132 lines
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132 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.384.6197" ID-GBIF-Dataset="f8902ab2-4591-440e-93f1-fb2d2f850112" ID-PMC="PMC3950428" ID-Pensoft-Pub="1313-2970-384-1" ID-PubMed="24624022" ID-ZBK="832EF8274642416895252C2AD202EB9B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-384-1" ModsDocOrigin="ZooKeys 384" ModsDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" checkinTime="1451246226561" checkinUser="pensoft" docAuthor="Hopkins, Heidi" docDate="2014" docId="89CEF6C673602BA3AAA4CA2B60B771FB" docLanguage="en" docName="ZooKeys 384: 1-256" docOrigin="ZooKeys 384" docSource="http://dx.doi.org/10.3897/zookeys.384.6197" docTitle="Arenivaga estelleae Hopkins, 2014, sp. n." docType="treatment" docUuid="0AF54DC2-9A96-494E-B967-976A5AAFD8ED" docUuidSource="ZooBank" docVersion="4" lastPageNumber="51" masterDocId="FF86641EFFB8B30DFFC9F429FFD4FFA1" masterDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" masterLastPageNumber="256" masterPageNumber="1" pageNumber="49" updateTime="1668157791821" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hopkins, Heidi</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>384</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>256</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.384.6197</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.384.6197</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-384-1</mods:identifier>
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<mods:identifier type="ZBK">832EF8274642416895252C2AD202EB9B</mods:identifier>
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<mods:identifier type="ZooBank">832EF8274642416895252C2AD202EB9B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152051400" LSID="urn:lsid:zoobank.org:act:0AF54DC2-9A96-494E-B967-976A5AAFD8ED" httpUri="http://treatment.plazi.org/id/89CEF6C673602BA3AAA4CA2B60B771FB" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49">
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<subSubSection pageId="48" pageNumber="49" type="nomenclature">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName LSID="http://zoobank.org/0AF54DC2-9A96-494E-B967-976A5AAFD8ED" class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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<taxonomicNameLabel pageId="48" pageNumber="49">sp. n.</taxonomicNameLabel>
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Figures 51-53
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type locality">
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<paragraph pageId="48" pageNumber="49">Type locality.</paragraph>
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<paragraph pageId="48" pageNumber="49">USA, California, San Diego Co., north of Fallbrook.</paragraph>
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</subSubSection>
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<subSubSection pageId="49" pageNumber="50" type="material examined">
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<paragraph pageId="49" pageNumber="50">
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<pageBreakToken pageId="49" pageNumber="50" start="start">Material</pageBreakToken>
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examined.
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</paragraph>
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<paragraph pageId="49" pageNumber="50">
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Holotype: ♂ in LACM labeled "July 27, 1957, 2.3 mi N. Fallbrook, San Diego Co., Calif., Lionel A Stange "HOLOTYPE
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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Hopkins, 2012" [red label with black border].
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</paragraph>
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<paragraph pageId="49" pageNumber="50">
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Paratypes (41): USA: CA, San Diego Co., NAS Miramar 4, 11/13/1996, N Bloomfield, blacklite (1, SDMC); CA, San Diego Co., MCAS Miramar, 9/18/1998, N Bloomfield, blacklite (1, SDMC); CA, San Diego Co., Mission Gorge Dam, 7/26/1976, illegible (1, SDMC); CA, San Diego Co., San Diego, 6/10/1953, J Powell, at light (1, EMEC); CA, San Diego Co., Lake Hodges, 8/24/1976, DK Faulkner (1, SDMC); CA, San Diego Co., N San Vicente Res, 8/19/1976, DK Faulkner (1, SDMC); CA, San Diego Co., 2.3 mi N of Fallbrook, 7/27/1957, LA Stange (1, LACM); CA, San Diego Co., Boulevard Manzanita, 10/10/1980, R Messner (1, SDMC); CA, San Diego Co., Boulevard Manzanita, 10/5/1979, R Messner (1, SDMC); CA, San Diego Co., Boulevard Manzanita, 8/7/1979, R Messner (1, SDMC); CA, San Diego Co., Boulevard Manzanita, 7/12/1979, R Messner (4, SDMC); CA, San Diego Co., Boulevard Manzanita, 6/3/1980, R Messner (1, SDMC); CA, San Diego Co., Boulevard Manzanita, 5/28/1980, R Messner (1, SDMC); CA, San Diego Co., Rancho Santa Fe, 8/25/1958, JR Northern (1, LACM); CA, San Diego Co., San Diego, 6/19/1974, Munzenmaier & Patten, Japanese beetle trap, 74524-38,
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Arenivaga</taxonomicName>
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sp. Det. AR Hardy 1974 (1, CSCA); CA, San Diego Co., Boulevard Manzanita, 7/29/1979, R Messner (1, SDMC); CA, San Diego Co., Alpine, 7/18/1990, J Mitchell (1, SDMC); CA, San Diego Co.,, NAS Miramar 2, 5/19/1997, N Bloomfield, blacklite (1, USNM); CA, San Diego Co., San Diego, 7/17/1912, FE Maisdell (1, CAS); MEXICO: BC, 4 mi SW of La Zapopita, Valle de Trinidad, 4/16/1961, FS Truxal (4, LACM); BC, 11 mi E of Ojos Negros on road to Laguna Hanson, 8/9/1988, 1160 m, Weissman & Lightfoot, Stop #88-85 (1, CAS); BC, 8 mi E of Tecate, 7/6/1984, Brown & Tocco, green dot (1, CAS); BC, Ensenada, 8/30/1952 (2, USNM); BC, turnoff Hwy 1 to Motel Durado, 3 km S of Ensenada, 7/18/1977, D Weissman, coastal sand dunes (1, CAS); BC, 17 mi S of Ensenada, 6/14/1938, Michelbacher & Ross (1, CAS); BC, Hwy 2, 5.7 km W of El Condor at KM 88.7, 8/19/1995, 1210 m, Weissman & Lightfoot, Stop #95-65 (1, CAS); BC, 13 mi SW of La Zapopita, 6/14/1963, EL Sleeper, blacklite (2, CAS); BC, 17 mi S of Ensenada, 6/14/1938, Michelbacher & Ross, photo.spec. (1, LACM); BC, Km 56 on road to Sierra San Pedro Martir Park off Hwy 1, 8/4/1981, 900 m, Lightfoot & Weissman, #81-64 (1, CAS). All paratypes labeled "Paratype
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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Hopkins 2012" [blue label with black border].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="49" pageNumber="50" type="etymology">
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<paragraph pageId="49" pageNumber="50">Etymology.</paragraph>
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<paragraph pageId="49" pageNumber="50">The name is a noun in the genitive case. This species is named for my grandmother, Estelle Beaumont, who taught me so much including how to be firm yet kind; also for the Latin stella, meaning star, because it is a lovely little roach.</paragraph>
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</subSubSection>
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<subSubSection pageId="49" pageNumber="50" type="distribution">
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<paragraph pageId="49" pageNumber="50">Distribution.</paragraph>
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<paragraph pageId="49" pageNumber="50">This species is found in far southwestern California and northwestern Baja California Norte, Mexico. See Fig. 53.</paragraph>
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</subSubSection>
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<subSubSection pageId="49" pageNumber="50" type="diagnosis">
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<paragraph pageId="49" pageNumber="50">Diagnosis.</paragraph>
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<paragraph pageId="49" pageNumber="50">
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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is smaller and browner than average for
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Arenivaga</taxonomicName>
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and could be confused with the parapatric species
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga gaiophanes" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="gaiophanes">Arenivaga gaiophanes</taxonomicName>
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(in its external appearance) and
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga paradoxa" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="paradoxa">Arenivaga paradoxa</taxonomicName>
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(in its genitalia). These three species are most likely closely related.
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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may be distinguished by the broad central field on the right dorsal phallomere and very wide gap on the right ventral phallomere. See Figs 52 and 124.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="description">
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<paragraph pageId="50" pageNumber="51">
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<pageBreakToken pageId="50" pageNumber="51" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="50" pageNumber="51">Male.Measurements. Holotype TL = 16.5 mm, GW = 8.9 mm, PW = 4.93 mm, PL = 3.31 mm, TL/GW = 1.85, PL/PW = 0.67. EW = 0.45 mm; OW = 0.45 mm. Among paratypes range of TL 15.1-20.6 mm; range of GW 7.2-9.8 mm; range of PW 4.57-5.91 mm; range of PL 3.31-4.25 mm.</paragraph>
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<paragraph pageId="50" pageNumber="51">
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Head. Two ocelli large, ovoid and protruding (0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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0.2 mm); vertex flat and medium brown with small ridges in rays around upper apex of eyes and extending onto ocellar tubercles; interocellar space concave with fine horizontal corrugations; medium brown laterally shading to light brown in concavity; deep vertical indentations medial to ocelli. Frons light brown, concave; anterior portion of frons bulbous, light brown medially shading darker laterally; pale beige anteclypeus with one horizontal corrugation. See Fig. 51d.
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</paragraph>
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<paragraph pageId="50" pageNumber="51">Pronotum. Pronotum translucent, waxy beige; dorsal surface of pronotum with short fine brown setae centrally and posteriorly grading to longer, thicker setae laterally and anteriorly; pronotal pattern brown "panther face"; brown maculations scattered across posterior 80% of dorsal surface of pronotum; no aura. See Fig. 51c.</paragraph>
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<paragraph pageId="50" pageNumber="51">Body. Wing brace present. Two tarsal claws present. Legs beige, body light brown with dark brown markings on lateral anterior portion of each sternite; subgenital plate light brown; strongly asymmetrical with rounded apices. See Fig. 51b.</paragraph>
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<paragraph pageId="50" pageNumber="51">Forewings. Wings extended well beyond abdominal apex (~40% of wing length); color light brown to dark medium brown depending on specimen, faintly blotchy; surface matte and opaque. See Fig. 51a.</paragraph>
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<paragraph pageId="50" pageNumber="51">Genitalia. Right dorsal phallomere composed of bulbous lightly sclerotized hook-shaped lobe, articulated with right ventral phallomere on lateral side; central field sclerotized; medial margin with slight concavity, shagreened with toothed margin; anterior and posterior margins deeply emarginate, posterior margin curving back out to several teeth; interior of lateral articulation setose; broad rolled sclerotized lip projecting posteriorly from lateral articulation to several teeth. Small central sclerite broad, flat and finely punctate with posteriorly projecting, sclerotized lip at anterior end; right ventral phallomere extends from articulation to form rounded shagreened structure with acute edge; attached anteriorly is mildly dorsally projecting flanged arm, shagreened with roughly toothed edge. Folded anterior portion of left phallomere dramatically modified into heavily setose, medially projecting, scoop shape. Genital hook with short extension to pointed head with average hook; arm robust. See Fig. 52.</paragraph>
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<caption pageId="50" pageNumber="51">
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<paragraph pageId="50" pageNumber="51">
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Figure 51.
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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, a dorsal habitus b ventral habitus c pronotum d head.
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</paragraph>
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</caption>
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<caption pageId="50" pageNumber="51">
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<paragraph pageId="50" pageNumber="51">
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Figure 52.
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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, genitalia: a right dorsal phallomere b right ventral phallomere c small central sclerite d genital hook e left phallomere. Arrow(s) indicate diagnostic characters (see text).
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</paragraph>
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</caption>
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<caption pageId="50" pageNumber="51">
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<paragraph pageId="50" pageNumber="51">
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Figure 53.
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<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga estelleae" order="Blattodea" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="estelleae">Arenivaga estelleae</taxonomicName>
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, distribution.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="habitat and natural history">
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<paragraph pageId="50" pageNumber="51">Habitat and natural history.</paragraph>
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<paragraph pageId="50" pageNumber="51">All life history elements remain unobserved.</paragraph>
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</subSubSection>
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</treatment>
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</document> |