treatments-xml/data/91/45/08/91450849DDF95F46B94A9DF53F701C3E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.975.55456" ID-GBIF-Dataset="bfcb4e07-aaea-419e-8bb5-08851e9f27dd" ID-PMC="PMC7572521" ID-Pensoft-Pub="1313-2970-975-11" ID-Pensoft-UUID="FE5DCB656614590DB4C35092D89F3F3D" ID-PubMed="33117063" ID-ZooBank="785EDF225A3242178C9B6433E0AD199F" ModsDocID="1313-2970-975-11" checkinTime="1602606874639" checkinUser="pensoft" docAuthor="Hendrich, Lars, Watts, Chris H. S. &amp; Balke, Michael" docDate="2020" docId="91450849DDF95F46B94A9DF53F701C3E" docLanguage="en" docName="ZooKeys 975: 11-49" docOrigin="ZooKeys 975" docSource="http://dx.doi.org/10.3897/zookeys.975.55456" docTitle="Gibbidessus davidi Hendrich &amp; Watts &amp; Balke 2020, sp. nov." docType="treatment" docUuid="4D235E7C-F517-4C1D-A3C7-0C87BDA34515" docUuidSource="ZooBank" docVersion="4" id="FE5DCB656614590DB4C35092D89F3F3D" lastPageNumber="11" masterDocId="FE5DCB656614590DB4C35092D89F3F3D" masterDocTitle="The &quot; minute diving beetles &quot; of southern Australia - taxonomic revision of Gibbidessus Watts, 1978, with description of six new species (Coleoptera, Dytiscidae, Bidessini)" masterLastPageNumber="49" masterPageNumber="11" pageNumber="11" updateTime="1668169601453" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The &quot; minute diving beetles &quot; of southern Australia - taxonomic revision of Gibbidessus Watts, 1978, with description of six new species (Coleoptera, Dytiscidae, Bidessini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>SNSB - Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D - 81247 Muenchen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8366-0749</mods:nameIdentifier>
<mods:nameIdentifier type="email">hendrich1@aol.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watts, Chris H. S.</mods:namePart>
<mods:affiliation>South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:affiliation>SNSB - Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D - 81247 Muenchen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>975</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>49</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.975.55456</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.975.55456</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-975-11</mods:identifier>
<mods:identifier type="ZooBank">785EDF225A3242178C9B6433E0AD199F</mods:identifier>
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<treatment ID-GBIF-Taxon="168338750" LSID="urn:lsid:zoobank.org:act:4D235E7C-F517-4C1D-A3C7-0C87BDA34515" httpUri="http://treatment.plazi.org/id/91450849DDF95F46B94A9DF53F701C3E" lastPageNumber="11" pageId="0" pageNumber="11">
<subSubSection pageId="0" pageNumber="11" type="nomenclature">
<paragraph pageId="0" pageNumber="11">
<taxonomicName LSID="91450849-DDF9-5F46-B94A-9DF53F701C3E" authority="Hendrich &amp; Watts &amp; Balke, 2020" authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus davidi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="davidi" status="sp. nov.">Gibbidessus davidi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="11">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">Figs 3</figureCitation>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">, 4</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F4" captionText="Figure 12. Head of G. davidi sp. nov. A with cervical line (red arrow), and head of G. pederzanii sp. nov. B without cervical line but rather a few punctures instead (red arrow)." figureDoi="10.3897/zookeys.975.55456.figure12" httpUri="https://binary.pensoft.net/fig/462663" pageId="0" pageNumber="11">, 12</figureCitation>
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">, 16</figureCitation>
<figureCitation captionStart="Figures 22, 23" captionStartId="F10" captionText="Figures 22, 23. Distribution of 22 Gibbidessus chipi (black dots), G. drikdrikensis sp. nov. (yellow square), G. kangarooensis sp. nov. (red square) 23 G. atomus sp. nov. (red square), G. davidi sp. nov. (yellow square)." figureDoi="10.3897/zookeys.975.55456.figures22-23" httpUri="https://binary.pensoft.net/fig/462669" pageId="0" pageNumber="11">, 23</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F13" captionText="Figure 27. Habitat of Gibbidessus atomus sp. nov. and Gibbidessus davidi sp. nov. A Seasonally flooded Melaleuca sedge swamp B small and shallow heathland pool along Windy Harbour Road, south of Northcliffe, south-western Australia." figureDoi="10.3897/zookeys.975.55456.figure27" httpUri="https://binary.pensoft.net/fig/462672" pageId="0" pageNumber="11">, 27</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F14" captionText="Figure 28. Habitat of Gibbidessus davidi sp. nov. and Gibbidessus rottnestensis sp. nov. A Shallow peaty pool with mats of floating grasses along Beeliar Swamps in Perth B same spot and habitat details with collecting methods." figureDoi="10.3897/zookeys.975.55456.figure28" httpUri="https://binary.pensoft.net/fig/462673" pageId="0" pageNumber="11">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type locality">
<paragraph pageId="0" pageNumber="11">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="11">
Western Australia, Perth, suburb Success, Beeliar Regional Park, shallow peaty puddle [
<geoCoordinate degrees="32" direction="south" minutes="8" orientation="latitude" precision="15" seconds="4" value="-32.134445">32°8'4S</geoCoordinate>
, 
<geoCoordinate degrees="115" direction="east" minutes="50" orientation="longitude" precision="15" seconds="22" value="115.83945">115°50'22E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type material">
<paragraph pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
,
</emphasis>
male: &quot;Australia, WA, Perth, Success, Beeliar RP, shallow peaty puddle 
<geoCoordinate degrees="32" direction="south" minutes="8" orientation="latitude" precision="1" seconds="4.97" value="-32.134716">32°8'4.97&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="115" direction="east" minutes="50" orientation="longitude" precision="1" seconds="22.78" value="115.83966">115°50'22.78&quot;E</geoCoordinate>
21.-31.10.2015 L. Hendrich (WA 1/15)&quot;, &quot;Holotype 
<taxonomicName authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus davidi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus davidi</emphasis>
</taxonomicName>
Hendrich, Watts &amp; Balke des. 2020&quot; (WAM) [red printed label]. 
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
(370 exs.)
</emphasis>
. 354 specimens with same data as holotype (ANIC, CGC, CLH, SAMA, WAM, ZSM); 4 exs, &quot;Australia: SW WA, 
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
NP, 11 km S, Northcliffe, 77m, 4.I.2007, 34.44.048S, 116.05.354E [
<geoCoordinate degrees="34" direction="south" minutes="44" orientation="latitude" precision="15" seconds="0" value="-34.733334">34°44'0S</geoCoordinate>
, 
<geoCoordinate degrees="116" direction="east" minutes="5" orientation="longitude" precision="15" seconds="13" value="116.086945">116°5'13E</geoCoordinate>
], L. &amp; E. Hendrich (WA 162)&quot; (ZSM); 10 exs., &quot;Australia, WA, Albany Hwy, Muir Lakes Nature Reserve, SW part of Byenup Lagoon, 4.&amp; 5.1.2000, 
<geoCoordinate degrees="34" direction="south" minutes="30" orientation="latitude" precision="15" seconds="4" value="-34.50111">34°30'4S</geoCoordinate>
, 
<geoCoordinate degrees="116" direction="east" minutes="44" orientation="longitude" precision="15" seconds="19" value="116.73861">116°44'19E</geoCoordinate>
, Hendrich leg. (loc. WA 11/157)&quot; (CLH, ZSM); 1 ex., &quot;Australia, WA, Barlee Brook Dickson Road (A) [
<geoCoordinate degrees="34" direction="south" minutes="12" orientation="latitude" precision="15" seconds="17" value="-34.204723">34°12'17S</geoCoordinate>
, 
<geoCoordinate degrees="115" direction="east" minutes="46" orientation="longitude" precision="15" seconds="18" value="115.77167">115°46'18E</geoCoordinate>
], DON03, 21/10/2005, South West Forest Monitoring&quot; (DPAW); 1 ex., &quot;Australia, WA, Fish Creek (A) [
<geoCoordinate degrees="34" direction="south" minutes="37" orientation="latitude" precision="15" seconds="29" value="-34.62472">34°37'29S</geoCoordinate>
, 
<geoCoordinate degrees="116" direction="east" minutes="26" orientation="longitude" precision="15" seconds="11" value="116.436386">116°26'11E</geoCoordinate>
], SHA22, 17/10/2010, South West Forest Monitoring&quot; (DPAW); 4 exs., &quot;Australia, WA, RVDLE03 Riverdale Wetland [
<geoCoordinate degrees="32" direction="south" minutes="59" orientation="latitude" precision="15" seconds="23" value="-32.989723">32°59'23S</geoCoordinate>
, 
<geoCoordinate degrees="115" direction="east" minutes="47" orientation="longitude" precision="15" seconds="23" value="115.78972">115°47'23E</geoCoordinate>
], 23/09/2008, South West Catchment Council Mon.&quot; (DPAW). All paratypes with red printed paratype labels.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="diagnosis">
<paragraph pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="11">
Small species which externally is characterised by a wide rounded body, shiny non-microreticulate dorsal surface, vague testaceous markings on elytra, and without habitus disruption between pronotum and elytron. Dorsoventrally rather domed. Cervical line present (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="description">
<paragraph pageId="0" pageNumber="11">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="11">Holotype: TL = 1.45 mm, TL-H = 1.35 mm, MW = 0.83 mm. Paratypes: TL = 1.35-1.5 mm, TL-H = 1.15-1.4 mm, MW = 0.8-0.9 mm.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Head</emphasis>
:
</emphasis>
Ferruginous, around eyes almost black. Cervical line present (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F4" captionText="Figure 12. Head of G. davidi sp. nov. A with cervical line (red arrow), and head of G. pederzanii sp. nov. B without cervical line but rather a few punctures instead (red arrow)." figureDoi="10.3897/zookeys.975.55456.figure12" httpUri="https://binary.pensoft.net/fig/462663" pageId="0" pageNumber="11">12A</figureCitation>
). Strongly and coarsely punctate, rather shiny, microreticulation present. Punctures weak anteriorly and strongly posteriorly between eyes. Antennae relatively short, stout. Antennomeres 1-8 ferruginous, 9-11 darkened anteriorly.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Pronotum</emphasis>
:
</emphasis>
Ferruginous, anterior and posterior margins darker. Disc of pronotum somewhat darkened, broadest at posterior corners. Punctation of pronotum very weak, almost evenly distributed, shiny and microsculpture absent. Sides of pronotum margined and almost evenly rounded. Angle between pronotum and elytra less pronounced, basal pronotal plicae present. Striae moderately defined, almost 1/2 length of pronotum, strongly incurved.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Elytra</emphasis>
:
</emphasis>
Dark brown with vague basolateral area ferruginous (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">3</figureCitation>
). Coarsely and densely punctate, shiny, microsculpture absent. Striae deeply impressed, straight but shorter than basal pronotal striae.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Ventral side</emphasis>
:
</emphasis>
Ferruginous. Prothorax and abdomen paler than other parts. Metacoxae and metaventrite covered with larger punctures, surface shiny, without microreticulation. Abdominal ventrites with dense and finer punctures, shiny, microreticulation absent. Metacoxal lines almost straight, anteriorly not divergent. Epipleuron ferruginous, with few coarse punctures, shiny, lacking microsculpture. Legs ferruginous with meta-/mesotibia and meta-/mesotarsi set in black.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Male</emphasis>
.
</emphasis>
Smaller and more elongate than female (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">4</figureCitation>
). Median lobe of aedeagus as in Fig. 
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16A, B</figureCitation>
. Shape of median lobe in lateral view, straight and fairly uniform, in ventral view broad, with a thorn on each side, and rounded at apex. Parameres bi-segmented and elongated with few setae at apex (Fig. 
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="affinities">
<paragraph pageId="0" pageNumber="11">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="11">
This species is similar to 
<taxonomicName lsidName="G. atomus" pageId="0" pageNumber="11" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">G. atomus</emphasis>
</taxonomicName>
sp. nov. but readily separated by its larger size, the different colour pattern and the form of the median lobe and parameres (Figs 
<figureCitation captionStart="Figures 14, 15" captionStartId="F6" captionText="Figures 14, 15. 14 Gibbidessus atomus sp. nov. 15 G. chipi A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures14-15" httpUri="https://binary.pensoft.net/fig/462665" pageId="0" pageNumber="11">14</figureCitation>
, 
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16</figureCitation>
). From 
<taxonomicName lsidName="G. pederzanii" pageId="0" pageNumber="11" rank="species" species="pederzanii">
<emphasis italics="true" pageId="0" pageNumber="11">G. pederzanii</emphasis>
</taxonomicName>
sp. nov. it can be distinguished by the less roundish body and the form of the median lobe and parameres (Figs 
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16</figureCitation>
, 
<figureCitation captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="etymology">
<paragraph pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="11">The beetle is named after the son of the first author, David Hendrich. The specific epithet is a substantive in the genitive case.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="distribution">
<paragraph pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="11">
South-western Australia. From Perth in the north to 
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
National Park in the south (Fig. 
<figureCitation captionStart="Figures 22, 23" captionStartId="F10" captionText="Figures 22, 23. Distribution of 22 Gibbidessus chipi (black dots), G. drikdrikensis sp. nov. (yellow square), G. kangarooensis sp. nov. (red square) 23 G. atomus sp. nov. (red square), G. davidi sp. nov. (yellow square)." figureDoi="10.3897/zookeys.975.55456.figures22-23" httpUri="https://binary.pensoft.net/fig/462669" pageId="0" pageNumber="11">23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="habitat">
<paragraph pageId="0" pageNumber="11">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="11">
In the Northcliffe area most specimens were obtained from an exposed, shallow and small roundish puddle, without any vegetation, except some algae. The other specimens were collected in a half-shaded pool in a 
<taxonomicName class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Melaleuca" order="Agaricales" pageId="0" pageNumber="11" phylum="Basidiomycota" rank="genus">Melaleuca</taxonomicName>
blackwater swamp, with few clumps of 
<taxonomicName class="Liliopsida" family="Juncaceae" genus="Juncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juncus" order="Poales" pageId="0" pageNumber="11" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Juncus</emphasis>
</taxonomicName>
spp. and extensive beds of macrophytes; depth up to 20 cm; bottom consisted of sedge-filled peat (pH 5.5), twigs and rotten leaves (Figs 
<figureCitation captionStart="Figure 27" captionStartId="F13" captionText="Figure 27. Habitat of Gibbidessus atomus sp. nov. and Gibbidessus davidi sp. nov. A Seasonally flooded Melaleuca sedge swamp B small and shallow heathland pool along Windy Harbour Road, south of Northcliffe, south-western Australia." figureDoi="10.3897/zookeys.975.55456.figure27" httpUri="https://binary.pensoft.net/fig/462672" pageId="0" pageNumber="11">27</figureCitation>
, 
<figureCitation captionStart="Figure 28" captionStartId="F14" captionText="Figure 28. Habitat of Gibbidessus davidi sp. nov. and Gibbidessus rottnestensis sp. nov. A Shallow peaty pool with mats of floating grasses along Beeliar Swamps in Perth B same spot and habitat details with collecting methods." figureDoi="10.3897/zookeys.975.55456.figure28" httpUri="https://binary.pensoft.net/fig/462673" pageId="0" pageNumber="11">28</figureCitation>
). The whole area south of Northcliffe is seasonally flooded with some permanent central water bodies in summer. In the 
<normalizedToken originalValue="D´Entrecasteaux">D'Entrecasteaux</normalizedToken>
NP the species is syntopic with 
<taxonomicName lsidName="G. atomus" pageId="0" pageNumber="11" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">G. atomus</emphasis>
</taxonomicName>
sp. nov., and around Perth in the Beeliar Regional Park with 
<taxonomicName lsidName="G. rottnestensis" pageId="0" pageNumber="11" rank="species" species="rottnestensis">
<emphasis italics="true" pageId="0" pageNumber="11">G. rottnestensis</emphasis>
</taxonomicName>
sp. nov. At the Riverdale Wetland Reserve 
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
sp. nov. was syntopic with 
<taxonomicName lsidName="G. atomus" pageId="0" pageNumber="11" rank="species" species="atomus">
<emphasis italics="true" pageId="0" pageNumber="11">G. atomus</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="G. rottnestensis" pageId="0" pageNumber="11" rank="species" species="rottnestensis">
<emphasis italics="true" pageId="0" pageNumber="11">G. rottnestensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph pageId="0" pageNumber="11">
Apart from the 
<taxonomicName authorityName="Watts" authorityYear="1978" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus</emphasis>
</taxonomicName>
, the water beetle coenosis at Northcliffe included the following species: 
<taxonomicName family="Dytiscidae" lsidName="" pageId="0" pageNumber="11" rank="family">Dytiscidae</taxonomicName>
: 
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Limbodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limbodessus inornatus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis italics="true" pageId="0" pageNumber="11">Limbodessus inornatus</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Watts" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Antiporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antiporus hollingsworthi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="hollingsworthi">
<emphasis italics="true" pageId="0" pageNumber="11">Antiporus hollingsworthi</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="A. mcraeae" pageId="0" pageNumber="11" rank="species" species="mcraeae">
<emphasis italics="true" pageId="0" pageNumber="11">A. mcraeae</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Brancuporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brancuporus gottwaldi" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="gottwaldi">
<emphasis italics="true" pageId="0" pageNumber="11">Brancuporus gottwaldi</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Lea" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Sternopriscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sternopriscus minimus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="minimus">
<emphasis italics="true" pageId="0" pageNumber="11">Sternopriscus minimus</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="S. eikei" pageId="0" pageNumber="11" rank="species" species="eikei">
<emphasis italics="true" pageId="0" pageNumber="11">S. eikei</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ater" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="11">Exocelina ater</emphasis>
</taxonomicName>
; 
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Hydrophilidae" lsidName="" pageId="0" pageNumber="11" rank="family">Hydrophilidae</taxonomicName>
: 
<taxonomicName baseAuthorityName="Blackburn" baseAuthorityYear="1895" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus eyrensis" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="eyrensis">
<emphasis italics="true" pageId="0" pageNumber="11">Enochrus eyrensis</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Broun" baseAuthorityYear="1880" class="Insecta" family="Hydrophilidae" genus="Limnoxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limnoxenus zealandicus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="zealandicus">
<emphasis italics="true" pageId="0" pageNumber="11">Limnoxenus zealandicus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Paracymus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracymus pygmaeus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis italics="true" pageId="0" pageNumber="11">Paracymus pygmaeus</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Hendrich, L" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="11" pagination="299 - 308" refId="B6" refString="Hendrich, L, 2001a. A new species of Antiporus Sharp, 1882 from peatland swamps of south-western Australia (Coleoptera: Dytiscidae). Linzer biologische Beitraege 33 (1): 299 - 308" title="A new species of Antiporus Sharp, 1882 from peatland swamps of south-western Australia (Coleoptera: Dytiscidae)." volume="33" year="2001 a">Hendrich 2001a</bibRefCitation>
). In the Beeliar Park the two 
<taxonomicName authorityName="Watts" authorityYear="1978" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus</emphasis>
</taxonomicName>
species share their habitat with 
<taxonomicName baseAuthorityName="Sharp" baseAuthorityYear="1882" class="Insecta" family="Dytiscidae" genus="Limbodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limbodessus inornatus" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis italics="true" pageId="0" pageNumber="11">Limbodessus inornatus</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Clark" baseAuthorityYear="1862" class="Insecta" family="Dytiscidae" genus="Paroster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroster insculptilis" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="insculptilis">
<emphasis italics="true" pageId="0" pageNumber="11">Paroster insculptilis</emphasis>
</taxonomicName>
(Clark, 1862), 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ater" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="11">Exocelina ater</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="W. S. MacLeay" baseAuthorityYear="1825" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus suturalis" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis italics="true" pageId="0" pageNumber="11">Rhantus suturalis</emphasis>
</taxonomicName>
(Macleay, 1825) and 
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus simulans" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="simulans">
<emphasis italics="true" pageId="0" pageNumber="11">Rhantus simulans</emphasis>
</taxonomicName>
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
, 1908.
</paragraph>
</subSubSection>
</treatment>
</document>