218 lines
33 KiB
XML
218 lines
33 KiB
XML
<document id="1B1B45D7E8C3072487BBC946BEDAA7B7" ID-CLB-Dataset="24234" ID-DOI="10.11646/zootaxa.4729.1.3" ID-GBIF-Dataset="22815f4d-7c51-45d1-b6b3-8216ea02e14c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3629007" ID-ZooBank="BBFF38A8-8BD6-4A70-A2DA-0804D5DE2C28" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580180119895" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2020" docId="03B6E13415380245259F8915FAE59743" docLanguage="en" docName="zootaxa.4729.1.3.pdf" docOrigin="Zootaxa 4729 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus pallidus Mikhaljova 2020, new species" docType="treatment" docVersion="7" lastPageNumber="59" masterDocId="FF8F994C1532024925088F48FFFB933C" masterDocTitle="New data on the fauna of China, part I: the taxonomy of the millipede family Julidae (Diplopoda)" masterLastPageNumber="66" masterPageNumber="47" pageNumber="57" updateTime="1698779498477" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3A4A1E30A64E5DD846DB660B05435623">New data on the fauna of China, part I: the taxonomy of the millipede family Julidae (Diplopoda)</mods:title>
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<mods:name id="E2D144C58D591CB07E40CD1062D2B982" type="personal">
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<mods:namePart id="AF90ED4C56CA60AC3B1DC8E880D2109E">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="23EF7588CCEC0D10558E2A0099C32658">Zootaxa</mods:title>
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<mods:part id="129DD669C0C720D222014E419B8ECD62">
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<mods:date id="F38186EF7FD557278109CFCF635B37FE">2020</mods:date>
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<mods:number id="F880A6F235AA751C3B277B9273EC196F">2020-01-28</mods:number>
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<mods:number id="2459D0CE55DAE0097D088BD1E3735051">4729</mods:number>
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<mods:identifier id="CE67FD9D3EA076779927BB065B69A681" type="DOI">10.11646/zootaxa.4729.1.3</mods:identifier>
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<treatment id="03B6E13415380245259F8915FAE59743" ID-DOI="http://doi.org/10.5281/zenodo.5918711" ID-GBIF-Taxon="161840491" ID-Zenodo-Dep="5918711" LSID="urn:lsid:plazi:treatment:03B6E13415380245259F8915FAE59743" httpUri="http://treatment.plazi.org/id/03B6E13415380245259F8915FAE59743" lastPageId="12" lastPageNumber="59" pageId="10" pageNumber="57">
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<subSubSection id="C30503A915380243259F8915FDAD954B" box="[151,598,1629,1656]" pageId="10" pageNumber="57" type="nomenclature">
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<paragraph id="8BA0502215380243259F8915FDAD954B" blockId="10.[151,598,1629,1691]" box="[151,598,1629,1656]" pageId="10" pageNumber="57">
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<heading id="D0E8E74E15380243259F8915FDAD954B" bold="true" box="[151,598,1629,1656]" fontSize="11" level="1" pageId="10" pageNumber="57" reason="1">
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<emphasis id="B96B8C3015380243259F8915FDAD954B" bold="true" box="[151,598,1629,1656]" pageId="10" pageNumber="57">
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<taxonomicName id="4C1F2BA115380243259F8915FE449544" ID-CoL="7b5e1d81-aae1-401f-95a3-e677cd5fa707" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[151,447,1629,1656]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
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<emphasis id="B96B8C3015380243259F8915FE449544" bold="true" box="[151,447,1629,1656]" italics="true" pageId="10" pageNumber="57">Nepalmatoiulus pallidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A258314B1538024324CE8915FDAD954B" box="[454,598,1629,1655]" pageId="10" pageNumber="57" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30503A915380243259F89C9FEEF95A7" box="[151,276,1664,1691]" pageId="10" pageNumber="57" type="description">
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<paragraph id="8BA0502215380243259F89C9FEEF95A7" blockId="10.[151,598,1629,1691]" box="[151,276,1664,1691]" pageId="10" pageNumber="57">
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<figureCitation id="13244CA715380243259F89C9FEEF95A7" box="[151,276,1664,1691]" captionStart-0="FIGURES 46–52" captionStart-1="FIGURES 53–57" captionStart-2="FIGURE 58" captionStartId-0="13.[151,264,1912,1937]" captionStartId-1="14.[151,264,1451,1476]" captionStartId-2="15.[151,250,1992,2017]" captionTargetBox-0="[204,1383,841,1889]" captionTargetBox-1="[151,1435,301,1423]" captionTargetBox-2="[439,1161,833,1953]" captionTargetId-0="figure@13.[204,1383,841,1892]" captionTargetId-1="figure@14.[151,1435,301,1431]" captionTargetId-2="figure@15.[423,1164,805,1973]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." captionText-1="FIGURES 53–57. Nepalmatoiulus pallidus sp. nov., male holotype. 53, promere, caudal view; 54, distal part of promere, caudal view; 55, central part of promere, caudal view; 56, distal portion of promere flagellum; 57, basal portion of promere flagellum. Scales: 2 μm (Figs 56–57), 10 μm (Fig. 55), 20 μm (Fig. 54), 100 μm (Fig. 53)." captionText-2="FIGURE 58. Nepalmatoiulus pallidus sp. nov., male holotype. Opisthomere, mesal view. Scale in mm." figureDoi-0="http://doi.org/10.5281/zenodo.3629025" figureDoi-1="http://doi.org/10.5281/zenodo.3629027" figureDoi-2="http://doi.org/10.5281/zenodo.3629029" httpUri-0="https://zenodo.org/record/3629025/files/figure.png" httpUri-1="https://zenodo.org/record/3629027/files/figure.png" httpUri-2="https://zenodo.org/record/3629029/files/figure.png" pageId="10" pageNumber="57">Figs 46–58</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30503A915380243259F8980FF3A9473" pageId="10" pageNumber="57" type="materials_examined">
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<paragraph id="8BA0502215380243259F8980FF3A9473" blockId="10.[151,1437,1736,2015]" pageId="10" pageNumber="57">
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<emphasis id="B96B8C3015380243259F8980FE8195DF" bold="true" box="[151,378,1736,1763]" pageId="10" pageNumber="57">Material examined</emphasis>
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:
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<materialsCitation id="3B775A7F1538024324828981FA66943B" ID-GBIF-Occurrence="2560337979" collectingDate="2013-05-30" collectionCode="ZMUM" collectorName="I. Belousov & Belousov, I & Kabak, G." country="China" elevation="3365" latitude="27.334167" location="Lijiang Shangrila" longLatPrecision="20" longitude="99.876114" pageId="10" pageNumber="57" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<emphasis id="B96B8C301538024324828981FE0A95DF" box="[394,497,1737,1763]" italics="true" pageId="10" pageNumber="57">
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<typeStatus id="54A4EE801538024324828981FE0A95DF" box="[394,497,1737,1763]" pageId="10" pageNumber="57" type="holotype">Holotype</typeStatus>
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</emphasis>
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: male (
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<collectionCode id="ED0EC8E715380243274E8981FD5895DF" box="[582,675,1737,1763]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="57">ZMUM</collectionCode>
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),
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<collectingCountry id="F30810B21538024327BF8981FD0695DF" box="[695,765,1737,1763]" name="China" pageId="10" pageNumber="57">China</collectingCountry>
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,
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<collectingRegion id="49DB9EC01538024326008981FC3795DF" box="[776,972,1737,1763]" country="China" name="Yunnan" pageId="10" pageNumber="57">Yunnan Province</collectingRegion>
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,
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<location id="8EC006F91538024326D08981FB6495DF" LSID="urn:lsid:plazi:treatment:03B6E13415380245259F8915FAE59743:8EC006F91538024326D08981FB6495DF" box="[984,1183,1737,1763]" country="China" latitude="27.334167" longLatPrecision="20" longitude="99.876114" name="Lijiang Shangrila" pageId="10" pageNumber="57" stateProvince="Yunnan">Lijiang Shangrila</location>
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, 214 Ntn. Road
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<locationDeviation id="9A1249001538024320508981FEC8943B" location="Edi village" pageId="10" pageNumber="57">WSW of Edi village</locationDeviation>
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||
,
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<geoCoordinate id="EE2B36E515380243243689A5FE31943B" box="[318,458,1773,1799]" degrees="27" direction="north" minutes="20" orientation="latitude" pageId="10" pageNumber="57" precision="15" seconds="3" value="27.334167">27°20´3´´ N</geoCoordinate>
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,
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<geoCoordinate id="EE2B36E51538024324DD89A4FD90943B" box="[469,619,1772,1799]" degrees="99" direction="east" minutes="52" orientation="longitude" pageId="10" pageNumber="57" precision="15" seconds="34" value="99.876114">99°52´34´´ E</geoCoordinate>
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, H =
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<elevation id="0032B7111538024327A289A4FCD1943B" box="[682,810,1772,1799]" metricMagnitude="3" metricUnit="m" metricValue="3.3649999999999998" pageId="10" pageNumber="57" unit="m" value="3365.0">
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||
<quantity id="4CE7FDC71538024327A289A4FD04943B" box="[682,767,1772,1799]" metricMagnitude="3" metricUnit="m" metricValue="3.3649999999999998" pageId="10" pageNumber="57" unit="m" value="3365.0">3365 m</quantity>
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asl
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</elevation>
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.,
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<date id="FFA176E215380243263389A5FC28943B" box="[827,979,1773,1799]" pageId="10" pageNumber="57" value="2013-05-30">
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||
<collectingDate id="EFE58F0A15380243263389A5FC28943B" box="[827,979,1773,1799]" pageId="10" pageNumber="57" value="2013-05-30">30 May 2013</collectingDate>
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</date>
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, leg.
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<collectorName id="26EA35F415380243210789A5FB69943B" box="[1039,1170,1773,1799]" pageId="10" pageNumber="57">I. Belousov</collectorName>
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||
, I.
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<collectorName id="26EA35F41538024321BD89A5FADC943B" box="[1205,1319,1773,1799]" pageId="10" pageNumber="57">Kabak, G.</collectorName>
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Davidian.
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</materialsCitation>
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<materialsCitation id="3B775A7F15380243259F885AFF3A9473" ID-GBIF-Occurrence="2560337983" collectingDate="2013-05-30" collectionCode="ZMUM" collectorName="I. Belousov & Belousov, I & Kabak, G." country="China" elevation="3365" latitude="27.334167" location="Lijiang Shangrila" longLatPrecision="20" longitude="99.876114" pageId="10" pageNumber="57" specimenCount="1" specimenCount-juvenile="1" stateProvince="Yunnan" typeStatus="paratype">
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<emphasis id="B96B8C3015380243259F885AFF059417" box="[151,254,1810,1835]" italics="true" pageId="10" pageNumber="57">
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<typeStatus id="54A4EE8015380243259F885AFF059417" box="[151,254,1810,1835]" pageId="10" pageNumber="57" type="paratype">Paratype</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D199BAB1538024324068859FE849417" box="[270,383,1809,1835]" pageId="10" pageNumber="57" type="juvenile">1 juvenile</specimenCount>
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(
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<collectionCode id="ED0EC8E71538024324878859FE179417" box="[399,492,1809,1835]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="57">ZMUM</collectionCode>
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), same locality as in
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<typeStatus id="54A4EE801538024327D68859FCB89417" box="[734,835,1809,1835]" pageId="10" pageNumber="57" type="holotype">holotype</typeStatus>
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,
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<date id="FFA176E21538024326478859FC119417" box="[847,1002,1809,1835]" pageId="10" pageNumber="57" value="2013-05-30">
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<collectingDate id="EFE58F0A1538024326478859FC119417" box="[847,1002,1809,1835]" pageId="10" pageNumber="57" value="2013-05-30">30 May 2013</collectingDate>
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</date>
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, leg.
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<collectorName id="26EA35F41538024321218859FB569417" box="[1065,1197,1809,1835]" pageId="10" pageNumber="57">I. Belousov</collectorName>
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, I.
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<collectorName id="26EA35F41538024321DA8859FABD9417" box="[1234,1350,1809,1835]" pageId="10" pageNumber="57">Kabak, G.</collectorName>
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Davidian.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30503A91538024325CF8810FB3E94E3" pageId="10" pageNumber="57" type="diagnosis">
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<paragraph id="8BA050221538024325CF8810FB3E94E3" blockId="10.[151,1437,1736,2015]" pageId="10" pageNumber="57">
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<emphasis id="B96B8C301538024325CF8810FEB9944F" bold="true" box="[199,322,1880,1907]" pageId="10" pageNumber="57">Diagnosis.</emphasis>
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The species seems to be particularly closely related to
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<taxonomicName id="4C1F2BA11538024326CC8811FB4A944F" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[964,1201,1881,1907]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="muli" status="sp. nov.">
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<emphasis id="B96B8C301538024326CC8811FB4A944F" box="[964,1201,1881,1907]" italics="true" pageId="10" pageNumber="57">Nepalmatoiulus muli</emphasis>
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</taxonomicName>
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<emphasis id="B96B8C301538024321B28811FAED944F" bold="true" box="[1210,1302,1881,1907]" pageId="10" pageNumber="57">
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<taxonomicNameLabel id="A258314B1538024321B28811FAED944F" box="[1210,1302,1881,1907]" pageId="10" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, but differs mainly in the rectangular apex of the anterior gonopod, broader posterior gonopod solenomere, shorter subapical branch of solenomere, nearly diamond-shaped penis, male coxae 2 with two setae, distal ring of male leg pair 1 with one seta, axial and near apical position of gland opening on male coxa 2, and lighter coloration.
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||
</paragraph>
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</subSubSection>
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||
<caption id="DF6000AA15390242259F8AD0FD8E9520" ID-DOI="http://doi.org/10.5281/zenodo.3629023" ID-Zenodo-Dep="3629023" httpUri="https://zenodo.org/record/3629023/files/figure.png" pageId="11" pageNumber="58" startId="11.[151,264,1432,1457]" targetBox="[174,1413,174,1406]" targetPageId="11">
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<paragraph id="8BA0502215390242259F8AD0FD8E9520" blockId="11.[151,1437,1432,1565]" pageId="11" pageNumber="58">
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<emphasis id="B96B8C3015390242259F8AD0FEAE968D" bold="true" box="[151,341,1432,1457]" pageId="11" pageNumber="58">FIGURES 38–45.</emphasis>
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<taxonomicName id="4C1F2BA11539024224538AD0FD49968C" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[347,690,1432,1456]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="tianbaoshanensis" status="sp. nov.">
|
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<emphasis id="B96B8C301539024224538AD0FD49968C" box="[347,690,1432,1456]" italics="true" pageId="11" pageNumber="58">Nepalmatoiulus tianbaoshanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B8C301539024227B18AD0FCF2968C" bold="true" box="[697,777,1432,1456]" pageId="11" pageNumber="58">
|
||
<taxonomicNameLabel id="A258314B1539024227B18AD0FCF2968C" box="[697,777,1432,1456]" pageId="11" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male and female paratypes. 38, promere, caudal view; 39, opisthomere, mesal view; 40, leg pair 2, front view; 41, penis, front view; 42, coxa 2, front view; 43, distal part of promere, caudal view; 44, vulva, caudal view; 45, leg pair 1, front view;
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<emphasis id="B96B8C301539024227D58AA8FD0296C4" bold="true" box="[733,761,1504,1528]" pageId="11" pageNumber="58">bb</emphasis>
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, basal blade of solenomere. Male: Figs 38–43, 45. Scales: 20 μm (Figs 41, 43, 45), 100 μm (Figs 38–40, 42, 44).
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</paragraph>
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</caption>
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||
<subSubSection id="C30503A91539024525CF8902FE6A9767" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58" type="description">
|
||
<paragraph id="8BA050221539024525CF8902FE2B90D3" blockId="11.[151,1437,1610,2034]" lastBlockId="12.[151,1437,153,1151]" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
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<emphasis id="B96B8C301539024225CF8902FEA29559" bold="true" box="[199,345,1610,1637]" pageId="11" pageNumber="58">Description.</emphasis>
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Male
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<typeStatus id="54A4EE801539024224A88903FDFD9559" box="[416,518,1611,1637]" pageId="11" pageNumber="58" type="holotype">holotype</typeStatus>
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. Length about
|
||
<quantity id="4CE7FDC71539024227A28903FD009559" box="[682,763,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="58" unit="mm" value="34.0">34 mm</quantity>
|
||
, vertical diameter about
|
||
<quantity id="4CE7FDC71539024221078903FB9C9559" box="[1039,1127,1611,1637]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="58" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, with 54 (–1) rings, excluding telson. Coloration yellow-grayish, with a narrow middorsal dark line and two broad lateral marbled brown stripes; the stripes gradually growing brighter and more noticeable toward telson. Preanal ring dark brown; anal valves brown. Venter (including basal parts of legs) yellowish. Anterior and ventral portions of head yellowish. Occiput and forehead marbled brown, dark brown stripe between eye patches. Eyes black. Antennae dark brown. Distal parts of legs marbled brown.
|
||
<emphasis id="B96B8C3015390242272789B7FD949425" bold="true" box="[559,623,1791,1817]" pageId="11" pageNumber="58">Head</emphasis>
|
||
smooth, 2 epicranial setae, 4 supralabral setae, and 28 labral setae. Eye patches almost oval, composed of about 80 ocelli. Antennae medium-sized, rather slender and clavate. Length ratios of antennomeres 3–7 as 4:3.4:4:2.6:1, width ratios as 1.7:1.8:1.9:1.7:1, respectively. Antennomeres 5 and 6 with incomplete distodorsal corolla of sensory bacilli (
|
||
<figureCitation id="13244CA71539024226148823FC8994B9" box="[796,882,1899,1925]" captionStart="FIGURES 46–52" captionStartId="13.[151,264,1912,1937]" captionTargetBox="[204,1383,841,1889]" captionTargetId="figure@13.[204,1383,841,1892]" captionTargetPageId="13" captionText="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." figureDoi="http://doi.org/10.5281/zenodo.3629025" httpUri="https://zenodo.org/record/3629025/files/figure.png" pageId="11" pageNumber="58">Fig. 48</figureCitation>
|
||
). Mandibular stipites with triangular lobes. Gnathochilarium: not less than 15 nonapical stipital setae; lamellae linguales each with not less than 10 setae.
|
||
<emphasis id="B96B8C3015390242204C88C6FA669495" bold="true" box="[1348,1437,1934,1961]" pageId="11" pageNumber="58">Collum</emphasis>
|
||
laterally with distinct lower striae of different length at posterior margin not reaching to anterior margin (excluding lowest 1–2 striae), dorsally with distinct short striae at posterior margin only. A transverse row of setae at hind edge of collum. Body rings circular. Prozona smooth except for ventral part with very shallow obliquely located striae and for dorsal part with transverse very indistinct striae. The metazonite gradually growing narrower toward body ventral part. Metazona with dense, regular, longitudinal striae reaching hind margin (23–24 striae in an approximate square with sides equal to metazonital length of a dorsal side of midbody ring). Limbus straight, smooth (of Type
|
||
<quantity id="4CE7FDC7153E024520868E4DFF56927F" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="59" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EF8E2DD3153E024525BB8E61FEAA927F" author="Enghoff, H." box="[179,337,297,323]" pageId="12" pageNumber="59" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
). A transverse row of sparse, thin setae at hind edge of metazonites, setae gradually growing denser and longer toward telson. Ozopores small, lying behind suture dividing pro- and metazona without touching the suture.
|
||
<emphasis id="B96B8C30153E024525E18E39FECD92B7" bold="true" box="[233,310,369,395]" pageId="12" pageNumber="59">Telson</emphasis>
|
||
with caudal dorsal projection straight and long, covered with setae and carrying at tip a claw-shaped process curved dorsally. Preanal ring, anal valves, and subanal scale densely setose.
|
||
<emphasis id="B96B8C30153E024521718EDDFB4A9293" bold="true" box="[1145,1201,405,431]" pageId="12" pageNumber="59">Legs</emphasis>
|
||
relatively short and slender. Very delicately serrate ventral pads present on postfemur and tibia, starting from legs 2. Claw of all legs at base with a long (longer than claw) setiform accessory claw ventrally but without accessory claw dorsally. Leg pair 1 forming a hook, the distal ring not coming into close contact with the basal rings (“open hook” type in
|
||
<bibRefCitation id="EF8E2DD3153E024520498D49FF289103" author="Enghoff, H." pageId="12" pageNumber="59" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
); postfemur with inflated scaly-rugose ventral surface, tip indistinctly wrinkled, coxa with one seta, distal ring with seta, without tarsal remnant (
|
||
<figureCitation id="13244CA7153E0245271E8D01FD91915F" box="[534,618,584,611]" captionStart="FIGURES 46–52" captionStartId="13.[151,264,1912,1937]" captionTargetBox="[204,1383,841,1889]" captionTargetId="figure@13.[204,1383,841,1892]" captionTargetPageId="13" captionText="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." figureDoi="http://doi.org/10.5281/zenodo.3629025" httpUri="https://zenodo.org/record/3629025/files/figure.png" pageId="12" pageNumber="59">Fig. 51</figureCitation>
|
||
). Coxa 2 with two mesapical oral setae, gland opening positioned in apical and axial position (
|
||
<figureCitation id="13244CA7153E0245249C8D25FE1E91BB" box="[404,485,621,647]" captionStart="FIGURES 46–52" captionStartId="13.[151,264,1912,1937]" captionTargetBox="[204,1383,841,1889]" captionTargetId="figure@13.[204,1383,841,1892]" captionTargetPageId="13" captionText="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." figureDoi="http://doi.org/10.5281/zenodo.3629025" httpUri="https://zenodo.org/record/3629025/files/figure.png" pageId="12" pageNumber="59">Fig. 47</figureCitation>
|
||
) according to
|
||
<bibRefCitation id="EF8E2DD3153E0245278E8D25FCC891BB" author="Enghoff, H." box="[646,819,621,647]" pageId="12" pageNumber="59" pagination="241 - 331" refId="ref10083" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff (1987)</bibRefCitation>
|
||
(the seta attached to one of the coxal gland openings is probably outside debris).
|
||
<emphasis id="B96B8C30153E024524BF8DD8FE0C9197" bold="true" box="[439,503,656,683]" pageId="12" pageNumber="59">Penis</emphasis>
|
||
subtrapezoidal, about 1.3 times longer than wide, with apical excavation (
|
||
<figureCitation id="13244CA7153E024520338DD9FA6B9190" box="[1339,1424,657,684]" captionStart="FIGURES 46–52" captionStartId="13.[151,264,1912,1937]" captionTargetBox="[204,1383,841,1889]" captionTargetId="figure@13.[204,1383,841,1892]" captionTargetPageId="13" captionText="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." figureDoi="http://doi.org/10.5281/zenodo.3629025" httpUri="https://zenodo.org/record/3629025/files/figure.png" pageId="12" pageNumber="59">Fig. 46</figureCitation>
|
||
). Ventral margin of body ring VII with ventrad subtriangular apically rounded lobes.
|
||
<emphasis id="B96B8C30153E0245214B8DFCFB4791F3" bold="true" box="[1091,1212,692,719]" pageId="12" pageNumber="59">Gonopods</emphasis>
|
||
slightly protruding. Anterior gonopod flattened, with parallel margins, apically obliquely rounded, in posterior view apically excavated for accommodation of mesomeral process, distal margins of the excavation papillate, subcentrally with knoblike flattened rudimentary telopodite (
|
||
<figureCitation id="13244CA7153E0245271C8C69FD639007" box="[532,664,800,827]" captionStart="FIGURES 53–57" captionStartId="14.[151,264,1451,1476]" captionTargetBox="[151,1435,301,1423]" captionTargetId="figure@14.[151,1435,301,1431]" captionTargetPageId="14" captionText="FIGURES 53–57. Nepalmatoiulus pallidus sp. nov., male holotype. 53, promere, caudal view; 54, distal part of promere, caudal view; 55, central part of promere, caudal view; 56, distal portion of promere flagellum; 57, basal portion of promere flagellum. Scales: 2 μm (Figs 56–57), 10 μm (Fig. 55), 20 μm (Fig. 54), 100 μm (Fig. 53)." figureDoi="http://doi.org/10.5281/zenodo.3629027" httpUri="https://zenodo.org/record/3629027/files/figure.png" pageId="12" pageNumber="59">Figs 53–55</figureCitation>
|
||
). Flagellum slender, middle length, caudally covered with cuticular conical obtuse denticles in the distal part (
|
||
<figureCitation id="13244CA7153E024527868C0DFD13905C" box="[654,744,837,864]" captionStart="FIGURES 53–57" captionStartId="14.[151,264,1451,1476]" captionTargetBox="[151,1435,301,1423]" captionTargetId="figure@14.[151,1435,301,1431]" captionTargetPageId="14" captionText="FIGURES 53–57. Nepalmatoiulus pallidus sp. nov., male holotype. 53, promere, caudal view; 54, distal part of promere, caudal view; 55, central part of promere, caudal view; 56, distal portion of promere flagellum; 57, basal portion of promere flagellum. Scales: 2 μm (Figs 56–57), 10 μm (Fig. 55), 20 μm (Fig. 54), 100 μm (Fig. 53)." figureDoi="http://doi.org/10.5281/zenodo.3629027" httpUri="https://zenodo.org/record/3629027/files/figure.png" pageId="12" pageNumber="59">Fig. 56</figureCitation>
|
||
); basal part of the flagellum covered with cuticular spikes (
|
||
<figureCitation id="13244CA7153E024525A88C21FF0390B8" box="[160,248,873,900]" captionStart="FIGURES 53–57" captionStartId="14.[151,264,1451,1476]" captionTargetBox="[151,1435,301,1423]" captionTargetId="figure@14.[151,1435,301,1431]" captionTargetPageId="14" captionText="FIGURES 53–57. Nepalmatoiulus pallidus sp. nov., male holotype. 53, promere, caudal view; 54, distal part of promere, caudal view; 55, central part of promere, caudal view; 56, distal portion of promere flagellum; 57, basal portion of promere flagellum. Scales: 2 μm (Figs 56–57), 10 μm (Fig. 55), 20 μm (Fig. 54), 100 μm (Fig. 53)." figureDoi="http://doi.org/10.5281/zenodo.3629027" httpUri="https://zenodo.org/record/3629027/files/figure.png" pageId="12" pageNumber="59">Fig. 57</figureCitation>
|
||
). Opisthomere with slender mesomeral process, its apex with several tiny papillae; velum steeply sloping (
|
||
<figureCitation id="13244CA7153E024525978CC5FEBC909B" box="[159,327,908,935]" captionStart="FIGURES 46–52" captionStartId="13.[151,264,1912,1937]" captionTargetBox="[204,1383,841,1889]" captionTargetId="figure@13.[204,1383,841,1892]" captionTargetPageId="13" captionText="FIGURES 46–52. Nepalmatoiulus pallidus sp. nov., male holotype. 46, penis, front view; 47, coxae 2, front view; 48, distal part of antenna; 49, gonopods, mesal view, slightly rotated around its axis; 50, central part of opisthomere, mesal view, slightly rotated around its axis; 51, leg pair 1, front view; 52, apex of opisthomere mesomeral process; m, accessory membrane; b, solenomere subapical branch. Scales: 20 μm (Figs 46–47, 50–52), 100 μm (Figs 48–49)." figureDoi="http://doi.org/10.5281/zenodo.3629025" httpUri="https://zenodo.org/record/3629025/files/figure.png" pageId="12" pageNumber="59">Figs 49–50, 52</figureCitation>
|
||
,
|
||
<figureCitation id="13244CA7153E0245245A8CC4FE89909B" box="[338,370,908,935]" captionStart="FIGURE 58" captionStartId="15.[151,250,1992,2017]" captionTargetBox="[439,1161,833,1953]" captionTargetId="figure@15.[423,1164,805,1973]" captionTargetPageId="15" captionText="FIGURE 58. Nepalmatoiulus pallidus sp. nov., male holotype. Opisthomere, mesal view. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.3629029" httpUri="https://zenodo.org/record/3629029/files/figure.png" pageId="12" pageNumber="59">58</figureCitation>
|
||
). Additional membrane (
|
||
<emphasis id="B96B8C30153E0245278E8CC5FD66909B" bold="true" box="[646,669,909,935]" pageId="12" pageNumber="59">m</emphasis>
|
||
) small, with a smooth edge. Solenomere densely spinose throughout, with relatively short subapical branch (
|
||
<emphasis id="B96B8C30153E024527478CF9FDA490F7" bold="true" box="[591,607,945,971]" pageId="12" pageNumber="59">b</emphasis>
|
||
) covered with spines throughout (the base of the subapical branch is hard to see in the figures 49–50).
|
||
</paragraph>
|
||
<paragraph id="8BA05022153E024525CF8CB1FEA9970B" blockId="12.[151,1437,153,1151]" pageId="12" pageNumber="59">
|
||
Juvenile. Length about
|
||
<quantity id="4CE7FDC7153E024524C38CB1FDE0972F" box="[459,539,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="59" unit="mm" value="17.0">17 mm</quantity>
|
||
, vertical diameter about 2.0 mm, with 32 (–2) rings, excluding telson. Legs without ventral pads.
|
||
</paragraph>
|
||
<paragraph id="8BA05022153E024525CF8B09FE6A9767" blockId="12.[151,1437,153,1151]" box="[199,401,1089,1115]" pageId="12" pageNumber="59">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30503A9153E024525CF8B2DFAE59743" box="[199,1310,1125,1151]" pageId="12" pageNumber="59" type="etymology">
|
||
<paragraph id="8BA05022153E024525CF8B2DFAE59743" blockId="12.[151,1437,153,1151]" box="[199,1310,1125,1151]" pageId="12" pageNumber="59">
|
||
<emphasis id="B96B8C30153E024525CF8B2DFEA99743" bold="true" box="[199,338,1125,1151]" pageId="12" pageNumber="59">Etymology:</emphasis>
|
||
The specific epithet refers to the lighter coloration as compared to the
|
||
<taxonomicName id="4C1F2BA1153E024521638B2EFB459743" authority="Mikhaljova, 2020" authorityName="Mikhaljova" authorityYear="2020" box="[1131,1214,1125,1151]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="muli" status="sp. nov.">
|
||
<emphasis id="B96B8C30153E024521638B2EFB459743" box="[1131,1214,1125,1151]" italics="true" pageId="12" pageNumber="59">N. muli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B8C30153E024521CD8B2DFAE59743" bold="true" box="[1221,1310,1125,1151]" pageId="12" pageNumber="59">
|
||
<taxonomicNameLabel id="A258314B153E024521CD8B2DFAE59743" box="[1221,1310,1125,1151]" pageId="12" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |