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<document id="A54A1D31C96F0169093E0F2C5940EACD" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFBAFFA3FF06AA732A6DFD68" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas iranicus Anker, Naderloo &amp; Marin 2010" docType="treatment" docVersion="2" lastPageNumber="191" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="189" updateTime="1702284319693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7C46B6DD346DA36A5E6902379AA219D5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="35EC9DD1487930F093DE9CFAE0E95981">
<mods:title id="55C5D87CD7C4EF194D480C52723059CA">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="06567CB4ED32F54018682F5A5781FE64" type="personal">
<mods:role id="6C241DA8A9578BBD6CB573144537EF93">
<mods:roleTerm id="C726279855BDFEF832A57BB898AA12FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="972AE9E5A51D43639C289AD3CC1738A5">Anker, Arthur</mods:namePart>
<mods:affiliation id="53C6D73E6D970033F8390CA53BDED590">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="ECD8F2F6082DB802D3761A659A799ACB">text</mods:typeOfResource>
<mods:relatedItem id="34C279CDE701CF9A283D8A50FACF14D0" type="host">
<mods:titleInfo id="89D493B8E217249E98AE39B8EAC34480">
<mods:title id="9094436D309BBED4AD995873B227502C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94DDD7BD02F2E83BDD35805B6C6E0067">
<mods:date id="33FBDA153815A13FAD22CEBE5DBBB9A6">2023</mods:date>
<mods:detail id="4645D490B6FDF7A59B2438FD964308CD" type="pubDate">
<mods:number id="9AD6518E8B124009987D90A7B10DA646">2023-12-11</mods:number>
</mods:detail>
<mods:detail id="FC6B25B200F6930FCBF10EDB3D7B0840" type="volume">
<mods:number id="22806CACA7213893FEC6C9B82D453A1D">5383</mods:number>
</mods:detail>
<mods:detail id="A63AA04D4408DC8391542A2772BDB445" type="issue">
<mods:number id="CEB1D8B289E7822C13F8534ED5825CDC">2</mods:number>
</mods:detail>
<mods:extent id="0CAD93928E5E2C25A9767064E60AAF11" unit="page">
<mods:start id="49E657F4D2651821282008CEE2964401">179</mods:start>
<mods:end id="C4592ADAF5643F50875DCDAD52879D65">215</mods:end>
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<mods:url id="CFF3C66CB49E9F7EA8A9ABFCB32B9D75">https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448</mods:url>
</mods:location>
<mods:classification id="C41C74FBBEFF4CEC7B4215999510B67F">journal article</mods:classification>
<mods:identifier id="3CE65FFE0032DE0B0183FD5E41528BE5" type="DOI">10.11646/zootaxa.5383.2.5</mods:identifier>
<mods:identifier id="92B07926F768716D67FCD1F45588FF74" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="42E14FEBBC0481CC97E07E197E36C3DF" type="Zenodo-Dep">10350767</mods:identifier>
<mods:identifier id="80D70D0488AB90093BE541E7A42A0368" type="ZooBank">7E434B40-6346-4E6D-BC55-38EBAE24BD52</mods:identifier>
</mods:mods>
<treatment id="386ABE42FFBAFFA3FF06AA732A6DFD68" ID-DOI="http://doi.org/10.5281/zenodo.10350895" ID-Zenodo-Dep="10350895" LSID="urn:lsid:plazi:treatment:386ABE42FFBAFFA3FF06AA732A6DFD68" httpUri="http://treatment.plazi.org/id/386ABE42FFBAFFA3FF06AA732A6DFD68" lastPageId="12" lastPageNumber="191" pageId="10" pageNumber="189">
<subSubSection id="F8D95CDFFFBAFFA5FF06AA732CDDFBFD" box="[151,765,1086,1113]" pageId="10" pageNumber="189" type="nomenclature">
<paragraph id="B07C0F54FFBAFFA5FF06AA732CDDFBFD" blockId="10.[151,765,1086,1149]" box="[151,765,1086,1113]" pageId="10" pageNumber="189">
<heading id="EB34B838FFBAFFA5FF06AA732CDDFBFD" bold="true" box="[151,765,1086,1113]" fontSize="11" level="1" pageId="10" pageNumber="189" reason="1">
<taxonomicName id="77C374D7FFBAFFA5FF06AA732CDDFBFD" ID-CoL="85MC5" authority="Anker, Naderloo &amp; Marin, 2010" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[151,765,1086,1113]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBAFFA5FF06AA732CDDFBFD" bold="true" box="[151,765,1086,1113]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF06AA732F47FBFC" bold="true" box="[151,359,1086,1113]" italics="true" pageId="10" pageNumber="189">Athanas iranicus</emphasis>
<bibRefCitation id="D45272A5FFBAFFA5FEFCAA732CDDFBFD" author="Anker, A. &amp; Naderloo, R. &amp; Marin, I." box="[365,765,1086,1113]" pageId="10" pageNumber="189" pagination="53 - 60" refId="ref20396" refString="Anker, A., Naderloo, R. &amp; Marin, I. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2372.1.6" type="journal article" year="2010">Anker, Naderloo &amp; Marin, 2010</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF06AA2E2ECBFBD8" box="[151,235,1122,1149]" pageId="10" pageNumber="189" type="description">
<paragraph id="B07C0F54FFBAFFA5FF06AA2E2ECBFBD8" blockId="10.[151,765,1086,1149]" box="[151,235,1122,1149]" pageId="10" pageNumber="189">
(
<figureCitation id="28F813D1FFBAFFA5FF0EAA2E2EC2FBD8" box="[159,226,1122,1149]" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="10" pageNumber="189">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF06AAEA2B0CFB1A" box="[151,1324,1191,1215]" pageId="10" pageNumber="189" type="reference_group">
<paragraph id="B07C0F54FFBAFFA5FF06AAEA2B0CFB1A" blockId="10.[151,1324,1191,1215]" box="[151,1324,1191,1215]" pageId="10" pageNumber="189">
<treatmentCitationGroup id="90D3287AFFBAFFA5FF06AAEA2B0CFB1A" box="[151,1324,1191,1215]" pageId="10" pageNumber="189">
<taxonomicName id="77C374D7FFBAFFA5FF06AAEA2C03FB1A" ID-CoL="85MC5" authority="Anker et al. 2010: 54" authorityName="Anker" authorityPageNumber="54" authorityYear="2010" box="[151,547,1191,1215]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBAFFA5FF06AAEA2F64FB1A" box="[151,324,1191,1215]" italics="true" pageId="10" pageNumber="189">Athanas iranicus</emphasis>
Anker
<emphasis id="82B7D346FFBAFFA5FE1FAAE52FE1FB1A" box="[398,449,1191,1215]" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
2010: 54
</taxonomicName>
, figs. 1–4; Naderloo &amp; Türkay 2012: 12;
<treatmentCitation id="31622945FFBAFFA5FC57AAEA2AE3FB1A" author="Ashrafi, H. &amp; Dehghani, A. &amp; Sari, A. &amp; Naderloo, R." box="[966,1219,1191,1215]" page="520" pageId="10" pageNumber="189" year="2020">
<bibRefCitation id="D45272A5FFBAFFA5FC57AAEA2AE3FB1A" author="Ashrafi, H. &amp; Dehghani, A. &amp; Sari, A. &amp; Naderloo, R." box="[966,1219,1191,1215]" pageId="10" pageNumber="189" pagination="521 - 534" refId="ref20514" refString="Ashrafi, H., Dehghani, A., Sari, A. &amp; Naderloo, R. (2020 a) An updated checklist of caridean shrimps of the Persian Gulf and Gulf of Oman. Zootaxa, 4747 (3), 521 - 534. https: // doi. org / 10.11646 / zootaxa. 4747.3.6" type="journal article" year="2020">
Ashrafi
<emphasis id="82B7D346FFBAFFA5FB88AAE52A65FB1A" box="[1049,1093,1191,1215]" italics="true" pageId="10" pageNumber="189">et al</emphasis>
. 2020a: 520
</bibRefCitation>
</treatmentCitation>
(
<tableCitation id="FD413AEFFFBAFFA5FB43AAEA2B3CFB1A" box="[1234,1308,1191,1215]" captionStart="TABLE 1" captionStartId="2.[152,244,1052,1076]" captionText="TABLE 1. Gill-exopod formula of Athanas Leach, 1814." pageId="10" pageNumber="189">Table 1</tableCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF06AAA02D7CFA45" pageId="10" pageNumber="189" type="materials_examined">
<paragraph id="B07C0F54FFBAFFA5FF06AAA02D7CFA45" blockId="10.[151,1437,1261,2008]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF06AAA02F56FAA2" bold="true" box="[151,374,1261,1287]" pageId="10" pageNumber="189">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFBAFFA5FE10AAA02C36FAEA" collectingDate="2022-01-02" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="23.508972" location="Bandar Khayran" longLatPrecision="1" longitude="58.732777" pageId="10" pageNumber="189" specimenCode="UF68857" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFBAFFA5FE10AAA02FFFFAA2" box="[385,479,1261,1287]" count="1" pageId="10" pageNumber="189" type="female">1 female</specimenCount>
(cl
<quantity id="773BA2B1FFBAFFA5FD95AAA02C7AFAA2" box="[516,602,1261,1287]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="189" unit="mm" value="3.6">3.6mm</quantity>
),
<collectionCode id="D6D29791FFBAFFA5FDFBAAA32CECFAAD" box="[618,716,1262,1288]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="10" pageNumber="189" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBAFFA5FD41AAA32D60FAAD" box="[720,832,1261,1288]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="10" pageNumber="189" type="Museum">UF68857</specimenCode>
,
<collectingCountry id="C8D44FC4FFBAFFA5FCD9AAA02DAFFAA2" box="[840,911,1261,1287]" name="Oman" pageId="10" pageNumber="189">Oman</collectingCountry>
, east of Muscat,
<location id="B51C598FFFBAFFA5FBAFAAA02AD6FAA2" LSID="urn:lsid:plazi:treatment:386ABE42FFBAFFA3FF06AA732A6DFD68:B51C598FFFBAFFA5FBAFAAA02AD6FAA2" box="[1086,1270,1261,1287]" country="Oman" latitude="23.508972" longLatPrecision="1" longitude="58.732777" name="Bandar Khayran" pageId="10" pageNumber="189">Bandar Khayran</location>
,
<geoCoordinate id="D5F76993FFBAFFA5FB6FAAA02BBCFAA2" box="[1278,1436,1261,1287]" degrees="23" direction="north" minutes="30" orientation="latitude" pageId="10" pageNumber="189" precision="1" seconds="32.3" value="23.508972">23°30’32.3”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFBAFFA5FF38AB5C2F67FA89" box="[169,327,1297,1324]" degrees="58" direction="east" minutes="43" orientation="longitude" pageId="10" pageNumber="189" precision="1" seconds="58.0" value="58.732777">58°43’58.0”E</geoCoordinate>
, mangrove, muddy sand and mud, depth
<quantity id="773BA2B1FFBAFFA5FCBAAB5C2D58FA89" box="[811,888,1297,1324]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="189" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="3BEEE867FFBAFFA5FCBAAB5C2D58FA89" box="[811,888,1297,1324]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="189" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</elevation>
</quantity>
, suction pump, in burrow, leg.
<collectorName id="1D366A82FFBAFFA5FB74AB5F2B6FFA8E" box="[1253,1359,1297,1324]" pageId="10" pageNumber="189">A. Anker</collectorName>
<emphasis id="82B7D346FFBAFFA5FACBAB5F2BAEFA8E" box="[1370,1422,1297,1323]" italics="true" pageId="10" pageNumber="189">et al</emphasis>
.,
<date id="C47D2994FFBAFFA5FF06AB782F36FAEA" box="[151,278,1333,1359]" pageId="10" pageNumber="189" value="2022-01-02">
<collectingDate id="D439D07CFFBAFFA5FF06AB782F36FAEA" box="[151,278,1333,1359]" pageId="10" pageNumber="189" value="2022-01-02">01.02.2022</collectingDate>
</date>
[fcn BOMAN-09986]
</materialsCitation>
;
<materialsCitation id="00AB0509FFBAFFA5FDB3AB782D5AFA1E" collectingDate="2022-04-02" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="23.509027" location="Bandar Khayran" longLatPrecision="1" longitude="58.73153" pageId="10" pageNumber="189" specimenCode="UF 68860, UF 68858" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFBAFFA5FDB3AB782CA3FAEA" box="[546,643,1333,1359]" count="1" pageId="10" pageNumber="189" type="female">1 female</specimenCount>
(cl
<quantity id="773BA2B1FFBAFFA5FD21AB782D2BFAEA" box="[688,779,1333,1359]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="189" unit="mm" value="2.8">2.8 mm</quantity>
),
<collectionCode id="D6D29791FFBAFFA5FC8FAB7B2DA0FAF5" box="[798,896,1334,1360]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="10" pageNumber="189" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBAFFA5FC19AB7B2DDCFAEA" box="[904,1020,1333,1360]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="10" pageNumber="189" type="Museum">UF 68860</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-09985]; 1 ovig. female (cl
<quantity id="773BA2B1FFBAFFA5FD75AB142D60FAD6" box="[740,832,1369,1395]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="189" unit="mm" value="3.6">3.6 mm</quantity>
),
<collectionCode id="D6D29791FFBAFFA5FCC5AB172D96FAD1" box="[852,950,1370,1396]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="10" pageNumber="189" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBAFFA5FC2EAB172A14FAD1" box="[959,1076,1369,1396]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="10" pageNumber="189" type="Museum">UF 68858</specimenCode>
,
<collectingCountry id="C8D44FC4FFBAFFA5FBD0AB142AA8FAD6" box="[1089,1160,1369,1395]" name="Oman" pageId="10" pageNumber="189">Oman</collectingCountry>
, east of Muscat,
<location id="B51C598FFFBAFFA5FADAAB142EDAFA32" LSID="urn:lsid:plazi:treatment:386ABE42FFBAFFA3FF06AA732A6DFD68:B51C598FFFBAFFA5FADAAB142EDAFA32" country="Oman" latitude="23.509027" longLatPrecision="1" longitude="58.73153" name="Bandar Khayran" pageId="10" pageNumber="189">Bandar Khayran</location>
,
<geoCoordinate id="D5F76993FFBAFFA5FE97AB302F83FA3D" box="[262,419,1405,1432]" degrees="23" direction="north" minutes="30" orientation="latitude" pageId="10" pageNumber="189" precision="1" seconds="32.5" value="23.509027">23°30’32.5”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFBAFFA5FE28AB302C77FA3D" box="[441,599,1405,1432]" degrees="58" direction="east" minutes="43" orientation="longitude" pageId="10" pageNumber="189" precision="1" seconds="53.5" value="58.73153">58°43’53.5”E</geoCoordinate>
, mangrove and adjacent mudflat, at night, depth at low tide
<quantity id="773BA2B1FFBAFFA5FB66AB302B60FA3D" box="[1271,1344,1405,1432]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="189" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="3BEEE867FFBAFFA5FB66AB302B60FA3D" box="[1271,1344,1405,1432]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="189" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</elevation>
</quantity>
, suction pump, in burrow, leg.
<collectorName id="1D366A82FFBAFFA5FE1DABEF2FD4FA1E" box="[396,500,1441,1468]" pageId="10" pageNumber="189">A. Anker</collectorName>
,
<date id="C47D2994FFBAFFA5FE6FABEC2C5DFA1E" box="[510,637,1441,1467]" pageId="10" pageNumber="189" value="2022-04-02">
<collectingDate id="D439D07CFFBAFFA5FE6FABEC2C5DFA1E" box="[510,637,1441,1467]" pageId="10" pageNumber="189" value="2022-04-02">04.02.2022</collectingDate>
</date>
[fcn BOMAN-11371]
</materialsCitation>
;
<materialsCitation id="00AB0509FFBAFFA5FC14ABEC2D78FA45" collectingDate="2022-04-02" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="23.509027" location="Bandar Khayran" longLatPrecision="1" longitude="58.73153" pageId="10" pageNumber="189" specimenCode="UF 68861" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFBAFFA5FC14ABEC2DC5FA1E" box="[901,997,1441,1467]" count="1" pageId="10" pageNumber="189" type="female">1 female</specimenCount>
(cl
<quantity id="773BA2B1FFBAFFA5FB81ABEC2A4AFA1E" box="[1040,1130,1441,1468]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="189" unit="mm" value="2.5">2.5 mm</quantity>
),
<collectionCode id="D6D29791FFBAFFA5FBEDABEF2AFEFA19" box="[1148,1246,1442,1468]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="10" pageNumber="189" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBAFFA5FB74ABEF2B78FA1E" box="[1253,1368,1441,1468]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="10" pageNumber="189" type="Museum">UF 68861</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-11355]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA4FF56ABA42CA0FF70" lastPageId="11" lastPageNumber="190" pageId="10" pageNumber="189" type="description">
<paragraph id="B07C0F54FFBAFFA5FF56ABA42E95F9E9" blockId="10.[151,1437,1261,2008]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF56ABA42F71F9A6" bold="true" box="[199,337,1513,1539]" pageId="10" pageNumber="189">Description</emphasis>
. See Anker
<emphasis id="82B7D346FFBAFFA5FE4CABA72C36F9A6" box="[477,534,1513,1539]" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
(2010) for description and illustrations; some taxonomic remarks on the Omani material are provided below, whereas the species’ colour pattern is described and illustrated for the first time (
<figureCitation id="28F813D1FFBAFFA5FAFEA8402E87F9E9" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="10" pageNumber="189">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="B07C0F54FFBAFFA4FF56A8182CA0FF70" blockId="10.[151,1437,1261,2008]" lastBlockId="11.[151,1436,151,429]" lastPageId="11" lastPageNumber="190" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF56A8182F46F9CA" bold="true" box="[199,358,1621,1647]" pageId="10" pageNumber="189">Colour in life</emphasis>
. Background semi-opaque whitish; carapace and pleon speckled with numerous blotches of red chromatophores, except for large part of branchial area of carapace; dorsal surface of carapace, including rostral and post-rostral area, with large patches of buff white chromatophores; pleon with broad mid-dorsal band of buff white colour stretching from anterior margin of first pleonite to posterior margin of sixth pleonite, and extending as transverse bands on first, third, fourth (slightly) and sixth (forming large transverse patch) pleonites; some red chromatophores visible within buff white mid-dorsal band or patches on carapace and pleon; telson with proximal two-thirds bright red and distal one-third conspicuously yellowish white; antennular peduncle with bright red chromatophores on stylocerite and some parts of first article, and bright yellow white chromatophores on most of remaining peduncle and proximal flagella; antennae largely translucent with bright red chromatophores on scaphocerite and carpocerite, distal area of scaphocerite yellow white, flagellum colourless; major cheliped hyaline white with patches of red chromatophores, especially on chela; minor cheliped and remaining pereiopods, as well as pleopods, mostly whitish; uropods with proximal half deep red and distal half conspicuously yellowish white (similar to telson); eggs red orange (
<figureCitation id="28F813D1FFBBFFA4FDBFAEF62C52FF70" box="[558,626,186,213]" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="11" pageNumber="190">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBBFFA4FF56AE932BB6FF5C" box="[199,1430,222,249]" pageId="11" pageNumber="190" type="distribution">
<paragraph id="B07C0F54FFBBFFA4FF56AE932BB6FF5C" blockId="11.[151,1436,151,429]" box="[199,1430,222,249]" pageId="11" pageNumber="190">
<emphasis id="82B7D346FFBBFFA4FF56AE932F79FF5D" bold="true" box="[199,345,222,248]" pageId="11" pageNumber="190">Distribution</emphasis>
.
<collectingCountry id="C8D44FC4FFBBFFA4FEF6AE922FB8FF5C" box="[359,408,223,249]" name="Iran" pageId="11" pageNumber="190">Iran</collectingCountry>
:
<collectingCountry id="C8D44FC4FFBBFFA4FE32AE922C1EFF5C" box="[419,574,223,249]" name="Qeshm Island" pageId="11" pageNumber="190">Qeshm Island</collectingCountry>
(Anker
<emphasis id="82B7D346FFBBFFA4FD0AAE922CEDFF5C" box="[667,717,223,249]" italics="true" pageId="11" pageNumber="190">et al</emphasis>
. 2010);
<collectingCountry id="C8D44FC4FFBBFFA4FCBBAE922D53FF5C" box="[810,883,223,249]" name="Oman" pageId="11" pageNumber="190">Oman</collectingCountry>
: Bandar Khayran east of
<collectingCountry id="C8D44FC4FFBBFFA4FB1FAE922AC0FF5C" box="[1166,1248,223,249]" name="Oman" pageId="11" pageNumber="190">Muscat</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBBFFA4FF56AF4E2DBDFE08" pageId="11" pageNumber="190" type="biology_ecology">
<paragraph id="B07C0F54FFBBFFA4FF56AF4E2DBDFE08" blockId="11.[151,1436,151,429]" pageId="11" pageNumber="190">
<emphasis id="82B7D346FFBBFFA4FF56AF4E2F06FEB8" bold="true" box="[199,294,259,285]" pageId="11" pageNumber="190">Ecology</emphasis>
. The four Omani specimens of
<taxonomicName id="77C374D7FFBBFFA4FD19AF4E2D23FEB9" box="[648,771,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBBFFA4FD19AF4E2D23FEB9" box="[648,771,259,285]" italics="true" pageId="11" pageNumber="190">A. iranicus</emphasis>
</taxonomicName>
were collected in muddy mangrove channels from burrows of unknown hosts. The syntopic macrofauna included large burrowing snapping shrimps (
<taxonomicName id="77C374D7FFBBFFA4FB07AF6A2B08FEE4" box="[1174,1320,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="82B7D346FFBBFFA4FB07AF6A2AD0FEE4" box="[1174,1264,295,321]" italics="true" pageId="11" pageNumber="190">Alpheus</emphasis>
spp.
</taxonomicName>
) and their associates (e.g.,
<taxonomicName id="77C374D7FFBBFFA4FEC4AF062CCEFEC0" authority="Barnard, 1962" authorityName="Barnard" authorityYear="1962" box="[341,750,331,357]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="82B7D346FFBBFFA4FEC4AF062C1CFEC1" box="[341,572,331,357]" italics="true" pageId="11" pageNumber="190">Salmoneus rostratus</emphasis>
<bibRefCitation id="D45272A5FFBBFFA4FDD6AF062CCEFEC0" author="Barnard, K. H." box="[583,750,331,357]" pageId="11" pageNumber="190" pagination="239 - 245" refId="ref21129" refString="Barnard, K. H. (1962) New records of marine Crustacea from the East African region. Crustaceana, 3, 239 - 245. https: // doi. org / 10.1163 / 156854062 X 00463" type="journal article" year="1962">Barnard, 1962</bibRefCitation>
</taxonomicName>
), several species of fossorial crabs (
<taxonomicName id="77C374D7FFBBFFA4FB04AF062BAFFEC0" box="[1173,1423,331,357]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="82B7D346FFBBFFA4FB04AF062B73FEC0" box="[1173,1363,331,357]" italics="true" pageId="11" pageNumber="190">Macrophthalmus</emphasis>
spp.
</taxonomicName>
), and callianassid ghost-shrimps. The
<typeStatus id="6F78B1F6FFBBFFA4FDABAF222CBCFE2C" box="[570,668,367,393]" pageId="11" pageNumber="190" type="holotype">holotype</typeStatus>
from
<collectingCountry id="C8D44FC4FFBBFFA4FD77AF222D33FE2C" box="[742,787,367,393]" name="Iran" pageId="11" pageNumber="190">Iran</collectingCountry>
was collected, according to the label accompanying the specimen, in “muddy sand with shell fragments” (Anker
<emphasis id="82B7D346FFBBFFA4FC87AFDE2D6EFE08" box="[790,846,403,429]" italics="true" pageId="11" pageNumber="190">et al.</emphasis>
2010).
</paragraph>
</subSubSection>
<caption id="E4BC5FDCFFBBFFA4FF06A8E92EFEF8A1" ID-DOI="http://doi.org/10.5281/zenodo.10350783" ID-Zenodo-Dep="10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="11" pageNumber="190" startId="11.[151,250,1700,1724]" targetBox="[151,1436,453,1677]" targetPageId="11" targetType="figure">
<paragraph id="B07C0F54FFBBFFA4FF06A8E92EFEF8A1" blockId="11.[151,1436,1700,1796]" pageId="11" pageNumber="190">
<emphasis id="82B7D346FFBBFFA4FF06A8E92F35F919" bold="true" box="[151,277,1700,1724]" pageId="11" pageNumber="190">FIGURE 5.</emphasis>
<taxonomicName id="77C374D7FFBBFFA4FE8CA8E92D05F919" authority="Anker, Naderloo &amp; Marin, 2010" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[285,805,1700,1724]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBBFFA4FE8CA8E92FECF919" box="[285,460,1700,1724]" italics="true" pageId="11" pageNumber="190">Athanas iranicus</emphasis>
<bibRefCitation id="D45272A5FFBBFFA4FE45A8E92D05F919" author="Anker, A. &amp; Naderloo, R. &amp; Marin, I." box="[468,805,1700,1724]" pageId="11" pageNumber="190" pagination="53 - 60" refId="ref20396" refString="Anker, A., Naderloo, R. &amp; Marin, I. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2372.1.6" type="journal article" year="2010">Anker, Naderloo &amp; Marin, 2010</bibRefCitation>
</taxonomicName>
, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author.
</paragraph>
</caption>
<subSubSection id="F8D95CDFFFBBFFA3FF56A91B2A6DFD68" lastPageId="12" lastPageNumber="191" pageId="11" pageNumber="190" type="discussion">
<paragraph id="B07C0F54FFBBFFA3FF56A91B2CF6FEC0" blockId="11.[151,1437,1878,2013]" lastBlockId="12.[151,1437,151,719]" lastPageId="12" lastPageNumber="191" pageId="11" pageNumber="190">
<emphasis id="82B7D346FFBBFFA4FF56A91B2F14F8D5" bold="true" box="[199,308,1878,1904]" pageId="11" pageNumber="190">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFBBFFA4FED5A91B2C25F8D5" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[324,517,1878,1904]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBBFFA4FED5A91B2C25F8D5" box="[324,517,1878,1904]" italics="true" pageId="11" pageNumber="190">Athanas iranicus</emphasis>
</taxonomicName>
is a distinctive member of the genus
<taxonomicName id="77C374D7FFBBFFA4FC2BA91B2A36F8D5" authorityName="Leach" authorityYear="1814" box="[954,1046,1878,1904]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFBBFFA4FC2BA91B2A36F8D5" box="[954,1046,1878,1904]" italics="true" pageId="11" pageNumber="190">Athanas</emphasis>
</taxonomicName>
, due to its unique combination of morphological features. Especially diagnostic for
<taxonomicName id="77C374D7FFBBFFA4FD54A9362D60F831" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[709,832,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="190" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBBFFA4FD54A9362D60F831" box="[709,832,1915,1941]" italics="true" pageId="11" pageNumber="190">A. iranicus</emphasis>
</taxonomicName>
are the very short rostrum; the greatly reduced orbital teeth; the robust, somewhat unequal and assymmetrical chelipeds, which are furnished with dense fields of fine setae, one extending from the carpus to the proximal portion of the propodus and one on the fingers; and the relatively robust walking legs (Anker
<emphasis id="82B7D346FFBCFFA3FDD6AEDA2CA1FF14" box="[583,641,151,177]" italics="true" pageId="12" pageNumber="191">et al.</emphasis>
2010). The species was hitherto known only from the type locality in
<collectingCountry id="C8D44FC4FFBCFFA3FF06AEF62EE4FF70" box="[151,196,187,213]" name="Iran" pageId="12" pageNumber="191">Iran</collectingCountry>
(
<collectingCountry id="C8D44FC4FFBCFFA3FF44AEF62F54FF70" box="[213,372,187,213]" name="Qeshm Island" pageId="12" pageNumber="191">Qeshm Island</collectingCountry>
north of Strait of
<collectingCountry id="C8D44FC4FFBCFFA3FDDDAEF62C8AFF70" box="[588,682,187,213]" name="Iran" pageId="12" pageNumber="191">Hormuz</collectingCountry>
) and based on a single male
<typeStatus id="6F78B1F6FFBCFFA3FC6FAEF62A44FF70" box="[1022,1124,187,213]" pageId="12" pageNumber="191" type="holotype">holotype</typeStatus>
. The material from Bandar Khayran represents the second only finding of
<taxonomicName id="77C374D7FFBCFFA3FD38AE922D05FF5D" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[681,805,223,249]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBCFFA3FD38AE922D05FF5D" box="[681,805,223,249]" italics="true" pageId="12" pageNumber="191">A. iranicus</emphasis>
</taxonomicName>
and extends the previously known distribution range of the species from
<collectingCountry id="C8D44FC4FFBCFFA3FECBAF4E2FA7FEB8" box="[346,391,259,285]" name="Iran" pageId="12" pageNumber="191">Iran</collectingCountry>
to
<collectingCountry id="C8D44FC4FFBCFFA3FE3FAF4E2FD6FEB8" box="[430,502,259,285]" name="Oman" pageId="12" pageNumber="191">Oman</collectingCountry>
. It is likely that
<taxonomicName id="77C374D7FFBCFFA3FD20AF4E2D0EFEB9" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[689,814,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBCFFA3FD20AF4E2D0EFEB9" box="[689,814,259,285]" italics="true" pageId="12" pageNumber="191">A. iranicus</emphasis>
</taxonomicName>
is more widely distributed in the north-western Indian Ocean, although it has not been collected at other sampled localities along the eastern and southern coasts of
<collectingCountry id="C8D44FC4FFBCFFA3FAC9AF6A2BBCFEE4" box="[1368,1436,295,321]" name="Oman" pageId="12" pageNumber="191">Oman</collectingCountry>
(
<collectingRegion id="7207C1B6FFBCFFA3FF0EAF062ED0FEC0" box="[159,240,331,357]" country="Oman" name="Zufar" pageId="12" pageNumber="191">Dhofar</collectingRegion>
and Masirah Island) (A. Anker, pers. obs.).
</paragraph>
<paragraph id="B07C0F54FFBCFFA3FF56AF222A6DFD68" blockId="12.[151,1437,151,719]" pageId="12" pageNumber="191">
All Omani specimens of
<taxonomicName id="77C374D7FFBCFFA3FE7AAF222C49FE2D" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[491,617,367,393]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBCFFA3FE7AAF222C49FE2D" box="[491,617,367,393]" italics="true" pageId="12" pageNumber="191">A. iranicus</emphasis>
</taxonomicName>
are females, including one ovigerous female (
<figureCitation id="28F813D1FFBCFFA3FB12AF222AEAFE2C" box="[1155,1226,366,393]" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="12" pageNumber="191">Fig. 5</figureCitation>
). They present no significant discrepancies from the description of the male
<typeStatus id="6F78B1F6FFBCFFA3FCA6AFDE2DBDFE08" box="[823,925,403,429]" pageId="12" pageNumber="191" type="holotype">holotype</typeStatus>
. In all but
<specimenCount id="A6C5C4DDFFBCFFA3FB8FAFDE2A9DFE08" box="[1054,1213,403,429]" count="1" pageId="12" pageNumber="191" type="generic">one specimen</specimenCount>
, the chelae possess two distinct areas of fine dense setae, as described above; these setae are only missing (or at least are very poorly developed) in the smallest female (cl
<quantity id="773BA2B1FFBCFFA3FDAFAF972CB6FE50" box="[574,662,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="191" unit="mm" value="2.5">2.5 mm</quantity>
) (FLMNH UF 68861). The previously unknown colour pattern of
<taxonomicName id="77C374D7FFBCFFA3FA15AF962ED3FDBD" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFBCFFA3FA15AF962ED3FDBD" italics="true" pageId="12" pageNumber="191">A. iranicus</emphasis>
</taxonomicName>
(
<figureCitation id="28F813D1FFBCFFA3FE95AFB22F68FDBC" box="[260,328,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="12" pageNumber="191">Fig. 5</figureCitation>
) is most similar to the colour patterns of
<taxonomicName id="77C374D7FFBCFFA3FCB7AFB22A49FDBC" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[806,1129,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="shawnsmithi">
<emphasis id="82B7D346FFBCFFA3FCB7AFB22DF3FDBC" box="[806,979,511,537]" italics="true" pageId="12" pageNumber="191">A. shawnsmithi</emphasis>
Anker, 2011
</taxonomicName>
,
<taxonomicName id="77C374D7FFBCFFA3FBE4AFB22B4DFDBC" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[1141,1389,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="82B7D346FFBCFFA3FBE4AFB22AFAFDBC" box="[1141,1242,511,537]" italics="true" pageId="12" pageNumber="191">A. daviei</emphasis>
Anker, 2011
</taxonomicName>
(cf. Anker 2011: fig. 8A–F; Anker
<emphasis id="82B7D346FFBCFFA3FE63AC6E2C0AFD98" box="[498,554,547,573]" italics="true" pageId="12" pageNumber="191">et al.</emphasis>
2015: fig. 9) and the above-described
<taxonomicName id="77C374D7FFBCFFA3FC4AAC6E2AA7FD98" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[987,1159,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFBCFFA3FC4AAC6E2AA7FD98" box="[987,1159,547,573]" italics="true" pageId="12" pageNumber="191">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFBCFFA3FB1FAC6E2AC7FD98" bold="true" box="[1166,1255,547,573]" pageId="12" pageNumber="191">
<taxonomicNameLabel id="99846E3DFFBCFFA3FB1FAC6E2AC7FD98" box="[1166,1255,547,573]" pageId="12" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="28F813D1FFBCFFA3FB66AC6E2B1BFD98" box="[1271,1339,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-34@6.[151,1435,181,1859]" captionTargetPageId="6" captionText="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="12" pageNumber="191">Fig. 3</figureCitation>
). On the other hand, the stomatopod-associated species
<taxonomicName id="77C374D7FFBCFFA3FD3EAC0A2A18FDC4" authority="Anker &amp; Komai, 2010" authorityName="Anker &amp; Komai" authorityYear="2010" box="[687,1080,583,609]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="ahyongi">
<emphasis id="82B7D346FFBCFFA3FD3EAC0A2D0AFDC4" box="[687,810,583,609]" italics="true" pageId="12" pageNumber="191">A. ahyongi</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FCA3AC0A2A18FDC4" author="Anker, A. &amp; Komai, T." box="[818,1080,583,609]" pageId="12" pageNumber="191" pagination="45 - 54" refId="ref20339" refString="Anker, A. &amp; Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker &amp; Komai, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="77C374D7FFBCFFA3FBD7AC0A2EEFFD20" authority="Duris &amp; Anker, 2014" authorityName="Duris &amp; Anker" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="manticolus">
<emphasis id="82B7D346FFBCFFA3FBD7AC0A2AC2FDC4" box="[1094,1250,583,609]" italics="true" pageId="12" pageNumber="191">A. manticolus</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FB7AAC052EEFFD20" author="Duris, Z. &amp; Anker, A." pageId="12" pageNumber="191" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="77C374D7FFBCFFA3FE99AC262CA0FD20" authority="Anker &amp; Duris 2022" authorityName="Anker &amp; Duris" authorityYear="2022" box="[264,640,619,647]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="philippei">
<emphasis id="82B7D346FFBCFFA3FE99AC262FACFD20" box="[264,396,619,645]" italics="true" pageId="12" pageNumber="191">A. philippei</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FE02AC262CA0FD20" author="Anker, A. &amp; Duris, Z." box="[403,640,619,647]" pageId="12" pageNumber="191" pagination="288 - 300" refId="ref20021" refString="Anker, A. &amp; Duris, Z. (2022) Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea). Zootaxa, 5094 (2), 288 - 300. https: // doi. org / 10.11646 / zootaxa. 5094.2.4" type="journal article" year="2022">Anker &amp; Ďuriš 2022</bibRefCitation>
</taxonomicName>
, as well as the
<taxonomicName id="77C374D7FFBCFFA3FCBEAC262DA9FD20" authorityName="Fabricius" authorityYear="1798" box="[815,905,619,645]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFBCFFA3FCBEAC262DA9FD20" box="[815,905,619,645]" italics="true" pageId="12" pageNumber="191">Alpheus</emphasis>
</taxonomicName>
-associated
<taxonomicName id="77C374D7FFBCFFA3FB9FAC262B4CFD20" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[1038,1388,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="alpheusophilus">
<emphasis id="82B7D346FFBCFFA3FB9FAC262AF8FD20" box="[1038,1240,619,645]" italics="true" pageId="12" pageNumber="191">A. alpheusophilus</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FB71AC262B4CFD20" author="Marin, I." box="[1248,1388,619,645]" pageId="12" pageNumber="191" pagination="50 - 62" refId="ref22879" refString="Marin, I. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin, 2017</bibRefCitation>
</taxonomicName>
and the above-reported
<taxonomicName id="77C374D7FFBCFFA3FEE3ACC22C17FD0C" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[370,567,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBCFFA3FEE3ACC22FABFD0C" box="[370,395,655,681]" italics="true" pageId="12" pageNumber="191">A.</emphasis>
cf.
<emphasis id="82B7D346FFBCFFA3FE26ACC22C17FD0C" box="[439,567,655,681]" italics="true" pageId="12" pageNumber="191">dentirostris</emphasis>
</taxonomicName>
, have very different colour patterns (cf.
<figureCitation id="28F813D1FFBCFFA3FB90ACC22A67FD0C" box="[1025,1095,655,681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1533,1557]" captionTargetBox="[151,1435,181,1510]" captionTargetId="figure-19@9.[151,1435,181,1510]" captionTargetPageId="9" captionText="FIGURE 4. Athanas cf. dentirostris Anker, Jeng &amp; Chan, 2001, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350781" httpUri="https://zenodo.org/record/10350781/files/figure.png" pageId="12" pageNumber="191">Fig. 4</figureCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FBC3ACC22B6DFD0C" author="Anker, A. &amp; Komai, T." box="[1106,1357,655,681]" pageId="12" pageNumber="191" pagination="45 - 54" refId="ref20339" refString="Anker, A. &amp; Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker &amp; Komai 2010</bibRefCitation>
: fig. 6;
<bibRefCitation id="D45272A5FFBCFFA3FF06ACF92FA1FD68" author="Duris, Z. &amp; Anker, A." box="[151,385,691,719]" pageId="12" pageNumber="191" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker 2014</bibRefCitation>
: fig. 6;
<bibRefCitation id="D45272A5FFBCFFA3FE44ACFE2C7DFD68" author="Marin, I." box="[469,605,691,717]" pageId="12" pageNumber="191" pagination="50 - 62" refId="ref22879" refString="Marin, I. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin 2017</bibRefCitation>
: fig. 6;
<bibRefCitation id="D45272A5FFBCFFA3FD21ACFE2DBBFD68" author="Anker, A. &amp; Duris, Z." box="[688,923,691,719]" pageId="12" pageNumber="191" pagination="288 - 300" refId="ref20021" refString="Anker, A. &amp; Duris, Z. (2022) Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea). Zootaxa, 5094 (2), 288 - 300. https: // doi. org / 10.11646 / zootaxa. 5094.2.4" type="journal article" year="2022">Anker &amp; Ďuriš 2022</bibRefCitation>
: figs. 5, 6A, B).
</paragraph>
</subSubSection>
</treatment>
</document>
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