treatments-xml/data/03/B6/E1/03B6E134153B0243259F8AB8FCA19537.xml
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<document id="3370E1627C11D5E61E3D283BDF7DCF49" ID-CLB-Dataset="24234" ID-DOI="10.11646/zootaxa.4729.1.3" ID-GBIF-Dataset="22815f4d-7c51-45d1-b6b3-8216ea02e14c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3629007" ID-ZooBank="BBFF38A8-8BD6-4A70-A2DA-0804D5DE2C28" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580180119895" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2020" docId="03B6E134153B0243259F8AB8FCA19537" docLanguage="en" docName="zootaxa.4729.1.3.pdf" docOrigin="Zootaxa 4729 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus tianbaoshanensis Mikhaljova 2020, new species" docType="treatment" docVersion="7" lastPageNumber="57" masterDocId="FF8F994C1532024925088F48FFFB933C" masterDocTitle="New data on the fauna of China, part I: the taxonomy of the millipede family Julidae (Diplopoda)" masterLastPageNumber="66" masterPageNumber="47" pageNumber="56" updateTime="1698779498477" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7C6A9908E9229FFD093653F0D9BF7359">New data on the fauna of China, part I: the taxonomy of the millipede family Julidae (Diplopoda)</mods:title>
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<mods:namePart id="68ACEE2240B8F20F9348B783AA8BA21E">Mikhaljova, Elena V.</mods:namePart>
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<mods:date id="D9B974A009DBBB87E3D8AAA3B44E04A2">2020</mods:date>
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<treatment id="03B6E134153B0243259F8AB8FCA19537" ID-DOI="http://doi.org/10.5281/zenodo.5918709" ID-GBIF-Taxon="161840490" ID-Zenodo-Dep="5918709" LSID="urn:lsid:plazi:treatment:03B6E134153B0243259F8AB8FCA19537" httpUri="http://treatment.plazi.org/id/03B6E134153B0243259F8AB8FCA19537" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
<subSubSection id="C30503A9153B0240259F8AB8FD339536" box="[151,712,1520,1547]" pageId="9" pageNumber="56" type="nomenclature">
<paragraph id="8BA05022153B0240259F8AB8FD339536" blockId="9.[151,712,1520,1583]" box="[151,712,1520,1547]" pageId="9" pageNumber="56">
<heading id="D0E8E74E153B0240259F8AB8FD339536" bold="true" box="[151,712,1520,1547]" fontSize="11" level="1" pageId="9" pageNumber="56" reason="1">
<emphasis id="B96B8C30153B0240259F8AB8FD339536" bold="true" box="[151,712,1520,1547]" pageId="9" pageNumber="56">
<taxonomicName id="4C1F2BA1153B0240259F8AB8FDCA9537" ID-CoL="1e8a37cd-0f00-4b14-bbbe-c15ed2d79567" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[151,561,1520,1547]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="tianbaoshanensis" status="sp. nov.">
<emphasis id="B96B8C30153B0240259F8AB8FDCA9537" bold="true" box="[151,561,1520,1547]" italics="true" pageId="9" pageNumber="56">Nepalmatoiulus tianbaoshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A258314B153B024027308AB9FD339536" box="[568,712,1520,1547]" pageId="9" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30503A9153B0240259F895CFEEE9513" box="[151,277,1556,1583]" pageId="9" pageNumber="56" type="description">
<paragraph id="8BA05022153B0240259F895CFEEE9513" blockId="9.[151,712,1520,1583]" box="[151,277,1556,1583]" pageId="9" pageNumber="56">
<figureCitation id="13244CA7153B0240259F895CFEEE9513" box="[151,277,1556,1583]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="9" pageNumber="56">Figs 3845</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30503A9153B0240259F8914FA7C943A" pageId="9" pageNumber="56" type="materials_examined">
<paragraph id="8BA05022153B0240259F8914FA7C943A" blockId="9.[151,1437,1628,2015]" pageId="9" pageNumber="56">
<emphasis id="B96B8C30153B0240259F8914FE81954B" bold="true" box="[151,378,1628,1655]" pageId="9" pageNumber="56">Material examined</emphasis>
:
<materialsCitation id="3B775A7F153B024024828914FEDD9582" ID-GBIF-Occurrence="2560337980" collectingDate="2015-05-20" collectionCode="ZMUM" collectorName="Shangrila &amp; Habaxue Shan &amp; NE Xiaozhongdian &amp; I. Belousov &amp; Belousov, I &amp; Kabak, G." country="China" elevation="4015" latitude="27.6" location="Tianbaoshan" longLatPrecision="20" longitude="99.90139" pageId="9" pageNumber="56" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B96B8C30153B024024828914FE0A954A" box="[394,497,1628,1654]" italics="true" pageId="9" pageNumber="56">
<typeStatus id="54A4EE80153B024024828914FE0A954A" box="[394,497,1628,1654]" pageId="9" pageNumber="56" type="holotype">Holotype</typeStatus>
</emphasis>
: male (
<collectionCode id="ED0EC8E7153B0240274F8915FD5F954B" box="[583,676,1629,1655]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="56">ZMUM</collectionCode>
),
<collectingCountry id="F30810B2153B024027B08914FD05954A" box="[696,766,1628,1654]" name="China" pageId="9" pageNumber="56">China</collectingCountry>
,
<collectingRegion id="49DB9EC0153B024026008915FC37954A" box="[776,972,1628,1655]" country="China" name="Yunnan" pageId="9" pageNumber="56">Yunnan Province</collectingRegion>
,
<location id="8EC006F9153B024026D08914FB9D954A" LSID="urn:lsid:plazi:treatment:03B6E134153B0243259F8AB8FCA19537:8EC006F9153B024026D08914FB9D954A" box="[984,1126,1628,1654]" country="China" latitude="27.6" longLatPrecision="20" longitude="99.90139" name="Tianbaoshan" pageId="9" pageNumber="56" stateProvince="Yunnan">Tianbaoshan</location>
between
<collectorName id="26EA35F4153B024021DB8914FAC5954A" box="[1235,1342,1628,1654]" pageId="9" pageNumber="56">Shangrila</collectorName>
&amp;
<collectorName id="26EA35F4153B0240206C8914FEF695A6" pageId="9" pageNumber="56">Habaxue Shan</collectorName>
,
<collectorName id="26EA35F4153B0240241F89C9FE0B95A6" box="[279,496,1664,1691]" pageId="9" pageNumber="56">NE Xiaozhongdian</collectorName>
,
<geoCoordinate id="EE2B36E5153B024024F289C8FD7F95A7" box="[506,644,1664,1691]" degrees="27" direction="north" minutes="36" orientation="latitude" pageId="9" pageNumber="56" precision="15" seconds="0" value="27.6">27°36´0´´ N</geoCoordinate>
,
<geoCoordinate id="EE2B36E5153B0240278689C8FCEF95A7" box="[654,788,1664,1691]" degrees="99" direction="east" minutes="54" orientation="longitude" pageId="9" pageNumber="56" precision="15" seconds="5" value="99.90139">99°54´5´´ E</geoCoordinate>
, H =
<elevation id="0032B711153B0240264789C8FC3695A6" box="[847,973,1664,1691]" metricMagnitude="3" metricUnit="m" metricValue="4.015" pageId="9" pageNumber="56" unit="m" value="4015.0">
<quantity id="4CE7FDC7153B0240264789C8FC5895A7" box="[847,931,1664,1691]" metricMagnitude="3" metricUnit="m" metricValue="4.015" pageId="9" pageNumber="56" unit="m" value="4015.0">4015 m</quantity>
asl
</elevation>
.,
<date id="FFA176E2153B024026D489C9FB8995A7" box="[988,1138,1664,1691]" pageId="9" pageNumber="56" value="2015-05-20">
<collectingDate id="EFE58F0A153B024026D489C9FB8995A7" box="[988,1138,1664,1691]" pageId="9" pageNumber="56" value="2015-05-20">20 May 2015</collectingDate>
</date>
, leg.
<collectorName id="26EA35F4153B024021A389C9FAD795A6" box="[1195,1324,1664,1691]" pageId="9" pageNumber="56">I. Belousov</collectorName>
, I.
<collectorName id="26EA35F4153B0240204489C8FF489583" pageId="9" pageNumber="56">Kabak, G.</collectorName>
Davidian.
</materialsCitation>
<materialsCitation id="3B775A7F153B0240242389EDFE8195DE" ID-GBIF-Occurrence="2560337990" collectingDate="2015-05-20" collectionCode="ZMUM" collectorName="I. Belousov &amp; Belousov, I &amp; Kabak, G" country="China" elevation="4015" latitude="27.6" location="Tianbaoshan" longLatPrecision="20" longitude="99.90139" pageId="9" pageNumber="56" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B96B8C30153B0240242389EDFE669582" box="[299,413,1701,1726]" italics="true" pageId="9" pageNumber="56">
<typeStatus id="54A4EE80153B0240242389EDFE669582" box="[299,413,1701,1726]" pageId="9" pageNumber="56" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D199BAB153B024024A289EDFDF99582" box="[426,514,1700,1727]" pageId="9" pageNumber="56" type="male">2 males</specimenCount>
,
<specimenCount id="9D199BAB153B0240270489EDFD8C9582" box="[524,631,1700,1727]" pageId="9" pageNumber="56" type="female">4 females</specimenCount>
(
<collectionCode id="ED0EC8E7153B0240278C89EDFD1A9583" box="[644,737,1701,1727]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="56">ZMUM</collectionCode>
), same locality as in
<typeStatus id="54A4EE80153B024026CC89ECFBD29582" box="[964,1065,1700,1726]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
,
<date id="FFA176E2153B0240213B89EDFB329583" box="[1075,1225,1700,1727]" pageId="9" pageNumber="56" value="2015-05-20">
<collectingDate id="EFE58F0A153B0240213B89EDFB329583" box="[1075,1225,1700,1727]" pageId="9" pageNumber="56" value="2015-05-20">20 May 2015</collectingDate>
</date>
, leg.
<collectorName id="26EA35F4153B0240200989EDFA799582" box="[1281,1410,1700,1727]" pageId="9" pageNumber="56">I. Belousov</collectorName>
, I.
<collectorName id="26EA35F4153B0240259F8980FEFE95DF" box="[151,261,1736,1763]" pageId="9" pageNumber="56">Kabak, G</collectorName>
. Davidian
</materialsCitation>
;
<materialsCitation id="3B775A7F153B0240248D8981FA7C943A" ID-GBIF-Occurrence="2560337988" collectingDate="2015-05-22" collectionCode="ZMUM" collectorName="Shangrila &amp; Habaxue Shan &amp; SE Niguge &amp; I. Belousov &amp; Belousov, I &amp; Kabak, G." country="China" elevation="4285" latitude="27.610556" location="Tianbaoshan" longLatPrecision="20" longitude="99.93694" pageId="9" pageNumber="56" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D199BAB153B0240248D8981FE2B95DE" box="[389,464,1736,1763]" pageId="9" pageNumber="56" type="male">1 male</specimenCount>
(
<collectionCode id="ED0EC8E7153B024024D78981FDC795DF" box="[479,572,1737,1763]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="9" pageNumber="56">ZMUM</collectionCode>
),
<collectingCountry id="F30810B2153B024027478980FD6E95DE" box="[591,661,1736,1762]" name="China" pageId="9" pageNumber="56">China</collectingCountry>
,
<collectingRegion id="49DB9EC0153B024027968981FC9A95DE" box="[670,865,1736,1763]" country="China" name="Yunnan" pageId="9" pageNumber="56">Yunnan Province</collectingRegion>
,
<location id="8EC006F9153B024026638980FC0295DE" LSID="urn:lsid:plazi:treatment:03B6E134153B0243259F8AB8FCA19537:8EC006F9153B024026638980FC0295DE" box="[875,1017,1736,1762]" country="China" latitude="27.610556" longLatPrecision="20" longitude="99.93694" name="Tianbaoshan" pageId="9" pageNumber="56" stateProvince="Yunnan">Tianbaoshan</location>
between
<collectorName id="26EA35F4153B0240216C8980FB3495DE" box="[1124,1231,1736,1762]" pageId="9" pageNumber="56">Shangrila</collectorName>
&amp;
<collectorName id="26EA35F4153B024021FB8980FA6395DE" box="[1267,1432,1736,1762]" pageId="9" pageNumber="56">Habaxue Shan</collectorName>
,
<collectorName id="26EA35F4153B0240259F89A5FEEF943A" box="[151,276,1772,1799]" pageId="9" pageNumber="56">SE Niguge</collectorName>
,
<geoCoordinate id="EE2B36E5153B0240241789A4FE43943B" box="[287,440,1772,1799]" degrees="27" direction="north" minutes="36" orientation="latitude" pageId="9" pageNumber="56" precision="15" seconds="38" value="27.610556">27°36´38´´ N</geoCoordinate>
,
<geoCoordinate id="EE2B36E5153B024024CC89A4FDA2943B" box="[452,601,1772,1799]" degrees="99" direction="east" minutes="56" orientation="longitude" pageId="9" pageNumber="56" precision="15" seconds="13" value="99.93694">99°56´13´´ E</geoCoordinate>
, H =
<elevation id="0032B711153B0240279F89A4FCEC943A" box="[663,791,1772,1799]" metricMagnitude="3" metricUnit="m" metricValue="4.285" pageId="9" pageNumber="56" unit="m" value="4285.0">
<quantity id="4CE7FDC7153B0240279F89A4FD17943B" box="[663,748,1772,1799]" metricMagnitude="3" metricUnit="m" metricValue="4.285" pageId="9" pageNumber="56" unit="m" value="4285.0">4285 m</quantity>
asl
</elevation>
.,
<date id="FFA176E2153B0240262F89A5FC44943B" box="[807,959,1772,1799]" pageId="9" pageNumber="56" value="2015-05-22">
<collectingDate id="EFE58F0A153B0240262F89A5FC44943B" box="[807,959,1772,1799]" pageId="9" pageNumber="56" value="2015-05-22">22 May 2015</collectingDate>
</date>
, leg.
<collectorName id="26EA35F4153B024026F289A5FB86943A" box="[1018,1149,1772,1799]" pageId="9" pageNumber="56">I. Belousov</collectorName>
, I.
<collectorName id="26EA35F4153B0240219789A4FAEA943B" box="[1183,1297,1772,1799]" pageId="9" pageNumber="56">Kabak, G.</collectorName>
Davidian.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30503A9153B024025CF8858FDAC94E2" pageId="9" pageNumber="56" type="diagnosis">
<paragraph id="8BA05022153B024025CF8858FDAC94E2" blockId="9.[151,1437,1628,2015]" pageId="9" pageNumber="56">
<emphasis id="B96B8C30153B024025CF8858FEB99417" bold="true" box="[199,322,1808,1835]" pageId="9" pageNumber="56">Diagnosis.</emphasis>
The species differs from
<taxonomicName id="4C1F2BA1153B024027618858FCAE9416" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[617,853,1808,1834]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="muli" status="sp. nov.">
<emphasis id="B96B8C30153B024027618858FCAE9416" box="[617,853,1808,1834]" italics="true" pageId="9" pageNumber="56">Nepalmatoiulus muli</emphasis>
</taxonomicName>
<emphasis id="B96B8C30153B024026558859FC439417" bold="true" box="[861,952,1809,1835]" pageId="9" pageNumber="56">
<taxonomicNameLabel id="A258314B153B024026558859FC439417" box="[861,952,1809,1835]" pageId="9" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the brown (dark brown or light brown) coloration, broader gonopods, absence of a subapical branch of the solenomere, strong broader dentiform mesomeral process of opisthomere, sharply curved margin of opisthomere velum, strong more broad anterior gonopods, less obliquely rounded apex of promere, distal position of promere with rudimentary telopodite, distal ring of male leg pair 1 with one seta, near diamond-shaped penis, axial and near apical position of gland opening on male coxa 2, and a longer setiform accessory claw.
</paragraph>
</subSubSection>
<subSubSection id="C30503A91538024325CF8FD0FBF296DB" pageId="10" pageNumber="57" type="description">
<paragraph id="8BA050221538024325CF8FD0FA91960F" blockId="10.[151,1437,152,1547]" pageId="10" pageNumber="57">
<emphasis id="B96B8C301538024325CF8FD0FEA2938F" bold="true" box="[199,345,152,179]" pageId="10" pageNumber="57">Description.</emphasis>
Male. Length
<quantity id="4CE7FDC71538024324F38FD1FD8E938F" box="[507,629,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.7" metricValueMin="2.7" pageId="10" pageNumber="57" unit="mm" value="32.0" valueMax="37.0" valueMin="27.0">2737 mm</quantity>
, vertical diameter 2.0
<quantity id="4CE7FDC715380243267C8FD1FC30938F" box="[884,971,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="57" unit="mm" value="2.9">2.9 mm</quantity>
, with 55(1) 56(1) rings, excluding telson. Coloration of
<typeStatus id="54A4EE8015380243246F8FF5FE3293EB" box="[359,457,189,215]" pageId="10" pageNumber="57" type="holotype">holotype</typeStatus>
in alcohol dark brown, with a broad dorsal light longitudinal stripe. Coloration of maleparatype from SE Niguge light brown with dark brown thin lateral strip but without a broad dorsal light longitudinal stripe (it is possible that this male is discolored). Legs and antennae of
<typeStatus id="54A4EE801538024326B88E4DFBE99223" box="[944,1042,261,287]" pageId="10" pageNumber="57" type="holotype">holotype</typeStatus>
and all
<typeStatus id="54A4EE801538024321628E4DFB2E9223" box="[1130,1237,261,287]" pageId="10" pageNumber="57" type="paratype">paratypes</typeStatus>
dark brown, eyes black. Labrum yellow. Gnathochilarium light brown with yellow distal part.
<emphasis id="B96B8C301538024326FA8E61FBC9927F" bold="true" box="[1010,1074,297,323]" pageId="10" pageNumber="57">Head</emphasis>
smooth, with 2 epicranial setae, 6 supralabral setae, not less than 30 labral setae and a group of setae on forehead and on each lateral side below antenna. Eye patches suboval, each composed of not less than 70 ocelli. Antennae medium-sized, rather slender and clavate; length and width ratios of the antennomeres as in majority of
<taxonomicName id="4C1F2BA11538024326AA8EDDFBA89293" authorityName="Mauries" authorityYear="1983" box="[930,1107,405,431]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B96B8C301538024326AA8EDDFBA89293" box="[930,1107,405,431]" italics="true" pageId="10" pageNumber="57">Nepalmatoiulus</emphasis>
</taxonomicName>
. Gnathochilarium with about 20 nonapical stipital setae. Mandibular stipite with subtriangular rounded smooth lobe; not less than 5 setae on each lamella linguales.
<emphasis id="B96B8C3015380243246C8E94FE4692CB" bold="true" box="[356,445,476,503]" pageId="10" pageNumber="57">Collum</emphasis>
at posterior margin with short striae; laterally with several longer lower striae; the lower striae not extending from posterior to anterior margin.A transverse row of sparse setae at hind edge of collum. Body cylindrical. Prozona smooth with indistinct obliquely located striae. Suture between pro- and metazonite distinct. Metazonite gradually growing narrower toward body ventral part. Metazona with dense, regular, longitudinal striae reaching hind margin (1516 striae in an approximate square with sides equal to metazonital length of a midbody ring dorsal part). A transverse row of thin setae at hind edge of metazonites, setae denser toward telson. Ozopores small, lying behind suture dividing pro- and metazona, in touch with suture only in anterior body part, set off from the suture in the second half of the body. Limbus straight, smooth (of Type
<quantity id="4CE7FDC71538024326D48D91FBFC91CF" box="[988,1031,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="57" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EF8E2DD31538024321058D91FB5091CF" author="Enghoff, H." box="[1037,1195,729,755]" pageId="10" pageNumber="57" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
).
<emphasis id="B96B8C301538024321B68D91FAF091CF" bold="true" box="[1214,1291,729,755]" pageId="10" pageNumber="57">Telson</emphasis>
with preanal ring covered with short sparse setae. Anal valves, subanal scale, and caudal process of epiproct densely setose. Epiproct caudal process of middle length, carrying at tip a claw-shaped process curved dorsally.
<emphasis id="B96B8C301538024321F98C69FAD29007" bold="true" box="[1265,1321,801,827]" pageId="10" pageNumber="57">Legs</emphasis>
relatively short and slender. Very delicately serrate ventral pads present on postfemur and tibia, starting from legs 2. Claw of all legs at base with a long (longer than claw) setiform accessory claw ventrally but without accessory claw dorsally. Leg pair 1 forming hook, the distal ring not coming into close contact with the basal rings (“open hook” type in
<bibRefCitation id="EF8E2DD315380243206E8CC5FEF290F7" author="Enghoff, H." pageId="10" pageNumber="57" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
), postfemur with inflated scaly-rugose ventral surface, coxa and distal ring with one seta, tarsal remnant minute (
<figureCitation id="13244CA71538024325FE8C9DFEB090D3" box="[246,331,980,1007]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Fig. 45</figureCitation>
). Coxa 2 with one mesapical oral seta, gland opening positioned in near apical and axial position (
<figureCitation id="13244CA71538024325A88CB1FEE0972F" box="[160,283,1017,1043]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Fig. 40, 42</figureCitation>
) according to
<bibRefCitation id="EF8E2DD31538024324B48CB1FD97972F" author="Enghoff, H." box="[444,620,1017,1043]" pageId="10" pageNumber="57" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff (1987)</bibRefCitation>
.
<emphasis id="B96B8C3015380243277F8CB0FD4C972F" bold="true" box="[631,695,1016,1043]" pageId="10" pageNumber="57">Penis</emphasis>
nearly diamond-shaped, about 2 times longer than wide (
<figureCitation id="13244CA71538024320348CB1FA74972F" box="[1340,1423,1017,1043]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Fig. 41</figureCitation>
). Ventral margins of body ring VII with ventrad subrectangular apically rounded lobes.
<emphasis id="B96B8C3015380243216B8B54FB27970B" bold="true" box="[1123,1244,1052,1079]" pageId="10" pageNumber="57">Gonopods</emphasis>
slightly protruding. Anterior gonopod flattened, robust, with subparallel margins, apically broadly rounded, subapically excavated for accommodation of mesomeral process, distal margin of the excavation slightly papillate; distal third of the posterior surface with knoblike, flattened, rudimentary telopodite (
<figureCitation id="13244CA71538024326808BC1FBF6979F" box="[904,1037,1161,1187]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Figs 38, 43</figureCitation>
). Flagellum slender, middle length, caudally covered with cuticular conical obtuse denticles in distal part. Basal part of the flagellum covered with cuticular spikes. Opisthomere with strong dentiform mesomeral process apically papillate (
<figureCitation id="13244CA71538024321A68B99FAFF97D7" box="[1198,1284,1233,1259]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Fig. 39</figureCitation>
). Velum with smooth, relatively sharply curved (with a small notch near the mesomeral process) margin. Margin of accessory membrane partly covered with denticles. Solenomere long, slender, spinose throughout, with basal blade (
<emphasis id="B96B8C301538024320338A51FAA1960F" bold="true" box="[1339,1370,1305,1331]" pageId="10" pageNumber="57">bb</emphasis>
).
</paragraph>
<paragraph id="8BA050221538024325C78A75FCE696A3" blockId="10.[151,1437,152,1547]" pageId="10" pageNumber="57">
Female. Length
<quantity id="4CE7FDC715380243248F8A75FDF8966B" box="[391,515,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="4.1" metricValueMin="2.7" pageId="10" pageNumber="57" unit="mm" value="34.0" valueMax="41.0" valueMin="27.0">2741 mm</quantity>
, vertical diameter 2.54.0 mm, with 52(2), 54(2), 53(1), 59(1) rings, excluding telson. Small females dark brown, without dorsal light longitudinal spripe. Head with 4 supralabral setae. The anterior ventral edge of ring 3 as low ridge with two tops.
</paragraph>
<paragraph id="8BA050221538024325CF8AE1FBF296DB" blockId="10.[151,1437,152,1547]" pageId="10" pageNumber="57">
Vulva with operculum and bursa of about equal height; apical excavation indistinct, apical lobes of bursa not long; posterior median plate with about 30 setae arranged irregularly (
<figureCitation id="13244CA71538024326A08A85FC0796DB" box="[936,1020,1485,1511]" captionStart="FIGURES 3845" captionStartId="11.[151,264,1432,1457]" captionTargetBox="[174,1413,174,1406]" captionTargetId="figure@11.[174,1413,174,1412]" captionTargetPageId="11" captionText="FIGURES 3845. Nepalmatoiulus tianbaoshanensis sp. nov., male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view; bb, basal blade of solenomere. Male: Figs 3843, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 3840, 42, 44)." figureDoi="http://doi.org/10.5281/zenodo.3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="10" pageNumber="57">Fig. 44</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30503A91538024325CF8AB9FCA19537" box="[199,858,1521,1547]" pageId="10" pageNumber="57" type="etymology">
<paragraph id="8BA050221538024325CF8AB9FCA19537" blockId="10.[151,1437,152,1547]" box="[199,858,1521,1547]" pageId="10" pageNumber="57">
<emphasis id="B96B8C301538024325CF8AB9FEA99537" bold="true" box="[199,338,1521,1547]" pageId="10" pageNumber="57">Etymology:</emphasis>
The specific epithet refers to the
<typeStatus id="54A4EE801538024327C18AB9FD029537" box="[713,761,1521,1547]" pageId="10" pageNumber="57">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>