899,980,283,309]" captionStart="FIGURE 2" captionStartId="5.[264,368,1870,1894]" captionTargetBox="[339,1251,320,1854]" captionTargetId="figure@5.[329,1259,314,1858]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Onthophagus notiodes Solís &amp; Kohlmann, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168691/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9E1D1EA624FEF5FD9FFD23" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="0E24353CFF9E1D1EA624FEF5FD9FFD23" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">
<emphasis id="3CEFE92EFF9E1D1EA624FEF5FE7FFE60" bold="true" box="[264,392,363,389]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
This species is distinguished by the following combination of characters: pronotal surface without setae; body colour uniform, reddish­brown with cupreous red or green metallic reflections, meso­and metafemora not yellow in their central part; setae of the apex of the metatibia always thick, forming a comb, stiff, always the same size and with fine setae alternating within them and with a size never exceeding 2.5 times the size of the stiff setae; elytra surface much sharpened and punctate; males have small teeth at the base of the head; short, broad, massive clypeal horn; a very square angle exists between the clypeal tooth and the gena; pronotal male process with lamellae that point slightly inwards and upwards.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9E1D1EA614FD4DFE7BF933" pageId="2" pageNumber="3" type="description">
<paragraph id="0E24353CFF9E1D1EA614FD4DFC85FCDB" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">
<emphasis id="3CEFE92EFF9E1D1EA614FD4DFD50FD08" bold="true" box="[312,679,723,749]" pageId="2" pageNumber="3">
Description. 
<typeStatus id="D1208B9EFF9E1D1EA6F6FD4DFDB9FD08" box="[474,590,723,749]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
. Male:
</emphasis>
Length: 
<quantity id="C96398D9FF9E1D1EA43BFD4AFC81FD0B" box="[791,886,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="3" unit="mm" value="7.8">7.8 mm</quantity>
. Humeral width: 
<quantity id="C96398D9FF9E1D1EA367FD4AFB5DFD0B" box="[1099,1194,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="3" unit="mm" value="4.1">4.1 mm</quantity>
. Head and pronotum cupreous red with intense golden­green reflections, elytra burgundy colours, rest of the body a dark reddish­brown with greenish cast.
</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FCD2FE4AFC3B" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">The head has a clypeal horn, thick set at its base and broadening toward the apex. The frontal area rugosely punctate, eyes small. There is a pair of small horns, teeth like, between the eyes and toward the base of the head. The angle between the clypeal tooth and the gena is 90°.</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FC72FB88FBCB" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">Pronotum slightly shagreened and uniformly punctate, prominences broadly bladelike, slightly pointing upwards, parallel and slightly converging at the apex.</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FBA2FD1DFB9B" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">Elytra with eight striae evident, each with large punctures, intervals flat and covered with fine punctation, surface shagreened.</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FB12FE95FB2B" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">Pygidium shagreened and covered with regular punctation, carina running through the middle.</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FB42FAE7FAFB" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">Protibiae long and slender, with tarsi present and four external teeth. Meso­ and metatibia dilated toward the apex. First metatarsus very long and almost rectangular in shape.</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FAB5FCCAFA03" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">
<emphasis id="3CEFE92EFF9E1D1EA614FAB5FDEAFAA0" bold="true" box="[312,541,1323,1349]" pageId="2" pageNumber="3">Allotype. Female:</emphasis>
Length 
<quantity id="C96398D9FF9E1D1EA5A3FAB2FD12FAA3" box="[655,741,1324,1350]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="3" unit="mm" value="6.9">6.9mm</quantity>
. Humeral width: 
<quantity id="C96398D9FF9E1D1EA4E4FAB2FBE9FAA3" box="[968,1054,1324,1350]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="3" unit="mm" value="3.9">3.9mm</quantity>
. Similar to male but unarmed, with a transversal carina at the middle of the head and the anterior border of the clypeus shows two well developed teeth separated by an open “v” notch, there are also two pointed small teeth, at the base of the head between the eyes. Pronotal process formed by two small blunt teeth. Foretibia short and broad.
</paragraph>
<paragraph id="0E24353CFF9E1D1EA614FA6DFE7BF933" blockId="2.[264,1325,363,1870]" pageId="2" pageNumber="3">
<emphasis id="3CEFE92EFF9E1D1EA614FA6DFE45F9E8" bold="true" box="[312,434,1523,1549]" pageId="2" pageNumber="3">Variation.</emphasis>
Length: 
<quantity id="C96398D9FF9E1D1EA504FA6AFD5BF9EB" box="[552,684,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.8" metricValueMin="6.1" pageId="2" pageNumber="3" unit="mm" value="6.949999999999999" valueMax="7.8" valueMin="6.1">6.1­7.8mm</quantity>
. Humeral width: 
<quantity id="C96398D9FF9E1D1EA4AAFA6AFBFCF9EB" box="[902,1035,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.9" metricValueMin="3.2" pageId="2" pageNumber="3" unit="mm" value="4.05" valueMax="4.9" valueMin="3.2">3.2­4.9mm</quantity>
. Specimen body colour can vary from an intense cupreous red to cupreous green. Males vary from having a well developed clypeal horn to just a small lamella; head horn and pronotal projections also vary from well to feebly developed. Less developed males lack the strongly developed angle between the clypeal teeth and the gena. Pygidium can go from a fine to a strong punctation.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9E1D1FA614F97DFE90FE1B" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="0E24353CFF9E1D1FA614F97DFB21FE4B" blockId="2.[264,1325,363,1870]" lastBlockId="3.[264,1324,284,1190]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="3CEFE92EFF9E1D1EA614F97DFDDEF918" bold="true" box="[312,553,1763,1789]" pageId="2" pageNumber="3">Examined material</emphasis>
(
<specimenCount id="189DFEB5FF9E1D1EA517F97AFD15F91B" box="[571,738,1764,1790]" pageId="2" pageNumber="3" type="generic">22 specimens</specimenCount>
). 
<emphasis id="3CEFE92EFF9E1D1EA5DAF97DFC9FF918" bold="true" box="[758,872,1763,1789]" pageId="2" pageNumber="3">
<typeStatus id="D1208B9EFF9E1D1EA5DAF97DFC9FF918" box="[758,872,1763,1789]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
, male: 
<collectingCountry id="768C75ACFF9E1D1EA4EEF97AFB84F91B" box="[962,1139,1764,1790]" name="Costa Rica" pageId="2" pageNumber="3">COSTA RICA</collectingCountry>
. Grano de Oro, 
<quantity id="C96398D9FF9E1D1EA624F892FE93F8C3" box="[264,356,1804,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="2" pageNumber="3" unit="m" value="1120.0">1120 m</quantity>
, Chirripó, Turrialba, Prov. Cartago, Feb, 1993, P. Campos. 
<emphasis id="3CEFE92EFF9E1D1EA30BF895FB66F8C0" bold="true" box="[1063,1169,1803,1829]" pageId="2" pageNumber="3">Allotype</emphasis>
, female: 
<emphasis id="3CEFE92EFF9E1D1EA22EF892FEB6F8A8" italics="true" pageId="2" pageNumber="3">ibidem,</emphasis>
Mar., 1993. 
<emphasis id="3CEFE92EFF9E1D1EA6F3F8ADFD93F8A8" bold="true" box="[479,612,1843,1869]" pageId="2" pageNumber="3">
<typeStatus id="D1208B9EFF9E1D1EA6F3F8ADFDA8F8A8" box="[479,607,1843,1869]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Cartago: Grano de Oro, 
<quantity id="C96398D9FF9E1D1EA4A5F8AAFC12F8AB" box="[905,997,1844,1870]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="2" pageNumber="3" unit="m" value="1120.0">1120 m</quantity>
, Chirripó, Turrialba, 
<date id="7A2513FCFF9E1D1FA3F3F8AAFEB2FED3" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" value="1993-03">Marzo 1993</date>
, P. Campos, 
<specimenCount id="189DFEB5FF9F1D1FA6F0FE82FD87FED3" box="[476,624,284,310]" pageId="3" pageNumber="4" type="generic">2 specimens</specimenCount>
; 
<date id="7A2513FCFF9F1D1FA557FE82FCFFFED3" box="[635,776,284,310]" pageId="3" pageNumber="4" value="1913-04-15">13­15 Abril</date>
, 
<specimenCount id="189DFEB5FF9F1D1FA43FFE82FC42FED3" box="[787,949,284,310]" pageId="3" pageNumber="4" type="generic">11 specimens</specimenCount>
; 
<date id="7A2513FCFF9F1D1FA4ECFE82FB5AFED3" box="[960,1197,284,310]" pageId="3" pageNumber="4" value="1992-09-01">
Septiembre 1992, 
<specimenCount id="189DFEB5FF9F1D1FA3B2FE82FB5AFED3" box="[1182,1197,284,310]" pageId="3" pageNumber="4" type="generic">1</specimenCount>
</date>
specimen; 
<date id="7A2513FCFF9F1D1FA624FEDAFE2EFEBB" box="[264,473,324,350]" pageId="3" pageNumber="4" value="1992-10-01">
Octubre 1992, 
<specimenCount id="189DFEB5FF9F1D1FA6E6FEDAFE2EFEBB" box="[458,473,324,350]" pageId="3" pageNumber="4" type="generic">1</specimenCount>
</date>
specimen; 
<date id="7A2513FCFF9F1D1FA55CFEDAFCCAFEBB" box="[624,829,324,350]" pageId="3" pageNumber="4" value="1993-02-01">
Febrero 1993, 
<specimenCount id="189DFEB5FF9F1D1FA403FEDAFCCAFEBB" box="[815,829,324,350]" pageId="3" pageNumber="4" type="generic">1</specimenCount>
</date>
specimen. Heredia: Estación 
<quantity id="C96398D9FF9F1D1FA393FEDAFAD2FEBB" box="[1215,1317,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="3" pageNumber="4" unit="m" value="1070.0">1070 m</quantity>
, transecto altitudinal Finca la Selva a Volcán Barva, Parque Nacional Braulio Carrillo, trampa de foso, 
<date id="7A2513FCFF9F1D1FA6E9FE0AFD93FE4B" box="[453,612,404,430]" pageId="3" pageNumber="4" value="2001-04-17">17 abril 2001</date>
, D. Brenes, 
<specimenCount id="189DFEB5FF9F1D1FA5D8FE0AFC70FE4B" box="[756,903,404,430]" pageId="3" pageNumber="4" type="generic">2 specimens</specimenCount>
, 
<date id="7A2513FCFF9F1D1FA4BFFE0AFBC6FE4B" box="[915,1073,404,430]" pageId="3" pageNumber="4" value="2001-04-18">18 abril 2001</date>
, 
<specimenCount id="189DFEB5FF9F1D1FA312FE0AFB25FE4B" box="[1086,1234,404,430]" pageId="3" pageNumber="4" type="generic">2 specimens</specimenCount>
.
</paragraph>
<paragraph id="0E24353CFF9F1D1FA614FE25FE90FE1B" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">
<emphasis id="3CEFE92EFF9F1D1FA614FE25FE68FE30" bold="true" box="[312,415,443,469]" pageId="3" pageNumber="4">Habitat.</emphasis>
<emphasis id="3CEFE92EFF9F1D1FA68BFE22FDBCFE30" box="[423,587,444,469]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="C99B4EBFFF9F1D1FA68BFE22FDD8FE30" box="[423,559,444,469]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="notiodes">O. notiodes</taxonomicName>
w
</emphasis>
as collected in cloud forest at elevations ranging from 
<quantity id="C96398D9FF9F1D1FA3FEFE22FE94FE1B" metricMagnitude="3" metricUnit="m" metricValue="1.095" metricValueMax="1.1199999999999999" metricValueMin="1.0699999999999998" pageId="3" pageNumber="4" unit="m" value="1095.0" valueMax="1120.0" valueMin="1070.0">
<date id="7A2513FCFF9F1D1FA3FEFE22FEB6FE1B" pageId="3" pageNumber="4" value="1070" valueMax="1120">1070 to 1120</date>
m
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9F1D1FA614FD95FDD5FBB3" pageId="3" pageNumber="4" type="description">
<paragraph id="0E24353CFF9F1D1FA614FD95FBEDFDAB" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">
<emphasis id="3CEFE92EFF9F1D1FA614FD95FD71FDC0" bold="true" box="[312,646,523,549]" pageId="3" pageNumber="4">Geographical distribution.</emphasis>
This species is known so far from Grano de Oro, in the province of Cartago and along the Atlantic slope of Barva volcano.
</paragraph>
<paragraph id="0E24353CFF9F1D1FA614FDC5FBA0FD7B" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">
<emphasis id="3CEFE92EFF9F1D1FA614FDC5FDAFFD90" bold="true" box="[312,600,603,629]" pageId="3" pageNumber="4">Chorological affinities.</emphasis>
The new species occupies higher terrain on the Caribbean slope, whereas 
<taxonomicName id="C99B4EBFFF9F1D1FA690FD1AFD9FFD78" box="[444,616,644,669]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA690FD1AFD9FFD78" box="[444,616,644,669]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
is found at lower altitudes on both slopes.
</paragraph>
<paragraph id="0E24353CFF9F1D1FA614FD35FDE5FBE3" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">
<emphasis id="3CEFE92EFF9F1D1FA614FD35FD85FD20" bold="true" box="[312,626,683,709]" pageId="3" pageNumber="4">Taxonomic relationships.</emphasis>
<taxonomicName id="C99B4EBFFF9F1D1FA555FD32FCF6FD20" box="[633,769,684,709]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="notiodes">
<emphasis id="3CEFE92EFF9F1D1FA555FD32FCF6FD20" box="[633,769,684,709]" italics="true" pageId="3" pageNumber="4">O. notiodes</emphasis>
</taxonomicName>
keys to 
<taxonomicName id="C99B4EBFFF9F1D1FA44BFD32FBE5FD20" box="[871,1042,684,709]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA44BFD32FBE5FD20" box="[871,1042,684,709]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
in 
<bibRefCitation id="6A0A48CDFF9F1D1FA315FD32FEADFD0B" author="Kohlmann" pageId="3" pageNumber="14" refString="Kohlmann, B. &amp; A. Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 9, 159 - 261." type="journal article" year="2001">Kohlmann and Solís (2001)</bibRefCitation>
. The males are easily distinguished by their short, broad, massive and not strongly backwardly curved clypeal horn, whereas 
<taxonomicName id="C99B4EBFFF9F1D1FA425FD62FC4FFCF0" box="[777,952,764,789]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA425FD62FC4FFCF0" box="[777,952,764,789]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
has a finer, slender and backwardly curved clypeal horn. The new species has a sharp angle between the clypeal tooth and the gena, whereas 
<taxonomicName id="C99B4EBFFF9F1D1FA53EFCD2FD4AFC80" box="[530,701,844,869]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA53EFCD2FD4AFC80" box="[530,701,844,869]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
has a rather open angle. 
<taxonomicName id="C99B4EBFFF9F1D1FA4CDFCD2FB9FFC80" box="[993,1128,844,869]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="notiodes">
<emphasis id="3CEFE92EFF9F1D1FA4CDFCD2FB9FFC80" box="[993,1128,844,869]" italics="true" pageId="3" pageNumber="4">O. notiodes</emphasis>
</taxonomicName>
has the pronotal process with lamellae that point slightly inwards and upwards, while those of 
<taxonomicName id="C99B4EBFFF9F1D1FA388FCEAFEC0FC50" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA388FCEAFEC0FC50" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
are parallel and level. In females the clypeal teeth are well developed, but the head horns are smaller than in 
<taxonomicName id="C99B4EBFFF9F1D1FA519FC5AFD17FC38" box="[565,736,964,989]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA519FC5AFD17FC38" box="[565,736,964,989]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
. Elytra are much more shagreened in 
<taxonomicName id="C99B4EBFFF9F1D1FA38FFC5AFADCFC38" box="[1187,1323,964,989]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="notiodes">
<emphasis id="3CEFE92EFF9F1D1FA38FFC5AFADCFC38" box="[1187,1323,964,989]" italics="true" pageId="3" pageNumber="4">O. notiodes</emphasis>
</taxonomicName>
than in 
<taxonomicName id="C99B4EBFFF9F1D1FA64CFC72FDFCFBE0" box="[352,523,1004,1029]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecellens">
<emphasis id="3CEFE92EFF9F1D1FA64CFC72FDFCFBE0" box="[352,523,1004,1029]" italics="true" pageId="3" pageNumber="4">O. praecellens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0E24353CFF9F1D1FA614FB8AFDD5FBB3" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">With the exception of elytral shagreenation, there are no characters separating the females of both species.</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9F1D1FA614FBFDFB7BFB43" pageId="3" pageNumber="4" type="etymology">
<paragraph id="0E24353CFF9F1D1FA614FBFDFB7BFB43" blockId="3.[264,1324,284,1190]" pageId="3" pageNumber="4">
<emphasis id="3CEFE92EFF9F1D1FA614FBFDFE32FB98" bold="true" box="[312,453,1123,1149]" pageId="3" pageNumber="4">Etymology.</emphasis>
<taxonomicName id="C99B4EBFFF9F1D1FA6E1FBFAFD36FB98" box="[461,705,1124,1150]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="6" phylum="Arthropoda" rank="species" species="notiodes">Notiodes ()</taxonomicName>
, a Greek word in apposition, meaning wet or damp, making reference to the very humid tropical forests, where this species lives.
</paragraph>
</subSubSection>
</treatment>
</document>