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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro &amp; Hirowatari, Toshiya" docDate="2014" docId="EAE6CBB2E9D5FE4ECEE85D10CCEC9576" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus simoni Grouvelle 1892" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="33" updateTime="1668158244412" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshida, Takahiro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hirowatari, Toshiya</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>403</mods:number>
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<mods:start>15</mods:start>
<mods:end>45</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
<mods:identifier type="ZBK">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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<treatment ID-GBIF-Taxon="152052613" LSID="urn:lsid:plazi:treatment:EAE6CBB2E9D5FE4ECEE85D10CCEC9576" httpUri="http://treatment.plazi.org/id/EAE6CBB2E9D5FE4ECEE85D10CCEC9576" lastPageId="19" lastPageNumber="34" pageId="18" pageNumber="33">
<subSubSection pageId="18" pageNumber="33" type="nomenclature">
<paragraph pageId="18" pageNumber="33">
<taxonomicName LSID="http://species-id.net/wiki/Psammoecus_simoni" authority="Grouvelle, 1892" authorityName="Grouvelle" authorityYear="1892" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus simoni" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="simoni">Psammoecus simoni Grouvelle, 1892</taxonomicName>
Figs 2C, 10and 14
<normalizedToken originalValue="JL">J-L</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="reference_group">
<paragraph pageId="18" pageNumber="33">
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus simonis" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="simonis">Psammoecus simonis</taxonomicName>
[sic.] Grouvelle, 1892: 287. Type locality: Philippine; Type deposition: Museum National
<normalizedToken originalValue="dHistoire">d'Histoire</normalizedToken>
Naturelle, Paris. (misspelling)
</paragraph>
<paragraph pageId="18" pageNumber="33">
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus simoni" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="simoni">Psammoecus simoni</taxonomicName>
:
<bibRefCitation author="Grouvelle, A" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="26" pageNumber="41" pagination="315 - 495" title="Coleopteres de la region Indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae." volume="77" year="1908">Grouvelle 1908</bibRefCitation>
: 476, 488. -
<bibRefCitation author="Hetschko, A" journalOrPublisher="W. Junk, Berlin" pageId="27" pageNumber="42" pagination="1 - 90" title="Coleopterorum Catalogus Pars 109" year="1930">Hetschko 1930</bibRefCitation>
: 81. (catalogue) -
<bibRefCitation author="Pal, TK" journalOrPublisher="Records of the Zoological Survey of India" pageId="27" pageNumber="42" pagination="1 - 54" title="A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae)." volume="71" year="1985">Pal 1985</bibRefCitation>
: 31, fig. 11. -
<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano 2009</bibRefCitation>
: 63, 65, 82, fig. 5. -
<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano 2010</bibRefCitation>
: 11, 14. -
<bibRefCitation author="Karner, M" journalOrPublisher="European Journal of Taxonomy" pageId="27" pageNumber="42" pagination="1 - 31" title="A revision of African Psammoecus from the collection of the Musee royal de l'Afrique central." url="10.5852/ejt.2012.17" volume="17" year="2012">Karner 2012</bibRefCitation>
: 25, fig. 11.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="diagnosis">
<paragraph pageId="18" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="33">
This species is distinguished from other Japanese
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
species by the black elytra with yellow maculae and the posterior teeth on lateral margins of pronotum longer than those of anterior margins.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="description">
<paragraph pageId="18" pageNumber="33">Description.</paragraph>
<paragraph pageId="18" pageNumber="33">Body length. 2.24-2.56 mm (n=8).</paragraph>
<paragraph pageId="18" pageNumber="33">9th abdominal sternite (Fig. 14J). Strut comparatively short, cut at anterior 2/5, diverging gradually, branches comparatively long and wide. Lateral sclerites elongate, slightly curved inwards, apical sclerites pointed.</paragraph>
<paragraph pageId="18" pageNumber="33">Aedeagus (Fig. 14K, L). Parameres fused with phallobase, wide, short, posterior and lateral portions punctuated sparsely, densely punctuated triangular areas in anterior portions, a few long setae and short setae on posterior half, several short folds around beginning of divergence of parameres, a few lines on upper densely punctuated areas. Phallobase stout, broad in lateral aspect, anterior margin straight, a few subparallel lines on posterior dorsal portion. Penis comparatively broad, flattened around apex, apical margin rounded, punctuated sparsely on apex.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="specimens examined">
<paragraph pageId="18" pageNumber="33">Specimens examined.</paragraph>
<paragraph pageId="18" pageNumber="33">
JAPAN: [Okinawa Pref.] 1 male, 4 females &amp; 2 exs., Inamine, Nago City, Okinawa Island, 26
<normalizedToken originalValue="IX">-IX-</normalizedToken>
2012, T. Yoshida leg. (ELKU); 1 ex., Mt. Yarabu-dake, Ishigaki Island, 16
<normalizedToken originalValue="VI">-VI-</normalizedToken>
2002, T. Watanabe leg. (ELKU).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="distribution">
<paragraph pageId="18" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="18" pageNumber="33">JAPAN: Okinawa, Hateruma and Ishigaki Islands; Madagascar; India; Sri Lanka; Malaysia; Indonesia; Philippines.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="33" type="biological notes">
<paragraph pageId="18" pageNumber="33">Biological notes.</paragraph>
<paragraph pageId="18" pageNumber="33">
This species is found in dead leaves to which fungi are attached and occurs sympatrically with other Silvanid species such as
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
,
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus scitus" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="scitus">Psammoecus scitus</taxonomicName>
sp. n.,
<taxonomicName class="Insecta" family="Silvanidae" genus="Cryptamorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptamorpha desjardinsi" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="desjardinsi">Cryptamorpha desjardinsi</taxonomicName>
(
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1844) and
<taxonomicName class="Insecta" family="Silvanidae" genus="Monanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monanus concinnulus" order="Coleoptera" pageId="18" pageNumber="33" phylum="Arthropoda" rank="species" species="concinnulus">Monanus concinnulus</taxonomicName>
(Walker, 1858) (Fig. 15B).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="34" type="remarks">
<paragraph pageId="19" pageNumber="34">
<pageBreakToken pageId="19" pageNumber="34" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="19" pageNumber="34">
Pal (1980) and
<bibRefCitation author="Karner, M" journalOrPublisher="European Journal of Taxonomy" pageId="27" pageNumber="42" pagination="1 - 31" title="A revision of African Psammoecus from the collection of the Musee royal de l'Afrique central." url="10.5852/ejt.2012.17" volume="17" year="2012">Karner (2012)</bibRefCitation>
redescribed this species.
<bibRefCitation author="Grouvelle, A" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="26" pageNumber="41" pagination="285 - 288" title="Cucujides (in Voyage de M. E. Simon a l'ie de Luzon Philippines)." volume="61" year="1892">Grouvelle (1892)</bibRefCitation>
described it as
<normalizedToken originalValue="simonis">'simonis'</normalizedToken>
. However,
<bibRefCitation author="Grouvelle, A" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="26" pageNumber="41" pagination="315 - 495" title="Coleopteres de la region Indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae." volume="77" year="1908">Grouvelle (1908)</bibRefCitation>
referred to it with the specific name
<normalizedToken originalValue="simoni">'simoni'</normalizedToken>
. Pal (1980),
<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano (2010)</bibRefCitation>
and
<bibRefCitation author="Karner, M" journalOrPublisher="European Journal of Taxonomy" pageId="27" pageNumber="42" pagination="1 - 31" title="A revision of African Psammoecus from the collection of the Musee royal de l'Afrique central." url="10.5852/ejt.2012.17" volume="17" year="2012">Karner (2012)</bibRefCitation>
used the latter spelling, which we adopt in the present study.
</paragraph>
</subSubSection>
</treatment>
</document>