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<document id="32BBF9891F10215BCCC3954239BBB093" ID-DOI="10.1206/0003-0082(2007)3599[1:CANETF]2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5389079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630309346201" checkinUser="carolina" docAuthor="GAFFNEY, EUGENE S., RICH, THOMAS H., VICKERS-RICH, PATRICIA, CONSTANTINE, ANDREW, VACCA, RAUL &amp; KOOL, LESLEY" docDate="2007" docId="8606879E7D1FE84E61AB5407FC084988" docLanguage="en" docName="N3599.pdf" docOrigin="American Museum Novitates 3599" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282007%293599%5B1%3ACANETF%5D2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Chubutemys copelloi GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL 2007, new species" docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="7A3FFFE67D1CE84863745077FFD9487F" masterDocTitle="Chubutemys, a New Eucryptodiran Turtle from the Early Cretaceous of Argentina, and the Relationships of the Meiolaniidae" masterLastPageNumber="36" masterPageNumber="1" pageNumber="4" updateTime="1699222336835" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5C6E30ECFEAC7007D60D3D9600F4461D">Chubutemys, a New Eucryptodiran Turtle from the Early Cretaceous of Argentina, and the Relationships of the Meiolaniidae</mods:title>
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<taxonomicName id="C9AF4D0B7D1FE84B61AB5407FC1F4CF7" authority="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL, 2007" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[735,966,1136,1160]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="copelloi" status="sp. nov.">
<emphasis id="3CDBEA9A7D1FE84B61AB5407FC1F4CF7" bold="true" box="[735,966,1136,1160]" italics="true" pageId="3" pageNumber="4">Chubutemys copelloi</emphasis>
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,
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<materialsCitation id="BEC73CD57D1FE84B61C454DBFC9B4D0A" collectionCode="SPECIMEN" pageId="3" pageNumber="4" specimenCode="MPEF-PV1236" specimenCount="1" typeStatus="holotype">
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:
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, a partial skull lacking much of the bone from its dorsal surface, but with an internal mold of it present, ventral surfaces preserved intact with stapes in situ, atlas elements disarticulated, and a piece of presumed hyoid in right temporal fossa
</materialsCitation>
.
</paragraph>
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<materialsCitation id="BEC73CD57D1FE84B61C4550CFC064DB1" collectionCode="LOCALITY" country="Argentina" latitude="-43.602497" location="Turtle Town'" longLatPrecision="19" longitude="-68.91917" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Chubut" typeStatus="holotype">
<typeStatus id="D114882A7D1FE84B61C4550CFD314DED" box="[688,744,1403,1427]" pageId="3" pageNumber="4">TYPE</typeStatus>
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:
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,
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43
<emphasis id="3CDBEA9A7D1FE84B6714550CFBB34DEC" bold="true" box="[1120,1130,1403,1427]" italics="true" pageId="3" pageNumber="4">°</emphasis>
36
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S
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,
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68
<emphasis id="3CDBEA9A7D1FE84B61C455EEFD634DCE" bold="true" box="[688,698,1433,1457]" italics="true" pageId="3" pageNumber="4">°</emphasis>
55
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W
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(figs. 13;
<bibRefCitation id="6A3E4B797D1FE84B600B55EEFB9E4DCE" author="Rich, T. H. &amp; P. Vickers-Rich &amp; F. E. Novas &amp; R. Cuneo &amp; P. Puerta &amp; R. Vacca" box="[895,1095,1433,1457]" pageId="3" pageNumber="4" pagination="111 - 115" refId="ref14960" refString="Rich, T. H., P. Vickers-Rich, F. E. Novas, R. Cuneo, P. Puerta, and R. Vacca. 1998. Theropods from the '' middle' ' Cretaceous Chubut Group of the San Jorge sedimentary basin, central Patagonia: a preliminary note. Gaia 15: 111 - 115." type="journal article" year="1998">Rich et al., 1998</bibRefCitation>
: fig. 1),
<collectingRegion id="CC6BF86A7D1FE84B61E255C1FC854DB1" box="[662,860,1462,1486]" country="Argentina" name="Chubut" pageId="3" pageNumber="4">Chubut Province</collectingRegion>
,
<collectingCountry id="76B876187D1FE84B601D55C1FC064DB1" box="[873,991,1462,1486]" name="Argentina" pageId="3" pageNumber="4">Argentina</collectingCountry>
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.
</paragraph>
<paragraph id="0E1036887D1FE84B61C455A3FCFE4EFF" blockId="3.[662,1188,1196,1754]" pageId="3" pageNumber="4">
HORIZON: Cerro Castaño Member (fig. 3), Cerro Barcino Formation,
<collectingRegion id="CC6BF86A7D1FE84B609C5585FBE64E75" box="[1000,1087,1522,1546]" country="Argentina" name="Chubut" pageId="3" pageNumber="4">Chubut</collectingRegion>
Group, Aptian (
<bibRefCitation id="6A3E4B797D1FE84B618C5678FBD64E58" author="Codignotto, J. &amp; F. Nullo &amp; J. Panza &amp; C. Proserpio" box="[760,1039,1551,1575]" pageId="3" pageNumber="4" pagination="471 - 480" refId="ref13373" refString="Codignotto, J., F. Nullo, J. Panza, and C. Proserpio. 1978. Estratigrafia del Grupo Chubut entre Paso de Indios y Las Plumas, Provincia del Chubut, Argentina. VII Congreso Geologico Argentino, Neuquen (9 - 15 abril 1978), Actas 1: 471 - 480." type="journal article" year="1978">Codignotto et al., 1978</bibRefCitation>
;
<bibRefCitation id="6A3E4B797D1FE84B676B5678FD174E3A" author="Rich, T. H. &amp; P. Vickers-Rich &amp; F. E. Novas &amp; R. Cuneo &amp; P. Puerta &amp; R. Vacca" pageId="3" pageNumber="4" pagination="111 - 115" refId="ref14960" refString="Rich, T. H., P. Vickers-Rich, F. E. Novas, R. Cuneo, P. Puerta, and R. Vacca. 1998. Theropods from the '' middle' ' Cretaceous Chubut Group of the San Jorge sedimentary basin, central Patagonia: a preliminary note. Gaia 15: 111 - 115." type="journal article" year="1998">Rich et al., 1998</bibRefCitation>
;
<bibRefCitation id="6A3E4B797D1FE84B61AF565AFC654E3A" author="Rauhut, O. W. M. &amp; G. Cladera &amp; P. Vickers-Rich &amp; T. H. Rich" box="[731,956,1581,1605]" pageId="3" pageNumber="4" pagination="487 - 497" refId="ref14913" refString="Rauhut, O. W. M., G. Cladera, P. Vickers-Rich, and T. H. Rich. 2003. Dinosaur remains from the Lower Cretaceous of the Chubut Group, Argentina. Cretaceous Research 24: 487 - 497." type="journal article" year="2003">Rauhut et al., 2003</bibRefCitation>
) based on the work of two of the authors,
<collectionCode id="68BEAE4D7D1FE84B60A7563DFC314E1D" box="[979,1000,1610,1634]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="4">T</collectionCode>
. Rich and A. Constantine.
</paragraph>
</subSubSection>
<subSubSection id="46B565037D1FE84B61C456F1FC004EE1" box="[688,985,1670,1694]" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="0E1036887D1FE84B61C456F1FC004EE1" blockId="3.[662,1188,1196,1754]" box="[688,985,1670,1694]" pageId="3" pageNumber="4">DIAGNOSIS: as for genus.</paragraph>
</subSubSection>
<subSubSection id="46B565037D1FE84E61C456D3FDCE4BAC" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="etymology">
<paragraph id="0E1036887D1FE84B61C456D3FC774EA5" blockId="3.[662,1188,1196,1754]" pageId="3" pageNumber="4">
ETYMOLOGY: For Maria Copello, who discovered the
<typeStatus id="D114882A7D1FE84B603656B5FC734EA5" box="[834,938,1730,1754]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="5AD066007D18E84C63E35619FD864EE1" ID-DOI="http://doi.org/10.5281/zenodo.5389081" ID-Zenodo-Dep="5389081" httpUri="https://zenodo.org/record/5389081/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,189,1646,1668]" targetBox="[207,1149,210,1611]" targetPageId="4">
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Fig. 1. Regional location maps showing the position of the type locality of
<emphasis id="3CDBEA9A7D18E84C60DF5619FB594EFC" box="[939,1152,1646,1667]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="C9AF4D0B7D18E84C60DF5619FBA54EFC" authority="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL, 2007" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[939,1148,1646,1667]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="copelloi" status="n. gen. et sp.">Chubutemys copelloi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="27E857E17D18E84C67F35619FF6E4EE1" pageId="4" pageNumber="5" rank="species">n. gen. et sp.</taxonomicNameLabel>
MPEF-PV1236 (from
<bibRefCitation id="6A3E4B797D18E84C62D256FFFD8D4EE1" author="Rich, T. H. &amp; P. Vickers-Rich &amp; F. E. Novas &amp; R. Cuneo &amp; P. Puerta &amp; R. Vacca" box="[422,596,1672,1694]" pageId="4" pageNumber="5" pagination="111 - 115" refId="ref14960" refString="Rich, T. H., P. Vickers-Rich, F. E. Novas, R. Cuneo, P. Puerta, and R. Vacca. 1998. Theropods from the '' middle' ' Cretaceous Chubut Group of the San Jorge sedimentary basin, central Patagonia: a preliminary note. Gaia 15: 111 - 115." type="journal article" year="1998">Rich et al., 1998</bibRefCitation>
).
</paragraph>
</caption>
<caption id="5AD066007D19E84D630455C0FDBE4D97" ID-DOI="http://doi.org/10.5281/zenodo.5389085" ID-Zenodo-Dep="5389085" httpUri="https://zenodo.org/record/5389085/files/figure.png" pageId="5" pageNumber="6" startId="5.[112,150,1463,1485]" targetBox="[125,1151,210,1428]" targetPageId="5">
<paragraph id="0E1036887D19E84D630455C0FB7A4DB2" blockId="5.[88,1187,1463,1512]" box="[112,1187,1463,1485]" pageId="5" pageNumber="6">Fig. 2. Detailed location map (from 250K topographic map) showing the position of the type locality of</paragraph>
<paragraph id="0E1036887D19E84D632C55A5FDBE4D97" blockId="5.[88,1187,1463,1512]" box="[88,615,1490,1512]" pageId="5" pageNumber="6">
<emphasis id="3CDBEA9A7D19E84D632C55A5FEF44D98" box="[88,301,1490,1511]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="C9AF4D0B7D19E84D632C55A5FEF34D98" authority="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL, 2007" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[88,298,1490,1511]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="copelloi" status="n. gen. et sp.">Chubutemys copelloi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="27E857E17D19E84D624155A5FE624D97" box="[309,443,1490,1512]" pageId="5" pageNumber="6" rank="species">n. gen. et sp.</taxonomicNameLabel>
MPEF-PV1236.
</paragraph>
</caption>
<paragraph id="0E1036887D19E84E63065667FEA34B7D" blockId="5.[88,614,1552,1754]" lastBlockId="6.[128,654,211,1753]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
REFERRED MATERIAL: The locality that yielded the skull of
<taxonomicName id="C9AF4D0B7D19E84D624A5659FE1E4E39" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[318,455,1582,1606]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D19E84D624A5659FE1E4E39" box="[318,455,1582,1606]" italics="true" pageId="5" pageNumber="6">Chubutemys</emphasis>
</taxonomicName>
also coughed up enough turtle remains to cause its giddy discoverers to call it Turtle Town (figs. 13;
<bibRefCitation id="6A3E4B797D19E84D632C56F0FEC24EE0" author="Rich, T. H. &amp; P. Vickers-Rich &amp; F. E. Novas &amp; R. Cuneo &amp; P. Puerta &amp; R. Vacca" box="[88,283,1671,1695]" pageId="5" pageNumber="6" pagination="111 - 115" refId="ref14960" refString="Rich, T. H., P. Vickers-Rich, F. E. Novas, R. Cuneo, P. Puerta, and R. Vacca. 1998. Theropods from the '' middle' ' Cretaceous Chubut Group of the San Jorge sedimentary basin, central Patagonia: a preliminary note. Gaia 15: 111 - 115." type="journal article" year="1998">Rich et al., 1998</bibRefCitation>
: fig. 1). The fossils from the site consist of shell and postcranial fragments of cryptodires and pleurodires, as well as the skull described here. No shell or postcranial material was found directly associated with the
<typeStatus id="D114882A7D19E84D61B6563BFD2B4E1B" box="[706,754,1612,1636]" pageId="5" pageNumber="6">type</typeStatus>
skull of
<taxonomicName id="C9AF4D0B7D19E84D602F563BFC3D4E1B" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[859,996,1612,1636]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D19E84D602F563BFC3D4E1B" box="[859,996,1612,1636]" italics="true" pageId="5" pageNumber="6">Chubutemys</emphasis>
</taxonomicName>
. However, there are postcranial elements that probably belong to
<taxonomicName id="C9AF4D0B7D19E84D61CA56F0FC9E4EE0" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[702,839,1671,1695]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D19E84D61CA56F0FC9E4EE0" box="[702,839,1671,1695]" italics="true" pageId="5" pageNumber="6">Chubutemys</emphasis>
</taxonomicName>
. The cryptodire material is identifiable because it consists of a partial carapace with anterior thoracic centra and an articulated eighth cervical, as well as peripherals, including bridge peripherals, that are very similar to the sinemydid/macrobaenid postcrania described for
<taxonomicName id="C9AF4D0B7D1AE84E628A515BFD5F493B" box="[510,646,300,324]" class="Reptilia" family="Macrobaenidae" genus="Dracochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E628A515BFD5F493B" box="[510,646,300,324]" italics="true" pageId="6" pageNumber="7">Dracochelys</emphasis>
</taxonomicName>
,
<taxonomicName id="C9AF4D0B7D1AE84E63F4513DFE7C491D" authority=", Hangaiemys" authorityName="Hangaiemys" box="[128,421,330,354]" class="Reptilia" family="Macrobaenidae" genus="Ordosemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E63F4513DFF22491D" box="[128,251,330,354]" italics="true" pageId="6" pageNumber="7">Ordosemys</emphasis>
,
<emphasis id="3CDBEA9A7D1AE84E626F513DFE7C491D" box="[283,421,330,354]" italics="true" pageId="6" pageNumber="7">Hangaiemys</emphasis>
</taxonomicName>
,
<taxonomicName id="C9AF4D0B7D1AE84E62B2513DFD9A491D" authorityName=": Gaffney" authorityYear="1996" box="[454,579,330,354]" class="Reptilia" family="Macrobaenidae" genus="Judithemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E62B2513DFD9A491D" box="[454,579,330,354]" italics="true" pageId="6" pageNumber="7">Judithemys</emphasis>
</taxonomicName>
, and
<taxonomicName id="C9AF4D0B7D1AE84E63F4511FFED849FF" box="[128,257,360,384]" class="Reptilia" genus="Otwayemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E63F4511FFED849FF" box="[128,257,360,384]" italics="true" pageId="6" pageNumber="7">Otwayemys</emphasis>
</taxonomicName>
. The shell fragment with an eighth cervical, MPEF PV1940, is identified as a cryptodire on the basis of the widely spaced zygapophyses and the close agreement of this cervical with those of
<taxonomicName id="C9AF4D0B7D1AE84E62BF51A8FD9F4988" box="[459,582,479,503]" class="Reptilia" family="Macrobaenidae" genus="Ordosemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E62BF51A8FD9F4988" box="[459,582,479,503]" italics="true" pageId="6" pageNumber="7">Ordosemys</emphasis>
</taxonomicName>
and
<taxonomicName id="C9AF4D0B7D1AE84E63F4518AFED14A6A" box="[128,264,509,533]" class="Reptilia" family="Macrobaenidae" genus="Dracochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E63F4518AFED14A6A" box="[128,264,509,533]" italics="true" pageId="6" pageNumber="7">Dracochelys</emphasis>
</taxonomicName>
(see below). Three sets of peripherals are also identified as cryptodire and probably belong to
<taxonomicName id="C9AF4D0B7D1AE84E620D524FFDDB4A2F" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[377,514,568,592]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E620D524FFDDB4A2F" box="[377,514,568,592]" italics="true" pageId="6" pageNumber="7">Chubutemys</emphasis>
</taxonomicName>
. These are MPEF PV1941, PV1942, and PV1943. They are very similar to peripherals figured by
<bibRefCitation id="6A3E4B797D1AE84E63F452E6FE9E4AD6" author="Brinkman, D. B." box="[128,327,657,681]" pageId="6" pageNumber="7" pagination="1645 - 1651" refId="ref12955" refString="Brinkman, D. B. 2001. New material of Dracochelys (Eucryptodira: Sinemydidae) from the Junggar Basin, Xinjiang, People's Republic of China. Canadian Journal of Earth Sciences 38: 1645 - 1651." type="journal article" year="2001">Brinkman (2001)</bibRefCitation>
for
<taxonomicName id="C9AF4D0B7D1AE84E62F152E6FDD44AD6" box="[389,525,657,681]" class="Reptilia" family="Macrobaenidae" genus="Dracochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E62F152E6FDD44AD6" box="[389,525,657,681]" italics="true" pageId="6" pageNumber="7">Dracochelys</emphasis>
</taxonomicName>
. They are relatively flat, show extensive costo-peripheral fontanelles, and have dorsally concave guttering as in
<taxonomicName id="C9AF4D0B7D1AE84E639F529CFEAA4B7C" box="[235,371,747,771]" class="Reptilia" family="Macrobaenidae" genus="Dracochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E639F529CFEAA4B7C" box="[235,371,747,771]" italics="true" pageId="6" pageNumber="7">Dracochelys</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0E1036887D1AE84E63EE537EFDCE4BAC" blockId="6.[128,654,211,1753]" pageId="6" pageNumber="7">
There is at least one other species of turtle at this locality, a chelid pleurodire much smaller than
<taxonomicName id="C9AF4D0B7D1AE84E63C85332FE9C4B22" authorityName="GAFFNEY &amp; RICH &amp; VICKERS-RICH &amp; CONSTANTINE &amp; VACCA &amp; KOOL" authorityYear="2007" box="[188,325,837,861]" class="Reptilia" family="Meiolaniidae" genus="Chubutemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3CDBEA9A7D1AE84E63C85332FE9C4B22" box="[188,325,837,861]" italics="true" pageId="6" pageNumber="7">Chubutemys</emphasis>
</taxonomicName>
. Most of these chelid bones differ from the cryptodire in being at least half the size and in having a very differently shaped shell with fused pelvis, no costo-peripheral fontanelles, and finely incised sulci.
</paragraph>
</subSubSection>
<subSubSection id="46B565037D1AE84E63EE53AEFC084988" pageId="6" pageNumber="7" type="discussion">
<paragraph id="0E1036887D1AE84E63EE53AEFC084988" blockId="6.[128,654,211,1753]" lastBlockId="6.[701,1228,211,503]" pageId="6" pageNumber="7">
DISCUSSION: Discovery of a definitive, skull-based eucryptodire in the Early Cretaceous of
<collectingCountry id="76B876187D1AE84E63935462FE804C52" box="[231,345,1045,1069]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
prompts reexamination of the published record, which has generally referred nonmarine turtles to the Pleurodira or
<taxonomicName id="C9AF4D0B7D1AE84E63DC5419FD534CF9" authority="(Broin and Fuente, 1993)" baseAuthorityName="Broin and Fuente" baseAuthorityYear="1993" box="[168,650,1134,1158]" class="Reptilia" family="Meiolaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="family">Meiolaniidae (Broin and Fuente, 1993)</taxonomicName>
. Much of this material is too fragmentary to reassess with confidence, but
<specimenCount id="18A9FD017D1AE84E629054DDFD574CBD" box="[484,654,1194,1218]" pageId="6" pageNumber="7" type="generic">two specimens</specimenCount>
previously identified as pleurodires are probably cryptodires. These were originally identified as grand
<taxonomicName id="C9AF4D0B7D1AE84E62595574FE494D64" box="[301,400,1283,1307]" class="Reptilia" family="Chelidae" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="7" phylum="Chordata" rank="family">Chelidae</taxonomicName>
primitif indetérmine´ (
<bibRefCitation id="6A3E4B797D1AE84E63F35556FE154D46" author="Broin, F. de &amp; M. de la Fuente" box="[135,460,1313,1337]" pageId="6" pageNumber="7" pagination="169 - 232" refId="ref13315" refString="Broin, F. de., and M. de la Fuente. 1993 b. Les tortues fossiles d'Argentine: synthese. Annales de Paleontologie 79 (3): 169 - 232." type="journal article" year="1993">Broin and Fuente, 1993b</bibRefCitation>
: fig. 5, 1819) from the Los Alamitos,
<collectingRegion id="CC6BF86A7D1AE84E62C85548FD9D4D28" box="[444,580,1343,1367]" country="Argentina" name="Rio Negro" pageId="6" pageNumber="7">Rio Negro</collectingRegion>
, Late Cretaceous. Both specimens are in the Museo Argentino de Ciencias Naturales and have been examined by the senior author (E.S.G.). MACN 907 was identified as a fifth cervical, but it is very similar to the undoubted eighth cervical of MPEF PV1940. Both are biconvex with a narrowed centrum and a wide, midway positioned transverse process, distinct from any described chelid. MACN 907 lacks the neural arch as well as the zygapophyses. The principal difference between them is that the anterior central articulation of MPEF PV1940 is slightly larger than the posterior one, whereas in MACN 907 they are closer in size. The other specimen, MACN 908, identified as an eighth cervical (Broin and Fuente, 1993), probably is an eighth; it agrees closely with MACN 907 but lacks transverse processes and a neural arch and only shows a biconvex centrum, so its identity must remain ambiguous. Nonetheless, it is very likely that MACN 908 is also a eucryptodire, along with MACN 907.
</paragraph>
</subSubSection>
</treatment>
</document>