175 lines
17 KiB
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175 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.68454" ID-GBIF-Dataset="67beb2cf-e7a0-4f7f-ab57-48badd4ea194" ID-PMC="PMC8292276" ID-Pensoft-Pub="1313-2970-1048-109" ID-Pensoft-UUID="CC5A95B22B135366B9E84778E8FC3020" ID-PubMed="34316266" ID-ZooBank="FC201D049877458392C284CC80BB75A5" ModsDocID="1313-2970-1048-109" checkinTime="1626283239882" checkinUser="pensoft" docAuthor="Evsyukov, Aleksandr P., Golovatch, Sergei I. & Antic, Dragan Z." docDate="2021" docId="9B65CB30B6B85DFBBE9206280BF3A26A" docLanguage="en" docName="ZooKeys 1048: 109-143" docOrigin="ZooKeys 1048" docPubDate="2021-07-13" docSource="http://dx.doi.org/10.3897/zookeys.1048.68454" docTitle="Amblyiulus Silvestri 1896" docType="treatment" docVersion="4" id="CC5A95B22B135366B9E84778E8FC3020" lastPageNumber="109" masterDocId="CC5A95B22B135366B9E84778E8FC3020" masterDocTitle="The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae)" masterLastPageNumber="143" masterPageNumber="109" pageNumber="109" updateTime="1668150576225" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The millipede genera Amblyiulus Silvestri, 1896 and Syrioiulus Verhoeff, 1914 in the Caucasus, with notes on their distributions (Diplopoda, Julida, Julidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Evsyukov, Aleksandr P.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5521-7563</mods:nameIdentifier>
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<mods:affiliation>Don State Technical University, Department of Biology and General Pathology, Rostov-on-Don, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">aevsukov@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Antic, Dragan Z.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1231-4213</mods:nameIdentifier>
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<mods:affiliation>University of Belgrade, Faculty of Biology, Institute of Zoology, Belgrade, Serbia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1048</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>109</mods:start>
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<mods:end>143</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1048.68454</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.68454</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1048-109</mods:identifier>
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<mods:identifier type="ZooBank">FC201D049877458392C284CC80BB75A5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">CC5A95B22B135366B9E84778E8FC3020</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="183542980" LSID="urn:lsid:plazi:treatment:9B65CB30B6B85DFBBE9206280BF3A26A" httpUri="http://treatment.plazi.org/id/9B65CB30B6B85DFBBE9206280BF3A26A" lastPageNumber="109" pageId="0" pageNumber="109">
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<subSubSection pageId="0" pageNumber="109" type="nomenclature">
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<paragraph pageId="0" pageNumber="109">
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Genus
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<taxonomicName LSID="9B65CB30-B6B8-5DFB-BE92-06280BF3A26A" authority="Silvestri, 1896" authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">Amblyiulus Silvestri, 1896</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="109" type="type species">
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<paragraph pageId="0" pageNumber="109">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="109">
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<taxonomicName authorityName="Porat" authorityYear="1893" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Julus barroisi" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="barroisi">
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<emphasis italics="true" pageId="0" pageNumber="109">Julus barroisi</emphasis>
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</taxonomicName>
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Porat, 1893, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="109" type="diagnostic remarks">
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<paragraph pageId="0" pageNumber="109">Diagnostic remarks.</paragraph>
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<paragraph pageId="0" pageNumber="109">
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Here we follow
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<bibRefCitation author="Tabacaru, I" journalOrPublisher="Travaux de l'Institut de Speologie " Emile Racovitza "" pageId="0" pageNumber="109" pagination="67 - 80" refId="B47" refString="Tabacaru, I, 1978. Sur la systematique des Pachyiulinae. Description d'une nouvelle espece de Geopachyiulus. Travaux de l'Institut de Speologie "Emile Racovitza" 17: 67 - 80" title="Sur la systematique des Pachyiulinae. Description d'une nouvelle espece de Geopachyiulus." volume="17" year="1978">
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<normalizedToken originalValue="Tabacaru’s">Tabacaru's</normalizedToken>
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(1978)
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</bibRefCitation>
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opinion that the genera of the subfamily/tribe
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<taxonomicName authorityName="Verhoeff" authorityYear="1899" lsidName="" pageId="0" pageNumber="109" rank="subFamily" subFamily="Pachyiulinae">Pachyiulinae</taxonomicName>
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/-ini are best to be diagnosed using a complex of characters, both gonopodal and somatic. In the latest review of this tribe (
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<bibRefCitation author="Mauries, J-P" journalOrPublisher="Pedobiologia" pageId="0" pageNumber="109" refId="B36" refString="Mauries, J-P, 1982. Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle, Serie 4, 4(3-4): 433-444." title="Dolichoiulus tongiorgii (Strasser), Diplopode halophile nouveau pour la faune de France. Remarques sur la classification des Pachyiulini (Myriapoda, Diplopoda, Iulida). Bulletin du Museum national d'Histoire naturelle, Serie 4, 4 (3 - 4): 433 - 444." year="1982">
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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1982
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</bibRefCitation>
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), all genera are divided into three groups depending on the structure of the apical part of the opisthomere, viz., the presence/absence of a fovea and the presence/absence of a pseudoflagellum. He mistakenly assigned the genus
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<taxonomicName authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus</emphasis>
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</taxonomicName>
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to group 3 (along with many other genera like
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<taxonomicName class="Diplopoda" family="Julidae" genus="Dolichoiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dolichoiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Dolichoiulus</emphasis>
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</taxonomicName>
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), which have neither a fovea nor a pseudoflagellum. However, in accordance with our and earlier descriptions (e.g.,
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<bibRefCitation DOI="https://doi.org/10.1134/S0044513418070073" author="Golovatch, SI" journalOrPublisher="Zoologicheskii zhurnal" pageId="0" pageNumber="109" pagination="791 - 805" refId="B23" refString="Golovatch, SI, 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805, DOI: https://doi.org/10.1134/S0044513418070073" title="The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae)." url="https://doi.org/10.1134/S0044513418070073" volume="97" year="2018">Golovatch 2018</bibRefCitation>
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),
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<taxonomicName authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus</emphasis>
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</taxonomicName>
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has a fovea, however small, on the top of the solenomere. By the presence of a fovea and the absence of a pseudoflagellum, the genera
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<taxonomicName authorityName="Golovatch" authorityYear="1979" class="Diplopoda" family="Julidae" genus="Parapachyiulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parapachyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Parapachyiulus</emphasis>
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</taxonomicName>
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Golovatch, 1979 and
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<taxonomicName authorityName="Golovatch" authorityYear="1979" class="Diplopoda" family="Julidae" genus="Dangaraiulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dangaraiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Dangaraiulus</emphasis>
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</taxonomicName>
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Golovatch, 1979 also join this group (
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<bibRefCitation author="Golovatch, SI" journalOrPublisher="Zoologicheskii zhurnal" pageId="0" pageNumber="109" pagination="1313 - 1325" refId="B20" refString="Golovatch, SI, 1979. [The composition and zoogeographic relationship of the diplopod fauna of Middle Asia. Part 2]. Zoologicheskii zhurnal 58 (9): 1313 - 1325" title="[The composition and zoogeographic relationship of the diplopod fauna of Middle Asia. Part 2]." volume="58" year="1979">Golovatch 1979</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.1134/S0044513418070073" author="Golovatch, SI" journalOrPublisher="Zoologicheskii zhurnal" pageId="0" pageNumber="109" pagination="791 - 805" refId="B23" refString="Golovatch, SI, 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805, DOI: https://doi.org/10.1134/S0044513418070073" title="The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae)." url="https://doi.org/10.1134/S0044513418070073" volume="97" year="2018">2018</bibRefCitation>
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). According to a number of other characters, such as the presence of frontal setae, apicoventral lobes on the male mandibles, and the mesomeral process being as high as the opisthomere,
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<taxonomicName authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus</emphasis>
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</taxonomicName>
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belongs to
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<normalizedToken originalValue="Mauriès’">Mauries'</normalizedToken>
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subgroup 3aa, together with the genera
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<taxonomicName authorityName="Verhoeff" authorityYear="1914" class="Diplopoda" family="Julidae" genus="Syrioiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrioiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Syrioiulus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Verhoeff" authorityYear="1943" class="Diplopoda" family="Julidae" genus="Promeritoconus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Promeritoconus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Promeritoconus</emphasis>
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</taxonomicName>
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. However, it seems noteworthy that sometimes frontal setae can be absent, while male mandibular stipites can remain unmodified.
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</paragraph>
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<paragraph pageId="0" pageNumber="109">
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The promere in
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<taxonomicName authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus</emphasis>
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</taxonomicName>
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is narrowed in the basal third, in contrast to that in
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<taxonomicName authorityName="Verhoeff" authorityYear="1943" class="Diplopoda" family="Julidae" genus="Promeritoconus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Promeritoconus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Promeritoconus</emphasis>
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</taxonomicName>
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, which is narrowed apically; in the apical part it may have one or two denticles, but sometimes none. The head can be with or without frontal setae. The eyes are mostly absent. The opisthomere of the posterior gonopod is tripartite: a solenomere, an anterior process, and an anteromesal or lateral rod, vs. bipartite in
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<taxonomicName authorityName="Verhoeff" authorityYear="1914" class="Diplopoda" family="Julidae" genus="Syrioiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrioiulus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="109">Syrioiulus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="109" type="species included">
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<paragraph pageId="0" pageNumber="109">Species included.</paragraph>
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<paragraph pageId="0" pageNumber="109">
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<taxonomicName baseAuthorityName="Porat" baseAuthorityYear="1893" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus barroisi" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="barroisi">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus barroisi</emphasis>
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</taxonomicName>
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(Porat, 1893),
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<taxonomicName authorityName="Attems" authorityYear="1926" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus cedrophilus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="cedrophilus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus cedrophilus</emphasis>
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</taxonomicName>
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(Attems, 1910),
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<taxonomicName baseAuthorityName="Silvestri" baseAuthorityYear="1895" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus festae" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="festae">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus festae</emphasis>
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</taxonomicName>
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(Silvestri, 1895),
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<taxonomicName authorityName="Lohmander" authorityYear="1932" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus georgicus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="georgicus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus georgicus</emphasis>
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</taxonomicName>
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Lohmander, 1932,
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<taxonomicName authorityName="Evsyukov & Golovatch & Antić" authorityYear="2021" class="Diplopoda" family="Julidae" genus="Amblyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyiulus hirtus" order="Julida" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="hirtus">
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<emphasis italics="true" pageId="0" pageNumber="109">Amblyiulus hirtus</emphasis>
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</taxonomicName>
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sp. nov., and possibly several others, but their identity requires verification (
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<bibRefCitation DOI="https://doi.org/10.1134/S0044513418070073" author="Golovatch, SI" journalOrPublisher="Zoologicheskii zhurnal" pageId="0" pageNumber="109" pagination="791 - 805" refId="B23" refString="Golovatch, SI, 2018. The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae). Zoologicheskii zhurnal 97 (7): 791 - 805, DOI: https://doi.org/10.1134/S0044513418070073" title="The millipede subfamily Pachyiulinae in Israel, with the description of a new species (Diplopoda, Julida, Julidae)." url="https://doi.org/10.1134/S0044513418070073" volume="97" year="2018">Golovatch 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |