711 lines
97 KiB
XML
711 lines
97 KiB
XML
<document id="379A97660841A09C27019CB3136E84C0" ID-CLB-Dataset="28158" ID-DOI="10.11646/zootaxa.4563.1.1" ID-GBIF-Dataset="3e3068c1-c158-4d97-9aec-31fc80551edb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2600858" ID-ZooBank="A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553170725873" checkinUser="plazi" docAuthor="Seeman, Owen D." docDate="2019" docId="03B63A508D67015FFF6014D7021D4691" docLanguage="en" docName="zootaxa.4563.1.1.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megisthanus womersleyi Seeman 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="33" masterDocId="FF8F42288D7D017FFFF7166307564009" masterDocTitle="The megisthanid mites (Mesostigmata: Megisthanidae) of Australia" masterLastPageNumber="40" masterPageNumber="1" pageNumber="27" updateTime="1698702915705" updateUser="ExternalLinkService">
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<mods:mods id="546AEC3B8164EC73C89515F82D08A899" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="07FB80E3A591F594942D19842223DACC">
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<mods:title id="14F695F887CCE78838E58197E440BE81">The megisthanid mites (Mesostigmata: Megisthanidae) of Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="940BCFDC04AA623E4DC096E26E41D0AF" type="personal">
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<mods:role id="7D9218257BCD7162D425A08F58A0E6ED">
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<mods:roleTerm id="F150612EEC367E4764FAEED00B2945E0">Author</mods:roleTerm>
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<mods:namePart id="4F0EFA26BF1C96955B8E52759994DAA4">Seeman, Owen D.</mods:namePart>
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<mods:typeOfResource id="1DE3CC428A4EB62DB0EB0256E9957AB9">text</mods:typeOfResource>
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<mods:titleInfo id="C70AA359D1AAAD76E529579085EC70DF">
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<mods:title id="01CC5D08583D45A51CCCA25F80AF04CB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2441A140C268E6F91046975A27786DEA">
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<mods:date id="52126902A96B33D7483AA7F1EC376882">2019</mods:date>
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<mods:detail id="EBBCE561AB5F3C18D6F11BAF8D6CB950" type="pubDate">
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<mods:number id="E3813B6D568BA6D59E89487990628427">2019-02-28</mods:number>
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</mods:detail>
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<mods:detail id="A5569F42B10FEC0E00D1FF9A26B6EE95" type="volume">
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<mods:number id="DD088B3CD516EAB68CB198B601003484">4563</mods:number>
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</mods:detail>
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<mods:detail id="E0FCCA5A271E4C382C0E61DC31B47CC4" type="issue">
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<mods:number id="2B350F3B20F2928CEC882235CCA5ABED">1</mods:number>
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<mods:start id="012B8C63363197135DB969C6C2722074">1</mods:start>
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<mods:end id="2D0C7FCD21C3F2285C06E909E033C329">40</mods:end>
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</mods:extent>
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<mods:classification id="C18D5F10B5E5C4EA11F63B138034BDF0">journal article</mods:classification>
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<mods:identifier id="039ED123A701C6339F670E76DC3523D8" type="CLB-Dataset">28158</mods:identifier>
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<mods:identifier id="345A001CA301634841564A5E05D654B8" type="DOI">10.11646/zootaxa.4563.1.1</mods:identifier>
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<mods:identifier id="5D879C1D62F83EB1A7772AF2E1147A21" type="GBIF-Dataset">3e3068c1-c158-4d97-9aec-31fc80551edb</mods:identifier>
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<mods:identifier id="2EBB46E1645A3C3DBBF0FD614B277E2E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="19F19DA0ABCC6DA3D0C12968B56554C2" type="Zenodo-Dep">2600858</mods:identifier>
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<mods:identifier id="11D1691878AD6F724DA16C4F14F93009" type="ZooBank">A44D4C11-ADA1-48A8-9F00-B46BDF7B2A49</mods:identifier>
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</mods:mods>
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<treatment id="03B63A508D67015FFF6014D7021D4691" ID-DOI="http://doi.org/10.5281/zenodo.5924271" ID-GBIF-Taxon="154474403" ID-Zenodo-Dep="5924271" LSID="urn:lsid:plazi:treatment:03B63A508D67015FFF6014D7021D4691" httpUri="http://treatment.plazi.org/id/03B63A508D67015FFF6014D7021D4691" lastPageId="32" lastPageNumber="33" pageId="26" pageNumber="27">
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<subSubSection id="C305D8CD8D670165FF6014D7057042C7" box="[151,550,692,718]" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="8BA08B468D670165FF6014D7057042C7" blockId="26.[151,550,692,751]" box="[151,550,692,718]" pageId="26" pageNumber="27">
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<heading id="D0E83C2A8D670165FF6014D7057042C7" bold="true" box="[151,550,692,718]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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||
<emphasis id="B96B57548D670165FF6014D7057042C7" bold="true" box="[151,550,692,718]" pageId="26" pageNumber="27">
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||
<taxonomicName id="4C1FF0C58D670165FF6014D7069442C7" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[151,450,692,718]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
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<emphasis id="B96B57548D670165FF6014D7069442C7" bold="true" box="[151,450,692,718]" italics="true" pageId="26" pageNumber="27">Megisthanus womersleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A258EA2F8D670165FE3E14D7057042C7" box="[457,550,692,718]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C305D8CD8D670165FF6014B5066E42E6" box="[151,312,726,751]" pageId="26" pageNumber="27" type="description">
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||
<paragraph id="8BA08B468D670165FF6014B5066E42E6" blockId="26.[151,550,692,751]" box="[151,312,726,751]" pageId="26" pageNumber="27">
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<figureCitation id="132497C38D670165FF6014B5066E42E6" box="[151,312,726,751]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="27.[151,250,1868,1890]" captionStartId-1="28.[151,250,1761,1783]" captionStartId-2="29.[151,250,974,996]" captionTargetBox-0="[157,1431,520,1838]" captionTargetBox-1="[182,1423,228,1666]" captionTargetBox-2="[170,1401,218,940]" captionTargetId-0="figure@27.[151,1436,517,1846]" captionTargetId-1="figure@28.[151,1436,193,1740]" captionTargetId-2="figure@29.[151,1436,183,953]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionText-0="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." captionText-1="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." captionText-2="FIGURE 18. Megisthanus womersleyi sp. nov. (A) female, subcapitulum and palp trochanter–femur; (B) major male, subcapitulum and palp trochanter–femur. Arrows indicate spurs." httpUri-0="https://zenodo.org/record/2600890/files/figure.png" httpUri-1="https://zenodo.org/record/2600892/files/figure.png" httpUri-2="https://zenodo.org/record/2600894/files/figure.png" pageId="26" pageNumber="27">Figures 16–18</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C305D8CD8D670165FF60157D07AE4406" pageId="26" pageNumber="27" type="materials_examined">
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||
<paragraph id="8BA08B468D670165FF60157D044F4376" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
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<emphasis id="B96B57548D670165FF60157D05BD433E" bold="true" box="[151,747,798,823]" pageId="26" pageNumber="27">
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||
Type material examined.
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||
<materialsCitation id="3B77811B8D670165FE2D157D05BD433E" ID-GBIF-Occurrence="2013950910" box="[474,747,798,823]" country="Australia" location="Queensland" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Queensland">
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<collectingCountry id="F308CBD68D670165FE2D157D051D433E" box="[474,587,798,823]" name="Australia" pageId="26" pageNumber="27">Australia</collectingCountry>
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,
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<collectingRegion id="49DB45A48D670165FDA9157D05BD433E" box="[606,747,798,823]" country="Australia" name="Queensland" pageId="26" pageNumber="27">Queensland</collectingRegion>
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</materialsCitation>
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</emphasis>
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, ex
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<taxonomicName id="4C1FF0C58D670165FCDF157D0274433E" authority="(Boisduval, 1835)" authorityName="Boisduval" authorityYear="1835" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[808,1314,798,823]" class="Insecta" family="Passallidae" genus="Protomocoelus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B96B57548D670165FCDF157D0316433F" box="[808,1088,798,822]" italics="true" pageId="26" pageNumber="27">Protomocoelus australis</emphasis>
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(Boisduval, 1835)
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</taxonomicName>
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||
.
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<materialsCitation id="3B77811B8D670165FAC3157D04D24352" ID-GBIF-Occurrence="2013950913" collectingDate="1999-07-20" collectionCode="QMS" collectorName="Iron Range & J. Skevington" country="Australia" pageId="26" pageNumber="27" specimenCode="QMS 109499" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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||
<typeStatus id="54A435E48D670165FAC3157D02CA433E" box="[1332,1436,798,823]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
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female,
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<collectorName id="26EAEE908D670165FF05152006264352" box="[242,368,835,859]" pageId="26" pageNumber="27">Iron Range</collectorName>
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NP,
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<date id="FFA1AD868D670165FE511521057F4352" box="[422,553,834,859]" pageId="26" pageNumber="27" value="1999-07-20">
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||
<collectingDate id="EFE5546E8D670165FE511521057F4352" box="[422,553,834,859]" pageId="26" pageNumber="27" value="1999-07-20">20.vii.1999</collectingDate>
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||
</date>
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,
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<collectorName id="26EAEE908D670165FDC21520059B4352" box="[565,717,834,859]" pageId="26" pageNumber="27">J. Skevington</collectorName>
|
||
(
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<specimenCode id="DBB9233D8D670165FD2A1520042A4352" box="[733,892,835,859]" collectionCode="QMS" pageId="26" pageNumber="27">QMS 109499</specimenCode>
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||
)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B77811B8D670165FC78152006134376" ID-GBIF-Occurrence="2013950911" collectingDate="1999-07-20" collectionCode="QMS" collectorName="Iron Range & J. Skevington" country="Australia" pageId="26" pageNumber="27" specimenCode="QMS 109500" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
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||
<typeStatus id="54A435E48D670165FC78152004A34352" box="[911,1013,835,859]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
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||
: 1 major male, same data as holotype (
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||
<specimenCode id="DBB9233D8D670165FF681504066B4376" box="[159,317,870,895]" collectionCode="QMS" pageId="26" pageNumber="27">QMS 109500</specimenCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B77811B8D670165FEB8150504434376" ID-GBIF-Occurrence="2013950901" box="[335,789,870,895]" collectionCode="QM" country="Australia" pageId="26" pageNumber="27" specimenCount="2" stateProvince="Queensland" typeStatus="holotype">
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||
<typeStatus id="54A435E48D670165FEB8150506E14376" box="[335,439,870,895]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
|
||
and
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||
<typeStatus id="54A435E48D670165FE19150405184376" box="[494,590,871,895]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
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||
deposited in
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<collectionCode id="ED0E13838D670165FD13150404434376" box="[740,789,871,895]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="26" pageNumber="27" type="Museum">QM</collectionCode>
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BA08B468D670165FF3015E907AE4406" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
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||
<emphasis id="B96B57548D670165FF3015E9045943AA" bold="true" box="[199,783,906,931]" pageId="26" pageNumber="27">
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||
Other material examined.
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||
<materialsCitation id="3B77811B8D670165FDE215E9045943AA" ID-GBIF-Occurrence="2013950905" box="[533,783,906,931]" collectorName="Misima Is. & Louisiade Archipelago & Rev. H. K. Bartlett" country="Papua New Guinea" location="Milne Bay Province" pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" stateProvince="Milne Bay">
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||
<collectingCountry id="F308CBD68D670165FDE215E9045C43AA" box="[533,778,906,931]" name="Papua New Guinea" pageId="26" pageNumber="27">Papua New Guinea</collectingCountry>
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||
.
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||
</materialsCitation>
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||
</emphasis>
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||
<specimenCount id="9D1940CF8D670165FCEA15E804D143AA" box="[797,903,906,931]" pageId="26" pageNumber="27" type="female">1 female</specimenCount>
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,
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<collectorName id="26EAEE908D670165FC6E15E9034E43AB" box="[921,1048,906,931]" pageId="26" pageNumber="27">Misima Is.</collectorName>
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,
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||
<collectingRegion id="49DB45A48D670165FBDC15E9024A43AA" box="[1067,1308,906,931]" country="Papua New Guinea" name="Milne Bay" pageId="26" pageNumber="27">Milne Bay Province</collectingRegion>
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,
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<collectorName id="26EAEE908D670165FAD915E9067343CE" pageId="26" pageNumber="27">Louisiade Archipelago</collectorName>
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||
,
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<collectorName id="26EAEE908D670165FEC715CC06AD43CE" box="[304,507,942,967]" pageId="26" pageNumber="27">Rev. H.K. Bartlett</collectorName>
|
||
(
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||
<accessionNumber id="944C16A58D670165FDFC15CD053443CE" box="[523,610,942,967]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17296" pageId="26" pageNumber="27" type="EnaNcbi">J17296</accessionNumber>
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||
).
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||
<materialsCitation id="3B77811B8D670165FD8215CC02CF43CE" ID-GBIF-Occurrence="2013950904" box="[629,1433,942,967]" collectorName="Misima Is. & Louisiade Archipelago & Rev. H. K. Bartlett" country="Papua New Guinea" location="Milne Bay Province" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2" stateProvince="Milne Bay">
|
||
<specimenCount id="9D1940CF8D670165FD8215CC05B343CE" box="[629,741,942,967]" pageId="26" pageNumber="27" type="female">2 females</specimenCount>
|
||
, same data (ex
|
||
<accessionNumber id="944C16A58D670165FC6415CC04B043CE" box="[915,998,943,967]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17373" pageId="26" pageNumber="27" type="EnaNcbi">J17373</accessionNumber>
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||
, slide mounted from spirit specimens)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B77811B8D670165FF6015B007A34406" ID-GBIF-Occurrence="2013950914" collectionCode="SAM" collectorName="Misima Is. & Rev. Bartlett" country="Papua New Guinea" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2">
|
||
<specimenCount id="9D1940CF8D670165FF6015B0065043E2" box="[151,262,978,1003]" pageId="26" pageNumber="27" type="female">2 females</specimenCount>
|
||
, 2 major males,
|
||
<collectorName id="26EAEE908D670165FE3715B1056C43E3" box="[448,570,978,1003]" pageId="26" pageNumber="27">Misima Is.</collectorName>
|
||
,
|
||
<collectorName id="26EAEE908D670165FDBF15B0058343E2" box="[584,725,978,1003]" pageId="26" pageNumber="27">Rev. Bartlett</collectorName>
|
||
(
|
||
<collectionCode id="ED0E13838D670165FD1215B0047543E2" box="[741,803,979,1003]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">SAM</collectionCode>
|
||
<accessionNumber id="944C16A58D670165FCDD15B1042843E2" box="[810,894,978,1003]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17295" pageId="26" pageNumber="27" type="EnaNcbi">J17295</accessionNumber>
|
||
,
|
||
<accessionNumber id="944C16A58D670165FC7D15B0048B43E2" box="[906,989,979,1003]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17297" pageId="26" pageNumber="27" type="EnaNcbi">J17297</accessionNumber>
|
||
–9,
|
||
<accessionNumber id="944C16A58D670165FBF115B1030C43E2" box="[1030,1114,978,1003]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J17364" pageId="26" pageNumber="27" type="EnaNcbi">J17364</accessionNumber>
|
||
), in very poor condition. All in
|
||
<collectionCode id="ED0E13838D670165FF43159407A34406" box="[180,245,1015,1039]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="26" pageNumber="27">SAM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D670165FF30127903D54526" pageId="26" pageNumber="27" type="diagnosis">
|
||
<paragraph id="8BA08B468D670165FF30127903D54526" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
|
||
<emphasis id="B96B57548D670165FF3012790617443A" bold="true" box="[199,321,1050,1075]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
|
||
<emphasis id="B96B57548D670165FEBF1278055C443A" box="[328,522,1051,1075]" italics="true" pageId="26" pageNumber="27">Female and male</emphasis>
|
||
. Sternal setae st4 present. Seta pd2 of femur IV on large projection. Dorsal shield finely punctate. Ventrianal shield widest posterior to anus.
|
||
<emphasis id="B96B57548D670165FCB0125D04CD445F" box="[839,923,1086,1110]" italics="true" pageId="26" pageNumber="27">Female</emphasis>
|
||
. Dorsal shield ovate, length
|
||
<quantity id="4CE726A38D670165FB18125D02C1445F" box="[1263,1431,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.875" metricValueMax="1.95" metricValueMin="1.8" pageId="26" pageNumber="27" unit="mm" value="1.875" valueMax="1.95" valueMin="1.8">1.80–1.95 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4CE726A38D670165FEA0120106AA4473" box="[343,508,1122,1147]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="26" pageNumber="27" unit="mm" value="1.0" valueMax="1.05" valueMin="0.95">0.95–1.05 mm</quantity>
|
||
, not including secondary sclerotisation. Most dorsal shield setae
|
||
<quantity id="4CE726A38D670165FB2F1201021E4472" box="[1240,1352,1122,1147]" metricMagnitude="0" metricUnit="m" metricValue="2.032" metricValueMax="2.7939999999999996" metricValueMin="1.27" pageId="26" pageNumber="27" unit="in" value="80.0" valueMax="110.0" valueMin="50.0">50–110 in</quantity>
|
||
length, with small barbs; some marginal setae thicker and longer; podonotum and opisthonotum densely setose. Sternogynal shields separate, each bearing 2 setae; internal genitalia with rudimentary latigynal element. Ventrianal shield moderately reniform.
|
||
<emphasis id="B96B57548D670165FE1512AC053C44EF" box="[482,618,1230,1254]" italics="true" pageId="26" pageNumber="27">Major male</emphasis>
|
||
. Broader than female, dorsal shield length
|
||
<quantity id="4CE726A38D670165FB9E12AC038544EF" box="[1129,1235,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="26" pageNumber="27" unit="mm" value="2.9">2.90 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D670165FADA12AD02CE44EF" box="[1325,1432,1230,1255]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="26" pageNumber="27" unit="mm" value="1.75">1.75 mm</quantity>
|
||
, without well-developed setae; setae on soft cuticle laterad dorsal shield short, length 25–70; 19 setae around genital opening; corniculi without medial inner tooth; setae h1–3 smooth, palp coxal seta barbed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D67015FFF301359043A416C" lastPageId="32" lastPageNumber="33" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="8BA08B468D670165FF3013590784469E" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
|
||
<emphasis id="B96B57548D670165FF3013590604455A" bold="true" box="[199,338,1338,1363]" pageId="26" pageNumber="27">Description</emphasis>
|
||
(Australian specimens).
|
||
<emphasis id="B96B57548D670165FD831359059D455A" bold="true" box="[628,715,1338,1363]" italics="true" pageId="26" pageNumber="27">Female</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D670165FD2A13580461455B" box="[733,823,1339,1362]" italics="true" pageId="26" pageNumber="27">Dorsum</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D670165FCBD13590488455A" box="[842,990,1338,1363]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="26" pageNumber="27">Figs 16A, C</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D670165FC1A13580376455A" box="[1005,1056,1339,1363]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">17C</figureCitation>
|
||
). Idiosomal length ca.
|
||
<quantity id="4CE726A38D670165FAD9135902C1455B" box="[1326,1431,1338,1363]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="26" pageNumber="27" unit="mm" value="2.6">2.60 mm</quantity>
|
||
, width ca.
|
||
<quantity id="4CE726A38D670165FEFC133C0622457F" box="[267,372,1375,1399]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="26" pageNumber="27" unit="mm" value="1.9">1.90 mm</quantity>
|
||
, body length (including
|
||
<taxonomicName id="4C1FF0C58D670165FD65133D044A457E" authorityName="Krefft" authorityYear="1871" box="[658,796,1374,1399]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="26" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
, to tip of gnathotectum)
|
||
<quantity id="4CE726A38D670165FBC8133C03FF457F" box="[1087,1193,1375,1399]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="26" pageNumber="27" unit="mm" value="3.2">3.20 mm</quantity>
|
||
. Dorsal shield ovate, length
|
||
<quantity id="4CE726A38D670165FF1313E1061F4593" box="[228,329,1410,1435]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="mm" value="2.5">2.50 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D670165FE6C13E005574593" box="[411,513,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="26" pageNumber="27" unit="mm" value="1.4">1.40 mm</quantity>
|
||
; secondary sclerotisation weakly formed but present laterally and posteriorly (
|
||
<figureCitation id="132497C38D670165FA9D13E107B845B6" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="26" pageNumber="27">Figs 16A, C</figureCitation>
|
||
); maximum length of dorsal shield including secondary sclerotisation
|
||
<quantity id="4CE726A38D670165FBED13C503D545B7" box="[1050,1155,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="26" pageNumber="27" unit="mm" value="2.55">2.55 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D670165FB2C13C5021245B7" box="[1243,1348,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="26" pageNumber="27" unit="mm" value="1.45">1.45 mm</quantity>
|
||
. Dorsal shield hypertrichous, densely setose excepting narrow band of bare cuticle posteriorly just within margin. Dorsal shield setae with small barbs, moderately short, length 50–110 (
|
||
<figureCitation id="132497C38D670165FC83138D0489460E" box="[884,991,1518,1543]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="26" pageNumber="27">Figs 16C</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D670165FC1A138C0349460E" box="[1005,1055,1519,1543]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">17C</figureCitation>
|
||
), excepting few anterolateral and posterolateral setae (up to 275); posterior margin with large setal sockets suggesting very long setae are present but broken off. Dorsal shield cuticle appearing smooth, with inconspicuous fine punctation. Setae in soft cuticle barbed, on small platelets, length 50–100, excepting some posterolateral setae close to dorsal shield long (up to 720).
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D670165FF3010C004E9470A" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
|
||
<emphasis id="B96B57548D670165FF3010C0065A46B3" box="[199,268,1699,1722]" italics="true" pageId="26" pageNumber="27">Venter</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D670165FED710C106DA46B2" box="[288,396,1698,1723]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="26" pageNumber="27">Figs 16B</figureCitation>
|
||
,
|
||
<figureCitation id="132497C38D670165FE6D10C006A646B2" box="[410,496,1699,1723]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">17A, B</figureCitation>
|
||
). Setae st1 broken. Sternal shield (
|
||
<figureCitation id="132497C38D670165FC7410C104BD46B2" box="[899,1003,1698,1723]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">Fig. 17A</figureCitation>
|
||
) reticulate, with 3 pairs of setae, 1–2 pairs of pores opening ventrally, 1–3 pores opening laterally, and one pair of lyrifissures; setae st2–3 sparsely barbed, st4 smooth, st2 ca. 100, st3 100, st4 55; shield medial length 85.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D670165FF30116D032D47BE" blockId="26.[151,1437,798,2011]" pageId="26" pageNumber="27">
|
||
Sternogynal shields (
|
||
<figureCitation id="132497C38D670165FE46116D054C472E" box="[433,538,1806,1831]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">Figs 17A</figureCitation>
|
||
) free posteriorly, genital opening length 260, width 320, each sternogynal shield length 240, weakly lineate-reticulate, each bearing 2 setae and one lyrifissure; anterior seta similar in form and length to other setae (20–25), smooth. Internal genitalia comprises three obvious elements (
|
||
<emphasis id="B96B57548D670165FB29113402564767" box="[1246,1280,1879,1902]" italics="true" pageId="26" pageNumber="27">mg</emphasis>
|
||
,
|
||
<emphasis id="B96B57548D670165FAE3113402784767" box="[1300,1326,1879,1902]" italics="true" pageId="26" pageNumber="27">va</emphasis>
|
||
,
|
||
<emphasis id="B96B57548D670165FAB41135020B4767" box="[1347,1373,1878,1902]" italics="true" pageId="26" pageNumber="27">vh</emphasis>
|
||
) and rudimentary latigynal (
|
||
<emphasis id="B96B57548D670165FE57111806E0479A" box="[416,438,1915,1939]" italics="true" pageId="26" pageNumber="27">lg</emphasis>
|
||
) elements (
|
||
<figureCitation id="132497C38D670165FDB7111905FC479A" box="[576,682,1914,1939]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="26" pageNumber="27">Fig. 17B</figureCitation>
|
||
); rudimentary latigynal elements fused with thickened margin of sternogynal shield, with small porose area near attachment of
|
||
<emphasis id="B96B57548D670165FCBB11FC043047BF" box="[844,870,1951,1974]" italics="true" pageId="26" pageNumber="27">va</emphasis>
|
||
with sternogynal shield.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D670164FF3011A104EB40F1" blockId="26.[151,1437,798,2011]" lastBlockId="27.[151,1437,151,464]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
Ventral shield (
|
||
<figureCitation id="132497C38D670165FE8411A1068147D2" box="[371,471,1986,2011]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="26" pageNumber="27">Fig. 16B</figureCitation>
|
||
) extends 1040 posteriorly from posterior margin of genital opening; maximum width behind coxa IV 430; shield with 13–15 pairs of setae around shield margin, one pair posteriorly within shield, and without setae off shield margin close to sternogynal shield; some setae expressed asymmetrically; setae smooth, length 40–100; shield reticulate posteriorly, otherwise lineate-reticulate.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D660164FF3017670565416C" blockId="27.[151,1437,151,464]" pageId="27" pageNumber="28">
|
||
Ventrianal shield trapezoidal (
|
||
<figureCitation id="132497C38D660164FDEF176705284114" box="[536,638,260,285]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="27" pageNumber="28">Fig. 16B</figureCitation>
|
||
), with anus in anterior half of shield; anterior width ca. 400, maximum width posterior to anus, 600, length 400; shield reticulate anteriorly, reticulation evanescent posteriorly; 16 loosely paired, smooth setae, length 75–120.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D660164FF30170C034541D9" blockId="27.[151,1437,151,464]" pageId="27" pageNumber="28">
|
||
Peritrematal shield hypertrichous, post-peritrematal region extends 760 behind coxa IV into metapodal region (
|
||
<figureCitation id="132497C38D660164FF6817F7065141A4" box="[159,263,404,429]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="27" pageNumber="28">Fig. 16B</figureCitation>
|
||
); setae laterad legs smooth, length 40–60, those directly above legs III–IV not much shorter than other setae; three pairs of long setae on anterior margin of peritrematal shield broken.
|
||
</paragraph>
|
||
<caption id="DF60DBCE8D660164FF60112F03B44796" httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,250,1868,1890]" targetBox="[157,1431,520,1838]" targetPageId="27">
|
||
<paragraph id="8BA08B468D660164FF60112F03B44796" blockId="27.[151,1436,1868,1951]" pageId="27" pageNumber="28">
|
||
<emphasis id="B96B57548D660164FF60112F06734768" bold="true" box="[151,293,1868,1890]" pageId="27" pageNumber="28">FIGURE 16.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D660164FEC6112F0567476B" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[305,561,1868,1890]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
|
||
<emphasis id="B96B57548D660164FEC6112F0567476B" box="[305,561,1868,1890]" italics="true" pageId="27" pageNumber="28">Megisthanus womersleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D660164FDC9112F05C24768" bold="true" box="[574,660,1868,1889]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="A258EA2F8D660164FDC9112F05C24768" box="[574,660,1868,1889]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D)
|
||
<taxonomicName id="4C1FF0C58D660164FB46110903BA4789" box="[1201,1260,1898,1920]" pageId="27" pageNumber="28" rank="species" species="major">major</taxonomicName>
|
||
male, chelicera, showing dendritic excrescences characteristic for
|
||
<taxonomicName id="4C1FF0C58D660164FD7811EA04714796" baseAuthorityName="Womersley" baseAuthorityYear="1937" box="[655,807,1929,1951]" class="Arachnida" family="Megisthanidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Megisthanidae</taxonomicName>
|
||
. Scale bars: 200 µm (A–B); 100 µm (C–D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF60DBCE8D610163FF6010820310475A" httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1761,1783]" targetBox="[182,1423,228,1666]" targetPageId="28">
|
||
<paragraph id="8BA08B468D610163FF6010820310475A" blockId="28.[151,1436,1761,1875]" pageId="28" pageNumber="29">
|
||
<emphasis id="B96B57548D610163FF601082067746FF" bold="true" box="[151,289,1761,1783]" pageId="28" pageNumber="29">FIGURE 17.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D610163FEDE1082057346FE" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[297,549,1761,1783]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
|
||
<emphasis id="B96B57548D610163FEDE1082057346FE" box="[297,549,1761,1783]" italics="true" pageId="28" pageNumber="29">Megisthanus womersleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D610163FDDA1082052946FF" bold="true" box="[557,639,1761,1782]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A258EA2F8D610163FDDA1082052946FF" box="[557,639,1761,1782]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF60DBCE8D600162FF6015AD0474440B" httpUri="https://zenodo.org/record/2600894/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,974,996]" targetBox="[170,1401,218,940]" targetPageId="29">
|
||
<paragraph id="8BA08B468D600162FF6015AD0474440B" blockId="29.[151,1436,974,1026]" pageId="29" pageNumber="30">
|
||
<emphasis id="B96B57548D600162FF6015AD067343EA" bold="true" box="[151,293,974,996]" pageId="29" pageNumber="30">FIGURE 18.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D600162FEC515AD056543ED" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[306,563,974,996]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
|
||
<emphasis id="B96B57548D600162FEC515AD056543ED" box="[306,563,974,996]" italics="true" pageId="29" pageNumber="30">Megisthanus womersleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EA2F8D600162FDB715AD05C143EA" box="[576,663,974,995]" pageId="29" pageNumber="30" rank="species">
|
||
<emphasis id="B96B57548D600162FDB715AD05C143EA" bold="true" box="[576,663,974,995]" pageId="29" pageNumber="30">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(A) female, subcapitulum and palp trochanter–femur; (B)
|
||
<taxonomicName id="4C1FF0C58D600162FAEB15AD020143ED" box="[1308,1367,974,996]" pageId="29" pageNumber="30" rank="species" species="major">major</taxonomicName>
|
||
male, subcapitulum and palp trochanter–femur. Arrows indicate spurs.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA08B468D600162FF30124C026744D1" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C1FF0C58D600162FF30124C0600444E" authorityName="Krefft" authorityYear="1871" box="[199,342,1071,1095]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="29" pageNumber="30" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B96B57548D600162FF30124C0600444E" box="[199,342,1071,1095]" italics="true" pageId="29" pageNumber="30">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132497C38D600162FE92124C069B4441" box="[357,461,1071,1096]" captionStart="FIGURE 18" captionStartId="29.[151,250,974,996]" captionTargetBox="[170,1401,218,940]" captionTargetId="figure@29.[151,1436,183,953]" captionTargetPageId="29" captionText="FIGURE 18. Megisthanus womersleyi sp. nov. (A) female, subcapitulum and palp trochanter–femur; (B) major male, subcapitulum and palp trochanter–femur. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600894/files/figure.png" pageId="29" pageNumber="30">Fig. 18A</figureCitation>
|
||
). Gnathotectum reticulate, medial length 700. Subcapitulum with setae h1 285, h2 180, h3 broken, pc 175, all setae sparsely barbed; medial subcapitulum with some fine striae, hypostome smooth. Corniculi length 265 with medial tooth. Palp trochanter with small ventro-distal spur, length 10–15, palp femur without distal spur. Chelicerae (
|
||
<figureCitation id="132497C38D600162FE0E12F8050944BD" box="[505,607,1179,1204]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="29" pageNumber="30">Fig. 16D</figureCitation>
|
||
) with entire fixed digit length 720, movable digit length 405. Morphology as per genus description. Fixed digit with 13 teeth; cheliceral seta length 30. Movable digit with 12 teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D600162FF30128706504585" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">
|
||
<emphasis id="B96B57548D600162FF30128707AA44F2" box="[199,252,1252,1275]" italics="true" pageId="29" pageNumber="30">Legs</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D600162FEE71280062844F5" box="[272,382,1251,1276]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="29" pageNumber="30">Fig. 17D</figureCitation>
|
||
). Setation as per genus description. Selected setal measurements: genu II setae ad1, pd1 subequal, length 140–150; seta pd1 on genu III long, 425, much longer than ad1, 185, tarsus III ad3 moderately long, 250, femur IV pd1 long,> 360, genu IV ad1 very long, ca. 600, pd1 broken, tibia IV pd1 long, 570 and tarsus IV ad4 long, 360, pd3 long, 370. Femur IV with large projections bearing setae pd2, ad3, small projection bearing setae ad2.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D600162FF3013F403424649" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">
|
||
<emphasis id="B96B57548D600162FF3013F4065545B9" bold="true" box="[199,259,1431,1456]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
|
||
(major) (n = 1).
|
||
<emphasis id="B96B57548D600162FE3813FB057F45A6" box="[463,553,1432,1455]" italics="true" pageId="29" pageNumber="30">Dorsum</emphasis>
|
||
. Idiosomal length ca. 3.00 mm, width ca.
|
||
<quantity id="4CE726A38D600162FBE913FB03D145A6" box="[1054,1159,1432,1456]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="mm" value="2.1">2.10 mm</quantity>
|
||
, body length (including
|
||
<taxonomicName id="4C1FF0C58D600162FF6013D8067745DD" authorityName="Krefft" authorityYear="1871" box="[151,289,1467,1492]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="29" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
, to tip of gnathotectum) ca.
|
||
<quantity id="4CE726A38D600162FDAC13DF059445DA" box="[603,706,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="29" pageNumber="30" unit="mm" value="3.7">3.70 mm</quantity>
|
||
. Dorsal shield ovate, length
|
||
<quantity id="4CE726A38D600162FBF713DF033045DA" box="[1024,1126,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="29" pageNumber="30" unit="mm" value="2.9">2.90 mm</quantity>
|
||
, width
|
||
<quantity id="4CE726A38D600162FB4F13D8024945DA" box="[1208,1311,1467,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="29" pageNumber="30" unit="mm" value="1.75">1.75 mm</quantity>
|
||
; secondary sclerotisation present but weakly formed. Dorsum hypertrichous. Setae on dorsal shield tiny, smooth, length <5. Setae in soft membrane close to dorsal shield smooth, becoming more barbed and longer laterally, length 25–70; some setae near posterolateral and posterior margin of shield also long (to 120).
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D600162FF30102F039E4735" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">
|
||
<emphasis id="B96B57548D600162FF30102F065A466A" box="[199,268,1612,1635]" italics="true" pageId="29" pageNumber="30">Venter</emphasis>
|
||
(
|
||
<figureCitation id="132497C38D600162FEEA10280626466D" box="[285,368,1611,1636]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="29" pageNumber="30">Figs 17</figureCitation>
|
||
E–F). Setae st1 barbed, length 150. Sternoventral shield reticulate, length from posterior margin of genital opening 1190. Sternal setae st2 and st3 barbed, length 130; setae st4 indistinguishable among other sternoventral setae; sternoventral shield (
|
||
<figureCitation id="132497C38D600162FD9710F0059746A5" box="[608,705,1683,1708]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="29" pageNumber="30">Fig. 17E</figureCitation>
|
||
) hypertrichous posterior to setae st3, with 19 setae around genital opening; posterior sternoventral shield with setae along margin; these setae smooth or sparsely barbed, shortest setae 70 (mesad coxa IV) and longest setae 140 (anterolaterad genital opening); posterior sternoventral shield (
|
||
<figureCitation id="132497C38D600162FA9810B807954711" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="29" pageNumber="30">Fig. 17F</figureCitation>
|
||
) with two large discs of porose cuticle (diameter 190), each bearing an anterior and posterior pair of unmodified pair of smooth setae 25–30, and a highly modified pair of setae centrally, length 50.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D600162FF3011240408478D" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">
|
||
Genital opening (
|
||
<figureCitation id="132497C38D600162FE67112406A14769" box="[400,503,1863,1888]" captionStart="FIGURE 17" captionStartId="28.[151,250,1761,1783]" captionTargetBox="[182,1423,228,1666]" captionTargetId="figure@28.[151,1436,193,1740]" captionTargetPageId="28" captionText="FIGURE 17. Megisthanus womersleyi sp. nov. female. (A) sternogenital region; (B) internal genitalia; (C) female, posterior dorsal shield, large setae broken (large sockets along margin remain), note submarginal porose band; (D) female, femur IV; (E) major male, sternogenital region; (F) major male, posterior margin of ventral shield. Abbreviations: g1 = anterior pair of genital setae; lg = latigynal remnant; mg = mesogynal shield; va = vaginal arms; vh = vaginal heads." httpUri="https://zenodo.org/record/2600892/files/figure.png" pageId="29" pageNumber="30">Fig. 17E</figureCitation>
|
||
) length 125, width 115, surrounded by a ring of heavily sclerotised cuticle; this cuticle with 11–12 loosely paired pores laterad genital opening.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D600162FF3011EC060A47F9" blockId="29.[151,1437,1071,2032]" pageId="29" pageNumber="30">Ventrianal shield a curved trapezoid, length 500, anterior width ca. 430, maximum width 660, widest posterolaterally, well behind anus; shield reticulate, with 12 loosely paired setae. Peritreme and peritrematal shield similar to female.</paragraph>
|
||
<caption id="DF60DBCE8D630161FF6011DD03A647FB" httpUri="https://zenodo.org/record/2600896/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1982,2004]" targetBox="[207,1374,193,1944]" targetPageId="30">
|
||
<paragraph id="8BA08B468D630161FF6011DD03A647FB" blockId="30.[151,1436,1982,2034]" pageId="30" pageNumber="31">
|
||
<emphasis id="B96B57548D630161FF6011DD064947DA" bold="true" box="[151,287,1982,2004]" pageId="30" pageNumber="31">FIGURE 19.</emphasis>
|
||
Photographs of the holotypes of
|
||
<taxonomicName id="4C1FF0C58D630161FD9811DD04B847DD" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" box="[623,1006,1982,2004]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D630161FD9811DD040A47DD" box="[623,860,1982,2004]" italics="true" pageId="30" pageNumber="31">Megisthanus doreianus</emphasis>
|
||
Thorell, 1882
|
||
</taxonomicName>
|
||
, (A) dorsum; (B) venter; and
|
||
<taxonomicName id="4C1FF0C58D630161FAEC11DD06E247FB" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="hatamanensis">
|
||
<emphasis id="B96B57548D630161FAEC11DD067447FB" italics="true" pageId="30" pageNumber="31">Megisthanus hatamanensis</emphasis>
|
||
Thorell, 1882
|
||
</taxonomicName>
|
||
, (C) dorsum; (D) venter. Photographs by Geoff Thompson, Queensland Museum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF60DBCE8D620160FF6011C603BA47D3" httpUri="https://zenodo.org/record/2600898/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1957,1979]" targetBox="[241,1339,193,1913]" targetPageId="31">
|
||
<paragraph id="8BA08B468D620160FF6011C603BA47D3" blockId="31.[151,1436,1957,2010]" pageId="31" pageNumber="32">
|
||
<emphasis id="B96B57548D620160FF6011C6067547B3" bold="true" box="[151,291,1957,1979]" pageId="31" pageNumber="32">FIGURE 20.</emphasis>
|
||
The holotype of
|
||
<taxonomicName id="4C1FF0C58D620160FE2911C6040D47B2" authority="Karg, 1997" authorityName="Karg" authorityYear="1997" box="[478,859,1957,1979]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D620160FE2911C6058A47B2" box="[478,732,1957,1979]" italics="true" pageId="31" pageNumber="32">Megisthanus postsetosus</emphasis>
|
||
Karg, 1997
|
||
</taxonomicName>
|
||
(ZMB 45476). (A) photograph; (B) detail of posterior idiosoma, drawn from photograph. Photograph by Anja Friederichs from the Museum für Naturkunde Berlin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA08B468D5D015FFF3016FB03FC4149" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
<taxonomicName id="4C1FF0C58D5D015FFF3016FB060040B9" authorityName="Krefft" authorityYear="1871" box="[199,342,152,176]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="32" pageNumber="33" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFF3016FB060040B9" box="[199,342,152,176]" italics="true" pageId="32" pageNumber="33">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132497C38D5D015FFE9216F4069A40B9" box="[357,460,151,176]" captionStart="FIGURE 18" captionStartId="29.[151,250,974,996]" captionTargetBox="[170,1401,218,940]" captionTargetId="figure@29.[151,1436,183,953]" captionTargetPageId="29" captionText="FIGURE 18. Megisthanus womersleyi sp. nov. (A) female, subcapitulum and palp trochanter–femur; (B) major male, subcapitulum and palp trochanter–femur. Arrows indicate spurs." httpUri="https://zenodo.org/record/2600894/files/figure.png" pageId="32" pageNumber="33">Fig. 18B</figureCitation>
|
||
). Much larger than female
|
||
<taxonomicName id="4C1FF0C58D5D015FFD0C16F404D040B9" authorityName="Krefft" authorityYear="1871" box="[763,902,151,176]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="32" pageNumber="37" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
. Gnathotectum similar to female, medial length 910; subcapitulum similar, except with large submedial projections (inner length 75). Setae h1 smooth, blade-like, 250; setae h2 smooth, 140, setae h3 smooth, 215, palp coxal seta barbed, 170. Corniculi without tooth-like processes, length 410. Palp trochanter with distoventral cusps on trochanter (length 30) and femur (length 15–20). Chelicerae similar to female except fixed digit with 17 teeth and movable digit with 13 teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5D015FFF30172E043A416C" blockId="32.[151,1437,151,1688]" box="[199,876,332,357]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96B57548D5D015FFF30172E07AA416D" box="[199,252,333,356]" italics="true" pageId="32" pageNumber="33">Legs</emphasis>
|
||
. Similar to female except distal spurs and spines larger.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D5D015FFF30170C027741D9" pageId="32" pageNumber="33" type="diagnosis">
|
||
<paragraph id="8BA08B468D5D015FFF30170C027741D9" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96B57548D5D015FFF30170C06874181" bold="true" box="[199,465,367,392]" pageId="32" pageNumber="33">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C1FF0C58D5D015FFE2B171305A34181" authorityName="Seeman" authorityYear="2019" box="[476,757,368,392]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="womersleyi">
|
||
<emphasis id="B96B57548D5D015FFE2B171305A34181" box="[476,757,368,392]" italics="true" pageId="32" pageNumber="33">Megisthanus womersleyi</emphasis>
|
||
</taxonomicName>
|
||
is most similar to five species of
|
||
<taxonomicName id="4C1FF0C58D5D015FFB7B1713024D4181" authorityName="Thorell" authorityYear="1882" box="[1164,1307,368,392]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFB7B1713024D4181" box="[1164,1307,368,392]" italics="true" pageId="32" pageNumber="33">Megisthanus</emphasis>
|
||
</taxonomicName>
|
||
from New
|
||
<collectingCountry id="F308CBD68D5D015FFF6017F707BD41A4" box="[151,235,404,429]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
, as discussed below.
|
||
<taxonomicName id="4C1FF0C58D5D015FFE2917F705A341A5" authorityName="Seeman" authorityYear="2019" box="[478,757,404,428]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="womersleyi">
|
||
<emphasis id="B96B57548D5D015FFE2917F705A341A5" box="[478,757,404,428]" italics="true" pageId="32" pageNumber="33">Megisthanus womersleyi</emphasis>
|
||
</taxonomicName>
|
||
can be most easily distinguished from these species by the density of its dorsal setation, which is far greater than all other species from
|
||
<collectingCountry id="F308CBD68D5D015FFC1817D4030041D9" box="[1007,1110,439,464]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
|
||
and New
|
||
<collectingCountry id="F308CBD68D5D015FFB3F17D4024B41D9" box="[1224,1309,439,464]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D5D015FFF3017BF04144659" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8BA08B468D5D015FFF3017BF043D42A1" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96B57548D5D015FFF3017BF066D41FC" bold="true" box="[199,315,476,501]" pageId="32" pageNumber="33">Remarks.</emphasis>
|
||
With the description of
|
||
<taxonomicName id="4C1FF0C58D5D015FFDB817BE05A341FD" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[591,757,476,500]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
|
||
<emphasis id="B96B57548D5D015FFDB817BE053041FD" box="[591,614,477,500]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD8117BF05A341FD" box="[630,757,476,500]" italics="true" pageId="32" pageNumber="33">womersleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D5D015FFD0A17BF040E41FC" bold="true" box="[765,856,476,501]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A258EA2F8D5D015FFD0A17BF040E41FC" box="[765,856,476,501]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, six species are now known from New
|
||
<collectingCountry id="F308CBD68D5D015FFAED17BF023841FC" box="[1306,1390,476,501]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
, the others being:
|
||
<taxonomicName id="4C1FF0C58D5D015FFED9146305834211" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" box="[302,725,511,536]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFED9146305654211" box="[302,563,512,536]" italics="true" pageId="32" pageNumber="33">Megisthanus doreianus</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFDCD179C05834211" author="Thorell, T." box="[570,725,511,536]" pageId="32" pageNumber="33" pagination="21 - 69" refId="ref22896" refString="Thorell, T. (1882) Descrizione di alcuni Arachnidi inferiori dell'Archipelago Malese. Annali del Museo civico di storia natural di Genova, 18, 21 - 69. pls. 6." type="journal article" year="1882">Thorell, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFD28146203724211" authority="Thorell, 1882" authorityName="Thorell" authorityYear="1882" box="[735,1060,511,536]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatamensis">
|
||
<emphasis id="B96B57548D5D015FFD28146205A04211" box="[735,758,513,536]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFCF3146304D44211" box="[772,898,512,536]" italics="true" pageId="32" pageNumber="33">hatamensis</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFC7E179C03724211" author="Thorell, T." box="[905,1060,511,536]" pageId="32" pageNumber="33" pagination="21 - 69" refId="ref22896" refString="Thorell, T. (1882) Descrizione di alcuni Arachnidi inferiori dell'Archipelago Malese. Annali del Museo civico di storia natural di Genova, 18, 21 - 69. pls. 6." type="journal article" year="1882">Thorell, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFBD9146202CE4211" authority="Oudemans, 1905" authorityName="Oudemans" authorityYear="1905" box="[1070,1432,511,536]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moaifensus">
|
||
<emphasis id="B96B57548D5D015FFBD9146203134211" box="[1070,1093,513,536]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFBA4146303874211" box="[1107,1233,512,536]" italics="true" pageId="32" pageNumber="33">moaifensus</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFB2F179C02CE4211" author="Oudemans, A. C." box="[1240,1432,511,536]" pageId="32" pageNumber="33" pagination="222 - 226" refId="ref22558" refString="Oudemans, A. C. (1905) Acarologische Aanteekeningen XVII. Entomologische Berichten, 1, 222 - 226. https: // doi. org / 10.5962 / bhl. part. 1124" type="journal article" year="1905">Oudemans, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFF60144606A24234" authority="Oudemans, 1905" authorityName="Oudemans" authorityYear="1905" box="[151,500,548,573]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B96B57548D5D015FFF60144607F84235" box="[151,174,549,572]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFF491447067F4235" box="[190,297,548,572]" italics="true" pageId="32" pageNumber="33">orientalis</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFEC5144706A24234" author="Oudemans, A. C." box="[306,500,548,573]" pageId="32" pageNumber="33" pagination="222 - 226" refId="ref22558" refString="Oudemans, A. C. (1905) Acarologische Aanteekeningen XVII. Entomologische Berichten, 1, 222 - 226. https: // doi. org / 10.5962 / bhl. part. 1124" type="journal article" year="1905">Oudemans, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFDF5144604354235" authority="Womersley, 1937" authorityName="Womersley" authorityYear="1937" box="[514,867,548,573]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="papuanus">
|
||
<emphasis id="B96B57548D5D015FFDF51446054F4235" box="[514,537,549,572]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFDDF144605C34235" box="[552,661,549,572]" italics="true" pageId="32" pageNumber="33">papuanus</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFD69144704354235" author="Womersley, H." box="[670,867,548,573]" pageId="32" pageNumber="33" pagination="175 - 180" refId="ref22977" refString="Womersley, H. (1937) Australian Acarina of the genus Megisthanus Thorell. Transactions of the Royal Society of South Australia, 61, 175 - 180." type="journal article" year="1937">Womersley, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFC6A144603994235" authority="Karg, 1997" authorityName="Karg" authorityYear="1997" box="[925,1231,548,573]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D5D015FFC6A144604E24235" box="[925,948,549,572]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC33144603124235" box="[964,1092,549,572]" italics="true" pageId="32" pageNumber="33">postsetosus</emphasis>
|
||
<bibRefCitation id="EF8EF6B78D5D015FFBBA144603994235" author="Karg, W." box="[1101,1231,548,573]" pageId="32" pageNumber="33" pagination="51 - 61" refId="ref22083" refString="Karg, W. (1997) Neue Raubmilbenarten der Antennophorina Berlese, 1892 (Acarina, Parasitiformes), phoretic on passalid beetles. Mitteilungen aus dem Zoologischen Museum in Berlin, 73, 51 - 61." type="journal article" year="1997">Karg, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
. All these species belong to a species group, previously found only in New
|
||
<collectingCountry id="F308CBD68D5D015FFCEF1424043A4269" box="[792,876,583,608]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
, which is defined by a large dorsal tubercle on leg IV that carries seta pd2. Unlike all other Australian species, these species also have eroded medial margins of the sternogynal shield, so that they appear to form C-shaped shields.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5D015FFF3014D7064D43E5" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
<bibRefCitation id="EF8EF6B78D5D015FFF3014D706CF42C5" author="Womersley, H." box="[199,409,692,717]" pageId="32" pageNumber="33" pagination="175 - 180" refId="ref22977" refString="Womersley, H. (1937) Australian Acarina of the genus Megisthanus Thorell. Transactions of the Royal Society of South Australia, 61, 175 - 180." type="journal article" year="1937">Womersley (1937)</bibRefCitation>
|
||
originally identified the specimens from Misima Island as
|
||
<taxonomicName id="4C1FF0C58D5D015FFBB314D6038D42C5" authorityName="Thorell" authorityYear="1882" box="[1092,1243,692,716]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFBB314D6030D42C5" box="[1092,1115,693,716]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFB9B14D7038D42C5" box="[1132,1243,692,716]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
. To determine if Womersley’s original decision was correct, I examined the
|
||
<typeStatus id="54A435E48D5D015FFC9D14B4048142F9" box="[874,983,727,752]" pageId="32" pageNumber="33" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1FF0C58D5D015FFBFB14BA03F142F9" authorityName="Thorell" authorityYear="1882" box="[1036,1191,728,752]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFBFB14BA037542F9" box="[1036,1059,729,752]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFBCE14BB03F142F9" box="[1081,1191,728,752]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFB4A14BA023E42F9" authorityName="Thorell" authorityYear="1882" box="[1213,1384,728,752]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatamensis">
|
||
<emphasis id="B96B57548D5D015FFB4A14BA038242F9" box="[1213,1236,729,752]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFB1D14BB023E42F9" box="[1258,1384,728,752]" italics="true" pageId="32" pageNumber="33">hatamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFA8A14BA0652431D" authorityName="Womersley" authorityYear="1937" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="papuanus">
|
||
<emphasis id="B96B57548D5D015FFA8A14BA02C242F9" box="[1405,1428,729,752]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFF60149E0652431D" box="[151,260,765,788]" italics="true" pageId="32" pageNumber="33">papuanus</emphasis>
|
||
</taxonomicName>
|
||
, photographs of the
|
||
<typeStatus id="54A435E48D5D015FFE18149F0507431C" box="[495,593,764,789]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1FF0C58D5D015FFD8F149E044B431D" authorityName="Karg" authorityYear="1997" box="[632,797,765,788]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D5D015FFD8F149E05D9431D" box="[632,655,765,788]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD69149E044B431D" box="[670,797,765,788]" italics="true" pageId="32" pageNumber="33">postsetosus</emphasis>
|
||
</taxonomicName>
|
||
, and the drawings of Oudemans’ species
|
||
<taxonomicName id="4C1FF0C58D5D015FFB00149E02CA431D" authorityName="Oudemans" authorityYear="1905" box="[1271,1436,764,788]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moaifensus">
|
||
<emphasis id="B96B57548D5D015FFB00149E0258431D" box="[1271,1294,765,788]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFAE9149F02CA431D" box="[1310,1436,764,788]" italics="true" pageId="32" pageNumber="33">moaifensus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFF3F1542060C4331" authorityName="Oudemans" authorityYear="1905" box="[200,346,800,824]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B96B57548D5D015FFF3F154207894331" box="[200,223,801,824]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFF181543060C4331" box="[239,346,800,824]" italics="true" pageId="32" pageNumber="33">orientalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C1FF0C58D5D015FFE9D154305394331" authorityName="Womersley" authorityYear="1937" box="[362,623,800,824]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="papuanus">
|
||
<emphasis id="B96B57548D5D015FFE9D154305394331" box="[362,623,800,824]" italics="true" pageId="32" pageNumber="33">Megisthanus papuanus</emphasis>
|
||
</taxonomicName>
|
||
is clearly different from all other species of
|
||
<taxonomicName id="4C1FF0C58D5D015FFB87154303A94331" authorityName="Thorell" authorityYear="1882" box="[1136,1279,800,824]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFB87154303A94331" box="[1136,1279,800,824]" italics="true" pageId="32" pageNumber="33">Megisthanus</emphasis>
|
||
</taxonomicName>
|
||
, as explained under the separate treatment for that species. However, all the other species are similar to those of Womersley’s specimens identified as
|
||
<taxonomicName id="4C1FF0C58D5D015FFE47150A051E4389" authorityName="Thorell" authorityYear="1882" box="[432,584,872,896]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFE47150A06914389" box="[432,455,873,896]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFE2E150B051E4389" box="[473,584,872,896]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
(i.e.
|
||
<taxonomicName id="4C1FF0C58D5D015FFD7D150A04624389" authority="Seeman, 2019" authorityName="Seeman" authorityYear="2019" box="[650,820,872,896]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="womersleyi" status="sp. nov.">
|
||
<emphasis id="B96B57548D5D015FFD7D150A05F74389" box="[650,673,873,896]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD43150B04624389" box="[692,820,872,896]" italics="true" pageId="32" pageNumber="33">womersleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96B57548D5D015FFCC8150404CB4389" bold="true" box="[831,925,871,896]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A258EA2F8D5D015FFCC8150404CB4389" box="[831,925,871,896]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), yet his specimens differ in one important aspect: the density of dorsal setation. Womersley’s specimens have a dense pelage of setae (
|
||
<figureCitation id="132497C38D5D015FFB4E15EF027543AC" box="[1209,1315,908,933]" captionStart="FIGURE 16" captionStartId="27.[151,250,1868,1890]" captionTargetBox="[157,1431,520,1838]" captionTargetId="figure@27.[151,1436,517,1846]" captionTargetPageId="27" captionText="FIGURE 16. Megisthanus womersleyi sp. nov. photomicrographs (A) female, dorsum; (B) female, venter; (C) female, posterior dorsal shield; arrows indicate sockets of broken large setae along posterior margin; (D) major male, chelicera, showing dendritic excrescences characteristic for Megisthanidae. Scale bars: 200 µm (A–B); 100 µm (C–D)." httpUri="https://zenodo.org/record/2600890/files/figure.png" pageId="32" pageNumber="33">Fig. 16A</figureCitation>
|
||
), while all other species have considerably sparser covering setae (
|
||
<figureCitation id="132497C38D5D015FFCC215CC04EB43C1" box="[821,957,943,968]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStartId-0="30.[151,250,1982,2004]" captionStartId-1="31.[151,250,1957,1979]" captionTargetBox-0="[207,1374,193,1944]" captionTargetBox-1="[241,1339,193,1913]" captionTargetId-0="figure@30.[206,1374,193,1964]" captionTargetId-1="figure@31.[241,1342,193,1932]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 19. Photographs of the holotypes of Megisthanus doreianus Thorell, 1882, (A) dorsum; (B) venter; and Megisthanus hatamanensis Thorell, 1882, (C) dorsum; (D) venter. Photographs by Geoff Thompson, Queensland Museum." captionText-1="FIGURE 20. The holotype of Megisthanus postsetosus Karg, 1997 (ZMB 45476). (A) photograph; (B) detail of posterior idiosoma, drawn from photograph. Photograph by Anja Friederichs from the Museum für Naturkunde Berlin." httpUri-0="https://zenodo.org/record/2600896/files/figure.png" httpUri-1="https://zenodo.org/record/2600898/files/figure.png" pageId="32" pageNumber="33">Figs 19–20</figureCitation>
|
||
; Oudemans Mite Collection, Wikimedia Commons).
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5D015FFF30159404F044EE" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
The specimens from Iron Range match those of the New
|
||
<collectingCountry id="F308CBD68D5D015FFC62159404B04419" box="[917,998,1015,1040]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
specimens closely. The host beetle,
|
||
<taxonomicName id="4C1FF0C58D5D015FFF60127F06FE443D" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[151,424,1052,1076]" class="Insecta" family="Passallidae" genus="Protomocoelus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B96B57548D5D015FFF60127F06FE443D" box="[151,424,1052,1076]" italics="true" pageId="32" pageNumber="33">Protomocoelus australis</emphasis>
|
||
</taxonomicName>
|
||
, is found in the isolated pockets of rainforest in northern Cape York (such as Iron Range), as well as New
|
||
<collectingCountry id="F308CBD68D5D015FFEA7125C06F24451" box="[336,420,1087,1112]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F308CBD68D5D015FFE2B125C05CB4451" box="[476,669,1087,1112]" name="Solomon Islands" pageId="32" pageNumber="33">Solomon Islands</collectingCountry>
|
||
, New Britain, Seram, Waigeo and the Aru Islands (
|
||
<bibRefCitation id="EF8EF6B78D5D015FFB18125C02C64451" author="Gravely, F. H." box="[1263,1424,1087,1112]" pageId="32" pageNumber="33" pagination="177 - 353" refId="ref21907" refString="Gravely, F. H. (1914) An account of the Oriental Passalidae (Coleoptera), based primarily on the collection in the Indian Museum. Memoirs of the Indian Museum, 3, 177 - 353. https: // doi. org / 10.5962 / bhl. title. 9406" type="journal article" year="1914">Gravely 1914</bibRefCitation>
|
||
). This biogeographic pattern is matched by other genera of Indonesian and New Guinean
|
||
<taxonomicName id="4C1FF0C58D5D015FFB4B120702654474" authorityName="Leach" authorityYear="1815" box="[1212,1331,1124,1149]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subFamily" subFamily="Passalinae">Passalinae</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="4C1FF0C58D5D015FFF6012EB07A04496" box="[151,246,1160,1183]" class="Insecta" family="Passallidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFF6012EB07A04496" box="[151,246,1160,1183]" italics="true" pageId="32" pageNumber="33">Gonatas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFEDB12EB06C544A9" authorityName="Kaup" authorityYear="1871" box="[300,403,1160,1184]" class="Insecta" family="Passallidae" genus="Labienus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFEDB12EB06C544A9" box="[300,403,1160,1184]" italics="true" pageId="32" pageNumber="33">Labienus</emphasis>
|
||
</taxonomicName>
|
||
, which are also found only in Cape York in
|
||
<collectingCountry id="F308CBD68D5D015FFC8812E404BF44A9" box="[895,1001,1159,1184]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
|
||
. Southern regions are instead occupied mostly by species of
|
||
<taxonomicName id="4C1FF0C58D5D015FFE7B12CF054244CD" authorityName="Kaup" authorityYear="1868" box="[396,532,1196,1220]" class="Insecta" family="Passallidae" genus="Pharochilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFE7B12CF054244CD" box="[396,532,1196,1220]" italics="true" pageId="32" pageNumber="33">Pharochilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFDB912CF058144CD" authorityName="Kaup" authorityYear="1868" box="[590,727,1196,1220]" class="Insecta" family="Passallidae" genus="Mastachilus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFDB912CF058144CD" box="[590,727,1196,1220]" italics="true" pageId="32" pageNumber="33">Mastachilus</emphasis>
|
||
</taxonomicName>
|
||
, as well as the widespread genus
|
||
<taxonomicName id="4C1FF0C58D5D015FFB9712CF03A344CD" authorityName="Kaup" authorityYear="1868" box="[1120,1269,1196,1220]" class="Insecta" family="Passallidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFB9712CF03A344CD" box="[1120,1269,1196,1220]" italics="true" pageId="32" pageNumber="33">Aulacocyclus</emphasis>
|
||
</taxonomicName>
|
||
, which occurs throughout southeast Asia, northern
|
||
<collectingCountry id="F308CBD68D5D015FFDD812AC05C044E1" box="[559,662,1231,1256]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
|
||
and along its east coast.
|
||
</paragraph>
|
||
<paragraph id="8BA08B468D5D015FFF30129704144659" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
The other species of
|
||
<taxonomicName id="4C1FF0C58D5D015FFE4D1297051F4505" authorityName="Thorell" authorityYear="1882" box="[442,585,1268,1292]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96B57548D5D015FFE4D1297051F4505" box="[442,585,1268,1292]" italics="true" pageId="32" pageNumber="33">Megisthanus</emphasis>
|
||
</taxonomicName>
|
||
from New
|
||
<collectingCountry id="F308CBD68D5D015FFD25129704704504" box="[722,806,1268,1293]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
, excepting
|
||
<taxonomicName id="4C1FF0C58D5D015FFC5E129603684505" authorityName="Womersley" authorityYear="1937" box="[937,1086,1269,1292]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="papuanus">
|
||
<emphasis id="B96B57548D5D015FFC5E129604964505" box="[937,960,1269,1292]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC26129603684505" box="[977,1086,1269,1292]" italics="true" pageId="32" pageNumber="33">papuanus</emphasis>
|
||
</taxonomicName>
|
||
, are more difficult to separate from each other. The species described by Oudemans,
|
||
<taxonomicName id="4C1FF0C58D5D015FFD01137A04CF4539" authorityName="Oudemans" authorityYear="1905" box="[758,921,1304,1328]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moaifensus">
|
||
<emphasis id="B96B57548D5D015FFD01137A045B4539" box="[758,781,1305,1328]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFCEC137B04CF4539" box="[795,921,1304,1328]" italics="true" pageId="32" pageNumber="33">moaifensus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFC39137A03084539" authorityName="Oudemans" authorityYear="1905" box="[974,1118,1304,1328]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B96B57548D5D015FFC39137A04B34539" box="[974,997,1305,1328]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC04137B03084539" box="[1011,1118,1304,1328]" italics="true" pageId="32" pageNumber="33">orientalis</emphasis>
|
||
</taxonomicName>
|
||
, are both depicted with long setae, while the dorsal setae of
|
||
<taxonomicName id="4C1FF0C58D5D015FFE03135E05DF455D" authorityName="Thorell" authorityYear="1882" box="[500,649,1340,1364]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFE03135E055D455D" box="[500,523,1341,1364]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFDED135F05DF455D" box="[538,649,1340,1364]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFD60135E046C455D" authorityName="Thorell" authorityYear="1882" box="[663,826,1340,1364]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatamensis">
|
||
<emphasis id="B96B57548D5D015FFD60135E05F8455D" box="[663,686,1341,1364]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD4B135F046C455D" box="[700,826,1340,1364]" italics="true" pageId="32" pageNumber="33">hatamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFC86135E0343455D" authorityName="Karg" authorityYear="1997" box="[881,1045,1341,1364]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D5D015FFC86135E04DE455D" box="[881,904,1341,1364]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC61135E0343455D" box="[918,1045,1341,1364]" italics="true" pageId="32" pageNumber="33">postsetosus</emphasis>
|
||
</taxonomicName>
|
||
are considerably shorter (
|
||
<figureCitation id="132497C38D5D015FFACE135F07E14571" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStartId-0="30.[151,250,1982,2004]" captionStartId-1="31.[151,250,1957,1979]" captionTargetBox-0="[207,1374,193,1944]" captionTargetBox-1="[241,1339,193,1913]" captionTargetId-0="figure@30.[206,1374,193,1964]" captionTargetId-1="figure@31.[241,1342,193,1932]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 19. Photographs of the holotypes of Megisthanus doreianus Thorell, 1882, (A) dorsum; (B) venter; and Megisthanus hatamanensis Thorell, 1882, (C) dorsum; (D) venter. Photographs by Geoff Thompson, Queensland Museum." captionText-1="FIGURE 20. The holotype of Megisthanus postsetosus Karg, 1997 (ZMB 45476). (A) photograph; (B) detail of posterior idiosoma, drawn from photograph. Photograph by Anja Friederichs from the Museum für Naturkunde Berlin." httpUri-0="https://zenodo.org/record/2600896/files/figure.png" httpUri-1="https://zenodo.org/record/2600898/files/figure.png" pageId="32" pageNumber="33">Figs 19– 20</figureCitation>
|
||
). These species may also differ in the form of the posterior setae on the dorsal shield, which comprise a row of short stiff setae in
|
||
<taxonomicName id="4C1FF0C58D5D015FFE9213E6055F4595" authorityName="Oudemans" authorityYear="1905" box="[357,521,1412,1436]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moaifensus">
|
||
<emphasis id="B96B57548D5D015FFE9213E6062A4595" box="[357,380,1413,1436]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFE7C13E7055F4595" box="[395,521,1412,1436]" italics="true" pageId="32" pageNumber="33">moaifensus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFDC813E605864595" authorityName="Oudemans" authorityYear="1905" box="[575,720,1412,1436]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B96B57548D5D015FFDC813E605004595" box="[575,598,1413,1436]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD9313E705864595" box="[612,720,1412,1436]" italics="true" pageId="32" pageNumber="33">orientalis</emphasis>
|
||
</taxonomicName>
|
||
, as well as several very long setae, which are also present in
|
||
<taxonomicName id="4C1FF0C58D5D015FFA8A13E6064145C9" authorityName="Karg" authorityYear="1997" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D5D015FFA8A13E602C24595" box="[1405,1428,1413,1436]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFF6013CA064145C9" box="[151,279,1449,1472]" italics="true" pageId="32" pageNumber="33">postsetosus</emphasis>
|
||
</taxonomicName>
|
||
; these setae are either absent or broken in
|
||
<taxonomicName id="4C1FF0C58D5D015FFD1A13CA04D645C9" authorityName="Thorell" authorityYear="1882" box="[749,896,1448,1472]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFD1A13CA045245C9" box="[749,772,1449,1472]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFCE613CB04D645C9" box="[785,896,1448,1472]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFC4113CA030E45C9" authorityName="Thorell" authorityYear="1882" box="[950,1112,1448,1472]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatamensis">
|
||
<emphasis id="B96B57548D5D015FFC4113CA049B45C9" box="[950,973,1449,1472]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC2C13CB030E45C9" box="[987,1112,1448,1472]" italics="true" pageId="32" pageNumber="33">hatamensis</emphasis>
|
||
</taxonomicName>
|
||
. The shape of the anal shield may also help separate species, with
|
||
<taxonomicName id="4C1FF0C58D5D015FFDA513AE05BC45ED" authorityName="Thorell" authorityYear="1882" box="[594,746,1484,1508]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFDA513AE053F45ED" box="[594,617,1485,1508]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD8B13AF05BC45ED" box="[636,746,1484,1508]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1FF0C58D5D015FFD0913AE04F045ED" authorityName="Oudemans" authorityYear="1905" box="[766,934,1484,1508]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moaifensus">
|
||
<emphasis id="B96B57548D5D015FFD0913AE044345ED" box="[766,789,1485,1508]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFCDF13AF04F045ED" box="[808,934,1484,1508]" italics="true" pageId="32" pageNumber="33">moaifensus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFC1013AE03C745ED" authorityName="Karg" authorityYear="1997" box="[999,1169,1485,1508]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="postsetosus">
|
||
<emphasis id="B96B57548D5D015FFC1013AE04A845ED" box="[999,1022,1485,1508]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFBE613AE03C745ED" box="[1041,1169,1485,1508]" italics="true" pageId="32" pageNumber="33">postsetosus</emphasis>
|
||
</taxonomicName>
|
||
having a broad shield approximately twice as wide as long, and
|
||
<taxonomicName id="4C1FF0C58D5D015FFD831392044F4601" authorityName="Thorell" authorityYear="1882" box="[628,793,1520,1544]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatamensis">
|
||
<emphasis id="B96B57548D5D015FFD83139205DD4601" box="[628,651,1521,1544]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD6C1393044F4601" box="[667,793,1520,1544]" italics="true" pageId="32" pageNumber="33">hatamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1FF0C58D5D015FFCA5139204B24601" authorityName="Oudemans" authorityYear="1905" box="[850,996,1520,1544]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B96B57548D5D015FFCA51392043F4601" box="[850,873,1521,1544]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFC8F139304B24601" box="[888,996,1520,1544]" italics="true" pageId="32" pageNumber="33">orientalis</emphasis>
|
||
</taxonomicName>
|
||
having a squarer shield approximately 1.5 x as wide as long. The species of New
|
||
<collectingCountry id="F308CBD68D5D015FFD64107705B24624" box="[659,740,1556,1581]" name="Guinea-Bissau" pageId="32" pageNumber="33">Guinea</collectingCountry>
|
||
require revision based on new material so that intraspecific variation, host beetles and male morphology become known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305D8CD8D5D015FFF30103F021D4691" pageId="32" pageNumber="33" type="etymology">
|
||
<paragraph id="8BA08B468D5D015FFF30103F021D4691" blockId="32.[151,1437,151,1688]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96B57548D5D015FFF30103F0618467C" bold="true" box="[199,334,1628,1653]" pageId="32" pageNumber="33">Etymology.</emphasis>
|
||
This species is named in honour of Australian acarologist Dr Herbert Womersley, who treated the specimens from Misima Island under
|
||
<taxonomicName id="4C1FF0C58D5D015FFDC910E205854691" authorityName="Thorell" authorityYear="1882" box="[574,723,1664,1688]" class="Arachnida" family="Megisthanidae" genus="Megisthanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doreianus">
|
||
<emphasis id="B96B57548D5D015FFDC910E205034691" box="[574,597,1665,1688]" italics="true" pageId="32" pageNumber="33">M</emphasis>
|
||
.
|
||
<emphasis id="B96B57548D5D015FFD9310E305854691" box="[612,723,1664,1688]" italics="true" pageId="32" pageNumber="33">doreianus</emphasis>
|
||
</taxonomicName>
|
||
in his revision of the
|
||
<taxonomicName id="4C1FF0C58D5D015FFC30101C02104691" authority="(Womersley 1937)" authorityName="Womersley" authorityYear="1937" baseAuthorityName="Womersley" baseAuthorityYear="1937" box="[967,1350,1663,1688]" class="Arachnida" family="Megisthanidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">
|
||
Megisthanidae (
|
||
<bibRefCitation id="EF8EF6B78D5D015FFB8D101C02684691" author="Womersley, H." box="[1146,1342,1663,1688]" pageId="32" pageNumber="33" pagination="175 - 180" refId="ref22977" refString="Womersley, H. (1937) Australian Acarina of the genus Megisthanus Thorell. Transactions of the Royal Society of South Australia, 61, 175 - 180." type="journal article" year="1937">Womersley 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |