treatments-xml/data/38/6A/BE/386ABE42FFA0FFBCFF06AD0B2F63F9AC.xml
2024-06-21 12:33:38 +02:00

586 lines
80 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="FF37A78D60ECF93879995216727640A5" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFA0FFBCFF06AD0B2F63F9AC" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanopsis brevirostris Banner &amp; Banner 1981" docType="treatment" docVersion="3" lastPageNumber="198" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="195" updateTime="1702658426235" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="D4C185EB1ED23822BFA88230B3451890" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="98FE4D579208C34F45EF44014B72259A">
<mods:title id="EAED8517F3C8A714C736E40ADF80C46B">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="7783C193C9E9D09F6E17F31C4B6545F7" type="personal">
<mods:role id="011AAEEF58E72A1C71784FECBE641A5A">
<mods:roleTerm id="47490CE1559E5D2D4294664B5A8647F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B05D6DE2A82CA7494A318D8F463A05B">Anker, Arthur</mods:namePart>
<mods:affiliation id="C53C41332833943DC459CB41986B275C">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="483308B345A09198EBB7EC804AED4CA7">text</mods:typeOfResource>
<mods:relatedItem id="EE68CFF8A83D9E643D0831951079EAA7" type="host">
<mods:titleInfo id="4BBCEA7708E3041815ED6AA8AC677950">
<mods:title id="D01A02DA2E64A65899D17DACAB9D866F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8CFA7F5822F9359B89429AD741DE5066">
<mods:date id="D95B64127DF5E96E79EC3083383D538A">2023</mods:date>
<mods:detail id="321DB8A2178B2B1E89347D69AB80B57E" type="pubDate">
<mods:number id="CEDB2354C07128D121C9E137A17DBD3C">2023-12-11</mods:number>
</mods:detail>
<mods:detail id="85A381D723B7805F729234EA8CDEF1BA" type="volume">
<mods:number id="C237C444CD281068C3D3857BAEBA0C4E">5383</mods:number>
</mods:detail>
<mods:detail id="87AF34D16B028580ED7B84C9E5F0A73F" type="issue">
<mods:number id="ADA6A38CC5D46871CFE3E5C9E44F7428">2</mods:number>
</mods:detail>
<mods:extent id="CC4DA2FC95DC49DFD67FA261F1D5D120" unit="page">
<mods:start id="950C1F7C513626D73A4CDA7D5EA0C6CD">179</mods:start>
<mods:end id="A79220A84AA0491DEFE3954F8463D4BF">215</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4D6E7143FCC0B530EB982EB2C06674DB">
<mods:url id="E8A33F622A19AF61AC06A4281F75E99E">https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448</mods:url>
</mods:location>
<mods:classification id="4BC983BC8E33007AC9DD45FF8D3DBE78">journal article</mods:classification>
<mods:identifier id="A4A32474388A69C5A1F18A7538FB5CA6" type="DOI">10.11646/zootaxa.5383.2.5</mods:identifier>
<mods:identifier id="1C5254CA4E5527820CBFE202897C3A2D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D27DA867DC3F78017D04750D6AD373E6" type="Zenodo-Dep">10350767</mods:identifier>
<mods:identifier id="6317AE519598DD0BCD0CA4C2F84A16C4" type="ZooBank">7E434B40-6346-4E6D-BC55-38EBAE24BD52</mods:identifier>
</mods:mods>
<treatment id="386ABE42FFA0FFBCFF06AD0B2F63F9AC" LSID="urn:lsid:plazi:treatment:386ABE42FFA0FFBCFF06AD0B2F63F9AC" httpUri="http://treatment.plazi.org/id/386ABE42FFA0FFBCFF06AD0B2F63F9AC" lastPageId="19" lastPageNumber="198" pageId="16" pageNumber="195">
<subSubSection id="F8D95CDFFFA0FFBFFF06AD0B2CFDFCC5" box="[151,733,838,865]" pageId="16" pageNumber="195" type="nomenclature">
<paragraph id="B07C0F54FFA0FFBFFF06AD0B2CFDFCC5" blockId="16.[151,733,838,900]" box="[151,733,838,865]" pageId="16" pageNumber="195">
<heading id="EB34B838FFA0FFBFFF06AD0B2CFDFCC5" bold="true" box="[151,733,838,865]" fontSize="11" level="1" pageId="16" pageNumber="195" reason="1">
<taxonomicName id="77C374D7FFA0FFBFFF06AD0B2CFDFCC5" ID-CoL="85MDB" authority="Banner &amp; Banner, 1981" authorityName="Banner &amp; Banner" authorityYear="1981" box="[151,733,838,865]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="195" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA0FFBFFF06AD0B2CFDFCC5" bold="true" box="[151,733,838,865]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF06AD0B2F8FFCC4" bold="true" box="[151,431,838,865]" italics="true" pageId="16" pageNumber="195">Athanopsis brevirostris</emphasis>
<bibRefCitation id="D45272A5FFA0FFBFFE27AD0B2CFDFCC5" author="Banner, D. M. &amp; Banner, A. H." box="[438,733,838,864]" pageId="16" pageNumber="195" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner, 1981</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA0FFBFFF06AD272F33FC21" box="[151,275,874,900]" pageId="16" pageNumber="195" type="description">
<paragraph id="B07C0F54FFA0FFBFFF06AD272F33FC21" blockId="16.[151,733,838,900]" box="[151,275,874,900]" pageId="16" pageNumber="195">
(
<figureCitation id="28F813D1FFA0FFBFFF0EAD272F2BFC21" box="[159,267,874,900]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="16" pageNumber="195">Fig. 8a, b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA0FFBFFF06ADE22A1DFC63" box="[151,1085,942,967]" pageId="16" pageNumber="195" type="reference_group">
<paragraph id="B07C0F54FFA0FFBFFF06ADE22A1DFC63" blockId="16.[151,1085,942,967]" box="[151,1085,942,967]" pageId="16" pageNumber="195">
<treatmentCitationGroup id="90D3287AFFA0FFBFFF06ADE22A1DFC63" box="[151,1085,942,967]" pageId="16" pageNumber="195">
<treatmentCitation id="31622945FFA0FFBFFF06ADE22CBEFC63" author="Banner, D. M. &amp; Banner, A. H." box="[151,670,942,967]" page="45" pageId="16" pageNumber="195" year="1981">
<taxonomicName id="77C374D7FFA0FFBFFF06ADE22CBEFC63" ID-CoL="85MDB" authority="Banner &amp; Banner 1981: 45" authorityName="Banner &amp; Banner" authorityPageNumber="45" authorityYear="1981" box="[151,670,942,967]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="195" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA0FFBFFF06ADE22FA2FC62" box="[151,386,943,967]" italics="true" pageId="16" pageNumber="195">Athanopsis brevirostris</emphasis>
<bibRefCitation id="D45272A5FFA0FFBFFE18ADE22CBEFC63" author="Banner, D. M. &amp; Banner, A. H." box="[393,670,942,967]" pageId="16" pageNumber="195" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 45</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 5;
<treatmentCitation id="31622945FFA0FFBFFD78ADE22DAAFC63" ID-DOI="http://doi.org/10.5281/zenodo.6178250" ID-Zenodo-Dep="6178250" author="Anker, A." box="[745,906,942,967]" httpUri="http://treatment.plazi.org/id/039B87C5FF91DF0BFF00E03B2C5039AF" page="51" pageId="16" pageNumber="195" year="2012">
<bibRefCitation id="D45272A5FFA0FFBFFD78ADE22DAAFC63" author="Anker, A." box="[745,906,942,967]" pageId="16" pageNumber="195" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012: 51</bibRefCitation>
</treatmentCitation>
, figs. 24, 7A, B.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA0FFBFFF06ADB82D6DF932" pageId="16" pageNumber="195" type="materials_examined">
<paragraph id="B07C0F54FFA0FFBFFF06ADB82FD7FADD" blockId="16.[151,1437,1013,1903]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF06ADB82F5BFBAA" bold="true" box="[151,379,1013,1039]" pageId="16" pageNumber="195">Material examined</emphasis>
. 1 ovig. female (cl 5.0 mm),
<collectionCode id="D6D29791FFA0FFBFFD41ADB82D12FBAA" box="[720,818,1013,1039]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<collectionCode id="D6D29791FFA0FFBFFCAAADB82D7FFBAA" box="[827,863,1013,1039]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF</collectionCode>
71444,
<collectingCountry id="C8D44FC4FFA0FFBFFC2CADB82A73FBAA" box="[957,1107,1013,1039]" name="Saudi Arabia" pageId="16" pageNumber="195">Saudi Arabia</collectingCountry>
, Thuwal, near
<collectionCode id="D6D29791FFA0FFBFFB6FADB82B7BFBAA" box="[1278,1371,1013,1039]" pageId="16" pageNumber="195">KAUST</collectionCode>
King Abdullah Monument,
<geoCoordinate id="D5F76993FFA0FFBFFE05AA542C12FB96" box="[404,562,1049,1075]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="16" pageNumber="195" precision="1" seconds="26.2" value="22.340612">22°2026.2”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA0FFBFFDDCAA542CCBFB91" box="[589,747,1049,1076]" degrees="39" direction="east" minutes="05" orientation="longitude" pageId="16" pageNumber="195" precision="1" seconds="15.1" value="39.08753">39°0515.1”E</geoCoordinate>
, shallow subtidal sandflat with seagrass and rubble patches, depth
<quantity id="773BA2B1FFA0FFBFFF4AAA702F6DFBF2" box="[219,333,1085,1112]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="16" pageNumber="195" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.51.5 m</quantity>
, suction pump, in echiuran burrow, leg. A. Anker,
<date id="C47D2994FFA0FFBFFCE6AA702DD5FBF2" box="[887,1013,1085,1111]" pageId="16" pageNumber="195" value="2023-01-20">20.01.2023</date>
[fcn AA-22-497];
<materialsCitation id="00AB0509FFA0FFBFFB55AA702BB8FBDE" collectionCode="FLMNH" pageId="16" pageNumber="195" specimenCode="UF 71426" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA0FFBFFB55AA702B2EFBF2" box="[1220,1294,1085,1111]" count="1" pageId="16" pageNumber="195" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA0FFBFFAA9AA702BB0FBF2" box="[1336,1424,1085,1111]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="16" pageNumber="195" unit="mm" value="4.1">4.1 mm</quantity>
), 1 ovig. female (cl
<quantity id="773BA2B1FFA0FFBFFEF8AA2C2FE4FBDE" box="[361,452,1121,1147]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="195" unit="mm" value="4.4">4.4 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFE49AA2C2C1AFBDE" box="[472,570,1121,1147]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFDD3AA2C2C97FBDE" box="[578,695,1121,1147]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF 71426</specimenCode>
, same collection data as for previous specimen [fcn AA-22-474]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFF06AAC82B65FB3A" box="[151,1349,1157,1183]" collectionCode="FLMNH" pageId="16" pageNumber="195" specimenCode="IUF 71429" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA0FFBFFF06AAC82EC2FB3A" box="[151,226,1157,1183]" count="1" pageId="16" pageNumber="195" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA0FFBFFE9EAAC82F49FB3A" box="[271,361,1157,1183]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="195" unit="mm" value="3.6">3.6 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFEEDAAC82FFEFB3A" box="[380,478,1157,1183]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFE74AAC82C43FB3A" box="[485,611,1157,1183]" collectionCode="IUF" pageId="16" pageNumber="195">IUF 71429</specimenCode>
, same collection data as for previous specimens [fcn AA-22-489]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFAC0AAC82F78FAA9" collectingDate="2023-01-21" collectionCode="FLMNH" collectorName="A. Anker" pageId="16" pageNumber="195" specimenCode="UF 71441, UF 71442" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA0FFBFFAC0AAC82BBCFB3A" box="[1361,1436,1157,1183]" count="1" pageId="16" pageNumber="195" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA0FFBFFF2DAAE42F37FB66" box="[188,279,1193,1219]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="195" unit="mm" value="3.9">3.9 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFEBBAAE42FACFB66" box="[298,396,1193,1219]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFE05AAE42C28FB66" box="[404,520,1193,1219]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF 71441</specimenCode>
, same collection data as for previous specimens [fcn AA-22-467]; 1 ovig. female (cl
<quantity id="773BA2B1FFA0FFBFFF2FAA802F3BFB42" box="[190,283,1229,1255]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="195" unit="mm" value="4.4">4.4 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFEA0AA802FB3FB42" box="[305,403,1229,1255]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFE0DAA802C33FB42" box="[412,531,1229,1255]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF 71442</specimenCode>
, same collection locality as for previous specimens, leg.
<collectorName id="1D366A82FFA0FFBFFB34AA802B31FB42" box="[1189,1297,1229,1255]" pageId="16" pageNumber="195">A. Anker</collectorName>
,
<date id="C47D2994FFA0FFBFFA8FAA802BBCFB42" box="[1310,1436,1229,1255]" pageId="16" pageNumber="195" value="2023-01-21">
<collectingDate id="D439D07CFFA0FFBFFA8FAA802BBCFB42" box="[1310,1436,1229,1255]" pageId="16" pageNumber="195" value="2023-01-21">21.01.2023</collectingDate>
</date>
[fcn AA-22-500]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFEF2AABC2ED2FA95" collectingDate="2023-01-21" collectionCode="FLMNH" collectorName="A. Anker" pageId="16" pageNumber="195" specimenCode="UF 71427" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA0FFBFFEF2AABC2F8EFAAE" box="[355,430,1265,1291]" count="1" pageId="16" pageNumber="195" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA0FFBFFE4BAABC2C14FAAE" box="[474,564,1265,1291]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="195" unit="mm" value="4.4">4.4 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFDD6AABC2C89FAAE" box="[583,681,1265,1291]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFD21AABC2D03FAAE" box="[688,803,1265,1291]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF 71427</specimenCode>
, same collection data as for previous specimen [fcn AA-22-511]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFF6EAB582FD2FADD" collectingDate="2013-06-03" collectionCode="FLMNH" collectorName="A. Anker" country="Saudi Arabia" elevation="8" latitude="18.220556" location="Farasan Islands" longLatPrecision="1" longitude="41.324387" municipality="Marca Island" pageId="16" pageNumber="195" specimenCode="UF 36637" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFA0FFBFFF6EAB582F41FA8A" box="[255,353,1301,1327]" count="1" pageId="16" pageNumber="195" type="female">1 female</specimenCount>
(cl ~
<quantity id="773BA2B1FFA0FFBFFE0FAB582FDAFA8A" box="[414,506,1301,1328]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="195" unit="mm" value="3.5">3.5 mm</quantity>
),
<collectionCode id="D6D29791FFA0FFBFFD9FAB582C50FA8A" box="[526,624,1301,1327]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="16" pageNumber="195" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA0FFBFFDE8AB582CCEFA8A" box="[633,750,1301,1327]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="16" pageNumber="195" type="Museum">UF 36637</specimenCode>
,
<collectingCountry id="C8D44FC4FFA0FFBFFD6BAB582DB0FA8A" box="[762,912,1301,1327]" name="Saudi Arabia" pageId="16" pageNumber="195">Saudi Arabia</collectingCountry>
,
<location id="B51C598FFFA0FFBFFC0DAB582A6EFA8A" LSID="urn:lsid:plazi:treatment:386ABE42FFA0FFBCFF06AD0B2F63F9AC:B51C598FFFA0FFBFFC0DAB582A6EFA8A" box="[924,1102,1301,1327]" country="Saudi Arabia" latitude="18.220556" longLatPrecision="1" longitude="41.324387" municipality="Marca Island" name="Farasan Islands" pageId="16" pageNumber="195">Farasan Islands</location>
,
<collectingMunicipality id="5018952EFFA0FFBFFBCBAB582AD1FA8A" box="[1114,1265,1301,1327]" pageId="16" pageNumber="195">Marca Island</collectingMunicipality>
,
<geoCoordinate id="D5F76993FFA0FFBFFB6FAB582BBCFA8A" box="[1278,1436,1301,1327]" degrees="18" direction="north" minutes="13" orientation="latitude" pageId="16" pageNumber="195" precision="1" seconds="14.0" value="18.220556">18°1314.0”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA0FFBFFF39AB742F63FAF6" box="[168,323,1337,1363]" degrees="41" direction="east" minutes="19" orientation="longitude" pageId="16" pageNumber="195" precision="1" seconds="27.8" value="41.324387">41°1927.8”E</geoCoordinate>
, subtidal sandflat, depth
<quantity id="773BA2B1FFA0FFBFFDF2AB742C9CFAF6" box="[611,700,1337,1364]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="16" pageNumber="195" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">
<elevation id="3BEEE867FFA0FFBFFDF2AB742C9CFAF6" box="[611,700,1337,1364]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="16" pageNumber="195" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">510 m</elevation>
</quantity>
, suction pump, in burrow of unknown host, leg.
<collectorName id="1D366A82FFA0FFBFFB7BAB742B73FAF6" box="[1258,1363,1337,1363]" pageId="16" pageNumber="195">A. Anker</collectorName>
<emphasis id="82B7D346FFA0FFBFFACDAB772BAEFAF6" box="[1372,1422,1337,1363]" italics="true" pageId="16" pageNumber="195">et al</emphasis>
.,
<date id="C47D2994FFA0FFBFFF06AB102F35FAD2" box="[151,277,1373,1399]" pageId="16" pageNumber="195" value="2013-06-03">
<collectingDate id="D439D07CFFA0FFBFFF06AB102F35FAD2" box="[151,277,1373,1399]" pageId="16" pageNumber="195" value="2013-06-03">06.03.2013</collectingDate>
</date>
[fcn BDJRS-1756]
</materialsCitation>
.
</paragraph>
<paragraph id="B07C0F54FFA0FFBFFF56ABCC2D6DF932" blockId="16.[151,1437,1013,1903]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF56ABCC2FF7FA3E" bold="true" box="[199,471,1409,1435]" pageId="16" pageNumber="195">Comparative material</emphasis>
.
<taxonomicName id="77C374D7FFA0FFBFFE7AABCC2D5EFA3E" authority="Anker, 2011: 1" authorityName="Anker" authorityPageNumber="1" authorityYear="2011" box="[491,894,1409,1435]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="195" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA0FFBFFE7AABCC2CE0FA3E" box="[491,704,1409,1435]" italics="true" pageId="16" pageNumber="195">Athanopsis saurus</emphasis>
Anker, 2011:
<materialsCitation id="00AB0509FFA0FFBFFCE1ABCC2D5EFA3E" box="[880,894,1409,1435]" collectingDate="2018-08-09" collectionCode="KM" collectorName="A. Anker" country="New Caledonia" county="Expedition Koumac" elevation="1" latitude="-20.583334" location="Koumac" longLatPrecision="929" longitude="164.275" municipality="Pointe Pandop" pageId="16" pageNumber="195" specimenCode="KM601" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFA0FFBFFCE1ABCC2D5EFA3E" box="[880,894,1409,1435]" count="1" pageId="16" pageNumber="195" type="female">1</specimenCount>
</materialsCitation>
</taxonomicName>
female (cl
<quantity id="773BA2B1FFA0FFBFFB9EABCC2A50FA3E" box="[1039,1136,1409,1435]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="195" unit="mm" value="3.6">3.6 mm</quantity>
), MNHN-2018-571,
<collectingCountry id="C8D44FC4FFA0FFBFFAF6ABCC2F2BFA1A" name="New Caledonia" pageId="16" pageNumber="195">New Caledonia</collectingCountry>
,
<location id="B51C598FFFA0FFBFFE88ABE82F5BFA1A" LSID="urn:lsid:plazi:treatment:386ABE42FFA0FFBCFF06AD0B2F63F9AC:B51C598FFFA0FFBFFE88ABE82F5BFA1A" box="[281,379,1445,1471]" country="New Caledonia" county="Expedition Koumac" latitude="-20.583334" longLatPrecision="929" longitude="164.275" municipality="Pointe Pandop" name="Koumac" pageId="16" pageNumber="195">Koumac</location>
,
<collectingMunicipality id="5018952EFFA0FFBFFE19ABE82C10FA1A" box="[392,560,1445,1471]" pageId="16" pageNumber="195">Pointe Pandop</collectingMunicipality>
,
<collectingCounty id="591D77D8FFA0FFBFFDAFABE82D01FA1A" box="[574,801,1445,1471]" pageId="16" pageNumber="195">Expedition Koumac</collectingCounty>
2.1 sta.
<specimenCode id="E065A72FFFA0FFBFFC17ABE82DC0FA1A" box="[902,992,1445,1471]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="16" pageNumber="195">KM601</specimenCode>
,
<geoCoordinate id="D5F76993FFA0FFBFFC7FABE82A6AFA65" box="[1006,1098,1445,1472]" degrees="20" direction="south" minutes="35" orientation="latitude" pageId="16" pageNumber="195" precision="925" value="-20.583334">20°35S</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA0FFBFFBF4ABE82AC7FA65" box="[1125,1255,1445,1472]" degrees="164" direction="east" minutes="16.5" orientation="longitude" pageId="16" pageNumber="195" precision="92" value="164.275">164°16.5E</geoCoordinate>
, partly exposed sand-mud intertidal with coral rubble fragments, depth
<quantity id="773BA2B1FFA0FFBFFC99AB842D47FA46" box="[776,871,1481,1507]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="16" pageNumber="195" unit="m" value="0.1" valueMax="0.2" valueMin="0.0">
<elevation id="3BEEE867FFA0FFBFFC99AB842D47FA46" box="[776,871,1481,1507]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="16" pageNumber="195" unit="m" value="0.1" valueMax="0.2" valueMin="0.0">00.2 m</elevation>
</quantity>
, suction pump, in echiuran burrow, leg.
<collectorName id="1D366A82FFA0FFBFFABEAB842BB8FA46" box="[1327,1432,1481,1507]" pageId="16" pageNumber="195">A. Anker</collectorName>
,
<date id="C47D2994FFA0FFBFFF06ABA02F39F9A2" box="[151,281,1517,1543]" pageId="16" pageNumber="195" value="2018-08-09">
<collectingDate id="D439D07CFFA0FFBFFF06ABA02F39F9A2" box="[151,281,1517,1543]" pageId="16" pageNumber="195" value="2018-08-09">08.09.2018</collectingDate>
</date>
;
<materialsCitation id="00AB0509FFA0FFBFFEB6ABA02B61F9EA" collectingDate="2018-09-18" collectionCode="KM" collectorName="A. Anker &amp; Z. Duris" country="New Caledonia" county="Pouebo river" elevation="3" latitude="-20.38" location="Saint-Mathieu" longLatPrecision="872" longitude="164.58333" municipality="Miscellanea" pageId="16" pageNumber="195" specimenCode="KM606" specimenCount="1" specimenCount-female="1">
<specimenCount id="A6C5C4DDFFA0FFBFFEB6ABA02FABF9A2" box="[295,395,1517,1543]" count="1" pageId="16" pageNumber="195" type="female">1 female</specimenCount>
(cl 5.0 mm), MNHN-2018-1012,
<collectingCountry id="C8D44FC4FFA0FFBFFC88ABA02DECF9A2" box="[793,972,1517,1543]" name="New Caledonia" pageId="16" pageNumber="195">New Caledonia</collectingCountry>
,
<location id="B51C598FFFA0FFBFFC4BABA02A5DF9A2" LSID="urn:lsid:plazi:treatment:386ABE42FFA0FFBCFF06AD0B2F63F9AC:B51C598FFFA0FFBFFC4BABA02A5DF9A2" box="[986,1149,1517,1543]" country="New Caledonia" county="Pouebo river" latitude="-20.38" longLatPrecision="872" longitude="164.58333" municipality="Miscellanea" name="Saint-Mathieu" pageId="16" pageNumber="195">Saint-Mathieu</location>
,
<location id="B51C598FFFA0FFBFFB1AABA02B4FF9A2" LSID="urn:lsid:plazi:treatment:386ABE42FFA0FFBCFF06AD0B2F63F9AC:B51C598FFFA0FFBFFB1AABA02B4FF9A2" box="[1163,1391,1517,1543]" country="New Caledonia" county="Pouebo river" latitude="-20.38" longLatPrecision="872" longitude="164.58333" municipality="Miscellanea" name="Expedition Koumac" pageId="16" pageNumber="195">Expedition Koumac</location>
2.1
<collectingMunicipality id="5018952EFFA0FFBFFF06A85C2F3EF98E" box="[151,286,1553,1579]" pageId="16" pageNumber="195">Miscellanea</collectingMunicipality>
sta.
<specimenCode id="E065A72FFFA0FFBFFEC6A85C2F91F98E" box="[343,433,1553,1579]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="16" pageNumber="195">KM606</specimenCode>
,
<geoCoordinate id="D5F76993FFA0FFBFFE2FA85C2C0FF98E" box="[446,559,1553,1579]" degrees="20" direction="south" minutes="22.8" orientation="latitude" pageId="16" pageNumber="195" precision="92" value="-20.38">20°22.8S</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA0FFBFFDD8A85C2C97F989" box="[585,695,1553,1580]" degrees="164" direction="east" minutes="35" orientation="longitude" pageId="16" pageNumber="195" precision="925" value="164.58333">164°35E</geoCoordinate>
, shallow sandflat with sparse seagrass near
<collectingCounty id="591D77D8FFA0FFBFFB21A85C2B62F98E" box="[1200,1346,1553,1579]" pageId="16" pageNumber="195">Pouébo river</collectingCounty>
estuary and mangroves, depth
<quantity id="773BA2B1FFA0FFBFFE08A8782FD8F9EA" box="[409,504,1589,1616]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="16" pageNumber="195" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">
<elevation id="3BEEE867FFA0FFBFFE08A8782FD8F9EA" box="[409,504,1589,1616]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="16" pageNumber="195" unit="m" value="0.25" valueMax="0.5" valueMin="0.0">00.5 m</elevation>
</quantity>
, suction pump, in echiuran burrow, leg.
<collectorName id="1D366A82FFA0FFBFFC52A8782A0BF9EA" box="[963,1067,1589,1615]" pageId="16" pageNumber="195">A. Anker</collectorName>
&amp;
<collectorName id="1D366A82FFA0FFBFFBC0A8782A8FF9F4" box="[1105,1199,1589,1617]" pageId="16" pageNumber="195">Z. Ďuriš</collectorName>
,
<date id="C47D2994FFA0FFBFFB2EA8782B61F9EA" box="[1215,1345,1589,1615]" pageId="16" pageNumber="195" value="2018-09-18">
<collectingDate id="D439D07CFFA0FFBFFB2EA8782B61F9EA" box="[1215,1345,1589,1615]" pageId="16" pageNumber="195" value="2018-09-18">18.09.2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFADCA8782A72F9D6" collectingDate="2018-09-18" collectionCode="KM" collectorName="A. Anker &amp; Z. Duris" country="New Caledonia" county="Pouebo river" elevation="3" latitude="-20.38" location="Saint-Mathieu" longLatPrecision="872" longitude="164.58333" municipality="Miscellanea" pageId="16" pageNumber="195" specimenCount="1">
1 ovig. female (cl
<quantity id="773BA2B1FFA0FFBFFE9CA8142F47F9D6" box="[269,359,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="195" unit="mm" value="4.9">4.9 mm</quantity>
), MNHN-2018-978, same collection data as for previous specimen
</materialsCitation>
;
<materialsCitation id="00AB0509FFA0FFBFFBCCA8142D68F932" collectingDate="2018-09-18" collectionCode="KM" collectorName="A. Anker &amp; Z. Duris" country="New Caledonia" county="Pouebo river" elevation="3" latitude="-20.38" location="Saint-Mathieu" longLatPrecision="872" longitude="164.58333" municipality="Miscellanea" pageId="16" pageNumber="195" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA0FFBFFBCCA8142A87F9D6" box="[1117,1191,1625,1651]" count="1" pageId="16" pageNumber="195" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA0FFBFFB43A8142B0BF9D6" box="[1234,1323,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="195" unit="mm" value="4.3">4.3 mm</quantity>
), MNHN- IU-2018-974, same collection data as for previous specimens
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA0FFBFFF56A8EC2AA8F882" pageId="16" pageNumber="195" type="description">
<paragraph id="B07C0F54FFA0FFBFFF56A8EC2EF5F8A6" blockId="16.[151,1437,1013,1903]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF56A8EC2F71F91E" bold="true" box="[199,337,1697,1723]" pageId="16" pageNumber="195">Description</emphasis>
. See
<bibRefCitation id="D45272A5FFA0FFBFFE06A8EC2C99F91E" author="Banner, D. M. &amp; Banner, A. H." box="[407,697,1697,1723]" pageId="16" pageNumber="195" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981)</bibRefCitation>
for original description and illustrations, and
<bibRefCitation id="D45272A5FFA0FFBFFB44A8EC2B51F91E" author="Anker, A." box="[1237,1393,1697,1723]" pageId="16" pageNumber="195" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012)</bibRefCitation>
for additional illustrations of the Madagascan material; colour pattern of the Saudi Arabian specimens is shown in
<figureCitation id="28F813D1FFA0FFBFFAE1A8882EF1F8A6" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="16" pageNumber="195">Fig. 8a, b</figureCitation>
.
</paragraph>
<paragraph id="B07C0F54FFA0FFBFFF56A9402AA8F882" blockId="16.[151,1437,1013,1903]" box="[199,1160,1805,1832]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF56A9402F45F882" bold="true" box="[199,357,1805,1831]" pageId="16" pageNumber="195">Colour in life</emphasis>
. Described in detail in
<bibRefCitation id="D45272A5FFA0FFBFFDF8A9402D04F88D" author="Anker, A." box="[617,804,1805,1832]" pageId="16" pageNumber="195" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012: 53</bibRefCitation>
, fig. 7A, B); see also
<figureCitation id="28F813D1FFA0FFBFFB86A9402AA4F882" box="[1047,1156,1805,1831]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="16" pageNumber="195">Fig. 8a, b</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA0FFBFFF56A97C2A26F8CA" pageId="16" pageNumber="195" type="distribution">
<paragraph id="B07C0F54FFA0FFBFFF56A97C2A26F8CA" blockId="16.[151,1437,1013,1903]" pageId="16" pageNumber="195">
<emphasis id="82B7D346FFA0FFBFFF56A97C2F79F8EE" bold="true" box="[199,345,1841,1867]" pageId="16" pageNumber="195">Distribution</emphasis>
.
<collectingRegion id="7207C1B6FFA0FFBFFEF9A97C2FEAF8EE" box="[360,458,1841,1867]" country="Eritrea" name="Semenawi Keyih Bahri" pageId="16" pageNumber="195">Red Sea</collectingRegion>
:
<collectingCountry id="C8D44FC4FFA0FFBFFE45A97C2C05F8EE" box="[468,549,1841,1867]" name="Eritrea" pageId="16" pageNumber="195">Eritrea</collectingCountry>
: Dahlak Archipelago (
<bibRefCitation id="D45272A5FFA0FFBFFCB1A97C2A0BF8EE" author="Banner, D. M. &amp; Banner, A. H." box="[800,1067,1841,1867]" pageId="16" pageNumber="195" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
),
<collectingCountry id="C8D44FC4FFA0FFBFFBD1A97C2AF7F8EE" box="[1088,1239,1841,1867]" name="Saudi Arabia" pageId="16" pageNumber="195">Saudi Arabia</collectingCountry>
: Thuwal, Farasan Islands (present study);
<collectingCountry id="C8D44FC4FFA0FFBFFE32A9182C0FF8CA" box="[419,559,1877,1903]" name="Madagascar" pageId="16" pageNumber="195">Madagascar</collectingCountry>
: Nosy-Bé;
<collectingCountry id="C8D44FC4FFA0FFBFFD3DA9182CD0F8CA" box="[684,752,1877,1903]" name="Japan" pageId="16" pageNumber="195">Japan</collectingCountry>
:
<collectingRegion id="7207C1B6FFA0FFBFFD6BA9182D7FF8CA" box="[762,863,1877,1903]" country="Japan" name="Okinawa" pageId="16" pageNumber="195">Okinawa</collectingRegion>
(
<bibRefCitation id="D45272A5FFA0FFBFFCFFA9182DDAF8CA" author="Anker, A." box="[878,1018,1877,1903]" pageId="16" pageNumber="195" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="E4BC5FDCFFA1FFBEFF06A9382B13F870" ID-DOI="http://doi.org/10.5281/zenodo.10350797" ID-Zenodo-Dep="10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="17" pageNumber="196" startId="17.[151,250,1909,1933]" targetBox="[151,1435,181,1885]" targetPageId="17" targetType="figure">
<paragraph id="B07C0F54FFA1FFBEFF06A9382B13F870" blockId="17.[151,1436,1909,2005]" pageId="17" pageNumber="196">
<emphasis id="82B7D346FFA1FFBEFF06A9382F38F828" bold="true" box="[151,280,1909,1933]" pageId="17" pageNumber="196">FIGURE 8.</emphasis>
<taxonomicName id="77C374D7FFA1FFBEFEB2A9382D00F828" authority="Banner &amp; Banner, 1981" authorityName="Banner &amp; Banner" authorityYear="1981" box="[291,800,1909,1933]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="196" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA1FFBEFEB2A9382C33F828" box="[291,531,1909,1933]" italics="true" pageId="17" pageNumber="196">Athanopsis brevirostris</emphasis>
<bibRefCitation id="D45272A5FFA1FFBEFD8FA93B2D00F828" author="Banner, D. M. &amp; Banner, A. H." box="[542,800,1909,1933]" pageId="17" pageNumber="196" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner, 1981</bibRefCitation>
</taxonomicName>
, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably
<taxonomicName id="77C374D7FFA1FFBEFF06A9F02F61F870" box="[151,321,1981,2005]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="17" pageNumber="196" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFA1FFBEFF06A9F02F3DF870" box="[151,285,1981,2005]" italics="true" pageId="17" pageNumber="196">Ochetostoma</emphasis>
sp.
</taxonomicName>
or
<taxonomicName id="77C374D7FFA1FFBEFEF3A9F02C20F870" box="[354,512,1981,2005]" class="Polychaeta" family="Thalassematidae" genus="Listriolobus" kingdom="Animalia" order="Echiuroidea" pageId="17" pageNumber="196" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFA1FFBEFEF3A9F02FFCF870" box="[354,476,1981,2005]" italics="true" pageId="17" pageNumber="196">Listriolobus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="77C374D7FFA1FFBEFD9FA9F02C9BF870" authorityName="Forbes &amp; Goodsir" authorityYear="1841" box="[526,699,1981,2005]" class="Polychaeta" family="Thalassematidae" kingdom="Animalia" order="Echiuroidea" pageId="17" pageNumber="196" phylum="Annelida" rank="family">Thalassematidae</taxonomicName>
), photographed shortly after death. Photographs by the author.
</paragraph>
</caption>
<caption id="E4BC5FDCFFA2FFBDFF06A91B2FC2F87F" ID-DOI="http://doi.org/10.5281/zenodo.10350801" ID-Zenodo-Dep="10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="18" pageNumber="197" startId="18.[151,250,1878,1902]" targetBox="[151,1436,151,1860]" targetPageId="18" targetType="figure">
<paragraph id="B07C0F54FFA2FFBDFF06A91B2FC2F87F" blockId="18.[151,1437,1878,2010]" pageId="18" pageNumber="197">
<emphasis id="82B7D346FFA2FFBDFF06A91B2F33F8CB" bold="true" box="[151,275,1878,1902]" pageId="18" pageNumber="197">FIGURE 9.</emphasis>
<taxonomicName id="77C374D7FFA2FFBDFE88A91B2C79F8CB" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[281,601,1878,1902]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="197" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA2FFBDFE88A91B2FF4F8CB" box="[281,468,1878,1902]" italics="true" pageId="18" pageNumber="197">Athanopsis saurus</emphasis>
Anker, 2011
</taxonomicName>
, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [ae] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably
<taxonomicName id="77C374D7FFA2FFBDFECBA9D32C22F810" box="[346,514,1950,1974]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="197" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFA2FFBDFECBA9D32FC0F813" box="[346,480,1950,1974]" italics="true" pageId="18" pageNumber="197">Ochetostoma</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="77C374D7FFA2FFBDFD81A9D32C9DF813" authorityName="Forbes &amp; Goodsir" authorityYear="1841" box="[528,701,1950,1974]" class="Polychaeta" family="Thalassematidae" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="197" phylum="Annelida" rank="family">Thalassematidae</taxonomicName>
), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author.
</paragraph>
</caption>
<subSubSection id="F8D95CDFFFA3FFBCFF56AEDA2B4BFEC0" pageId="19" pageNumber="198" type="biology_ecology">
<paragraph id="B07C0F54FFA3FFBCFF56AEDA2B4BFEC0" blockId="19.[151,1437,151,1545]" pageId="19" pageNumber="198">
<emphasis id="82B7D346FFA3FFBCFF56AEDA2F06FF14" bold="true" box="[199,294,151,177]" pageId="19" pageNumber="198">Ecology</emphasis>
. Most Saudi Arabian specimens of
<taxonomicName id="77C374D7FFA3FFBCFD40AEDA2D55FF14" authorityName="Banner &amp; Banner" authorityYear="1981" box="[721,885,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFD40AEDA2D55FF14" box="[721,885,151,177]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
were collected together with echiuran hosts, a large reddish species possibly belonging to either the genus
<taxonomicName id="77C374D7FFA3FFBCFCD1AEF62A96FF70" authority="Ruppel &amp; Leuckart" authorityName="Ruppel &amp; Leuckart" box="[832,1206,187,213]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="19" pageNumber="198" phylum="Annelida" rank="genus">
<emphasis id="82B7D346FFA3FFBCFCD1AEF62DF4FF70" box="[832,980,187,213]" italics="true" pageId="19" pageNumber="198">Ochetostoma</emphasis>
Rüppel &amp; Leuckart
</taxonomicName>
or the related genus
<taxonomicName id="77C374D7FFA3FFBCFF06AE922F57FF5C" authority="Fischer" authorityName="Fischer" authorityYear="1926" box="[151,375,223,249]" class="Polychaeta" family="Thalassematidae" genus="Listriolobus" kingdom="Animalia" order="Echiuroidea" pageId="19" pageNumber="198" phylum="Annelida" rank="genus">
<emphasis id="82B7D346FFA3FFBCFF06AE922F3EFF5C" box="[151,286,223,249]" italics="true" pageId="19" pageNumber="198">Listriolobus</emphasis>
Fischer
</taxonomicName>
(
<taxonomicName id="77C374D7FFA3FFBCFE16AE922C64FF5C" authorityName="Forbes &amp; Goodsir" authorityYear="1841" box="[391,580,223,249]" class="Polychaeta" family="Thalassematidae" kingdom="Animalia" order="Echiuroidea" pageId="19" pageNumber="198" phylum="Annelida" rank="family">Thalassematidae</taxonomicName>
) (
<figureCitation id="28F813D1FFA3FFBCFDC8AE922C89FF5C" box="[601,681,223,249]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="19" pageNumber="198">Fig. 8c</figureCitation>
). Some of the previously reported specimens of
<taxonomicName id="77C374D7FFA3FFBCFB50AE922B40FF5C" authorityName="Banner &amp; Banner" authorityYear="1981" box="[1217,1376,223,249]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFB50AE922B40FF5C" box="[1217,1376,223,249]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
were also extracted from burrows of
<taxonomicName id="77C374D7FFA3FFBCFE6DAF4E2C97FEB8" box="[508,695,259,285]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="19" pageNumber="198" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFA3FFBCFE6DAF4E2CB0FEB8" box="[508,656,259,285]" italics="true" pageId="19" pageNumber="198">Ochetostoma</emphasis>
sp.
</taxonomicName>
or unknown hosts, however, the
<typeStatus id="6F78B1F6FFA3FFBCFBBEAF4E2AB1FEB8" box="[1071,1169,259,285]" pageId="19" pageNumber="198" type="holotype">holotype</typeStatus>
came from a poisoning station (
<bibRefCitation id="D45272A5FFA3FFBCFF7EAF6A2FD5FEE4" author="Banner, D. M. &amp; Banner, A. H." box="[239,501,295,321]" pageId="19" pageNumber="198" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
;
<bibRefCitation id="D45272A5FFA3FFBCFE6FAF6A2CA7FEE4" author="Anker, A." box="[510,647,295,321]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
). Thus, the present evidence suggests that all species of
<taxonomicName id="77C374D7FFA3FFBCFB61AF6A2B4CFEE4" authorityName="Coutiere" authorityYear="1897" box="[1264,1388,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFA3FFBCFB61AF6A2B4CFEE4" box="[1264,1388,295,321]" italics="true" pageId="19" pageNumber="198">Athanopsis</emphasis>
</taxonomicName>
live in symbiosis with echiurans (
<bibRefCitation id="D45272A5FFA3FFBCFE4FAF062C7DFEC0" author="Miya, Y." box="[478,605,331,357]" pageId="19" pageNumber="198" pagination="117 - 131" refId="ref23018" refString="Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 5, 117 - 131." type="journal article" year="1980">Miya 1980</bibRefCitation>
;
<bibRefCitation id="D45272A5FFA3FFBCFDF9AF062D32FEC0" author="Berggren, M." box="[616,786,331,357]" pageId="19" pageNumber="198" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991</bibRefCitation>
; Anker 2011, 2012;
<bibRefCitation id="D45272A5FFA3FFBCFC64AF062A9BFEC0" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[1013,1211,331,357]" pageId="19" pageNumber="198" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFA3FFBCFBD1AF062A52FEC0" box="[1088,1138,331,357]" italics="true" pageId="19" pageNumber="198">et al</emphasis>
. 2014
</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA3FFBCFF56AF222F63F9AC" pageId="19" pageNumber="198" type="discussion">
<paragraph id="B07C0F54FFA3FFBCFF56AF222BAEFD0C" blockId="19.[151,1437,151,1545]" pageId="19" pageNumber="198">
<emphasis id="82B7D346FFA3FFBCFF56AF222F14FE2C" bold="true" box="[199,308,367,393]" pageId="19" pageNumber="198">Remarks</emphasis>
.
<bibRefCitation id="D45272A5FFA3FFBCFED4AF222FC0FE2C" author="Anker, A." box="[325,480,367,393]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012)</bibRefCitation>
separated
<taxonomicName id="77C374D7FFA3FFBCFDF1AF222D24FE2C" authorityName="Banner &amp; Banner" authorityYear="1981" box="[608,772,367,393]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFDF1AF222D24FE2C" box="[608,772,367,393]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
from the closely related
<taxonomicName id="77C374D7FFA3FFBCFBBEAF222ABDFE2D" authorityName="Anker" authorityYear="2011" box="[1071,1181,367,393]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFBBEAF222ABDFE2D" box="[1071,1181,367,393]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
(type locality: Lizard Island, Great Barrier Reef) by the following four features: (1) rostrum short, with lateral margins broadly concave in
<emphasis id="82B7D346FFA3FFBCFF24AFFA2F5DFE75" box="[181,381,439,465]" italics="true" pageId="19" pageNumber="198">
<taxonomicName id="77C374D7FFA3FFBCFF24AFFA2F77FE74" authorityName="Banner &amp; Banner" authorityYear="1981" box="[181,343,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">A. brevirostris</taxonomicName>
vs.
</emphasis>
longer, with lateral margins proximally convex in
<taxonomicName id="77C374D7FFA3FFBCFC2EAFFA2A0AFE75" authorityName="Anker" authorityYear="2011" box="[959,1066,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFC2EAFFA2A0AFE75" box="[959,1066,439,465]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
; (2) second antennular peduncle much wider than long in
<emphasis id="82B7D346FFA3FFBCFE27AF962C5DFE51" box="[438,637,475,501]" italics="true" pageId="19" pageNumber="198">
<taxonomicName id="77C374D7FFA3FFBCFE27AF962C77FE50" authorityName="Banner &amp; Banner" authorityYear="1981" box="[438,599,475,501]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">A. brevirostris</taxonomicName>
vs.
</emphasis>
only slightly wider than long (almost square-shaped) in
<taxonomicName id="77C374D7FFA3FFBCFA90AF962B4CFE51" authorityName="Anker" authorityYear="2011" box="[1281,1388,475,501]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFA90AF962B4CFE51" box="[1281,1388,475,501]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
; (3) major chela palm with small field of setae distally in
<emphasis id="82B7D346FFA3FFBCFD62AFB22D99FDBD" box="[755,953,511,537]" italics="true" pageId="19" pageNumber="198">
<taxonomicName id="77C374D7FFA3FFBCFD62AFB22DB4FDBC" authorityName="Banner &amp; Banner" authorityYear="1981" box="[755,916,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">A. brevirostris</taxonomicName>
vs.
</emphasis>
with more extensive setation in
<taxonomicName id="77C374D7FFA3FFBCFABBAFB22BB5FDBD" authorityName="Anker" authorityYear="2011" box="[1322,1429,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFABBAFB22BB5FDBD" box="[1322,1429,511,537]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
; and (4) colour pattern with more white on the second and third pleonites and less red on the carapace in
<taxonomicName id="77C374D7FFA3FFBCFABAAC6E2BB6FD99" authorityName="Anker" authorityYear="2011" box="[1323,1430,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFABAAC6E2BB6FD99" box="[1323,1430,547,573]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
. As pointed out by
<bibRefCitation id="D45272A5FFA3FFBCFEF7AC0A2C21FDC4" author="Anker, A." box="[358,513,583,609]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012)</bibRefCitation>
, in the
<typeStatus id="6F78B1F6FFA3FFBCFDC2AC0A2C95FDC4" box="[595,693,583,609]" pageId="19" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFA3FFBCFD4AAC0A2D5CFDC4" authorityName="Banner &amp; Banner" authorityYear="1981" box="[731,892,583,609]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFD4AAC0A2D5CFDC4" box="[731,892,583,609]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
, as well as in the Madagascan material assigned to that species, the merus of the third to fifth pereiopods has a small distolateral tooth, which was not overlooked by
<bibRefCitation id="D45272A5FFA3FFBCFF06ACC22F92FD0C" author="Banner, D. M. &amp; Banner, A. H." box="[151,434,655,681]" pageId="19" pageNumber="198" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981)</bibRefCitation>
; this tooth is present, and sometimes quite conspicuous, in all Saudi Arabian specimens.
</paragraph>
<paragraph id="B07C0F54FFA3FFBCFF56ACFE2F63F9AC" blockId="19.[151,1437,151,1545]" pageId="19" pageNumber="198">
The New Caledonian material is morphologically somewhat intermediate between
<taxonomicName id="77C374D7FFA3FFBCFBCAACFE2ADAFD68" authorityName="Banner &amp; Banner" authorityYear="1981" box="[1115,1274,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFBCAACFE2ADAFD68" box="[1115,1274,691,717]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFA3FFBCFABCACFE2BB6FD69" authorityName="Anker" authorityYear="2011" box="[1325,1430,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFABCACFE2BB6FD69" box="[1325,1430,691,717]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
, especially in the features of the rostrum and major chela, raising at least the possibility that the two species may be synonymous. Indeed, the aforementioned morphological differences between
<taxonomicName id="77C374D7FFA3FFBCFBBDACB62AEDFCB0" authorityName="Banner &amp; Banner" authorityYear="1981" box="[1068,1229,763,789]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFBBDACB62AEDFCB0" box="[1068,1229,763,789]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFA3FFBCFA97ACB62B52FCB1" authorityName="Anker" authorityYear="2011" box="[1286,1394,763,789]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFA97ACB62B52FCB1" box="[1286,1394,763,789]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
are slight and somewhat subjective, and need to be verified when more material becomes available. In addition, the colour pattern of the Saudi Arabian specimens of
<taxonomicName id="77C374D7FFA3FFBCFD4BAD0E2D5DFCF8" authorityName="Banner &amp; Banner" authorityYear="1981" box="[730,893,835,861]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFD4BAD0E2D5DFCF8" box="[730,893,835,861]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
(
<figureCitation id="28F813D1FFA3FFBCFC1EAD0E2A20FCF8" box="[911,1024,835,861]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="19" pageNumber="198">Fig. 8a, b</figureCitation>
) appears to be closer to the pattern of
<taxonomicName id="77C374D7FFA3FFBCFF26AD2A2F02FC25" authorityName="Anker" authorityYear="2011" box="[183,290,871,897]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFF26AD2A2F02FC25" box="[183,290,871,897]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
from Lizard Island (Anker 2011: fig. 8G, H;
<bibRefCitation id="D45272A5FFA3FFBCFCB8AD2A2D93FC24" author="Anker, A." box="[809,947,871,897]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
: fig. 7E), whereas the colour pattern of the Madagascan specimens reported as
<taxonomicName id="77C374D7FFA3FFBCFDA7ADC62CFAFC00" authorityName="Banner &amp; Banner" authorityYear="1981" box="[566,730,907,933]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFDA7ADC62CFAFC00" box="[566,730,907,933]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
(
<bibRefCitation id="D45272A5FFA3FFBCFD7AADC62D5AFC00" author="Anker, A." box="[747,890,907,933]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
: fig. 7A, B) is very similar to that of the New Caledonian specimens herein identified as
<taxonomicName id="77C374D7FFA3FFBCFDECADE22CC8FC6D" authorityName="Anker" authorityYear="2011" box="[637,744,943,969]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFDECADE22CC8FC6D" box="[637,744,943,969]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
(
<figureCitation id="28F813D1FFA3FFBCFD69ADE22D47FC6C" box="[760,871,943,969]" captionStart="FIGURE 9" captionStartId="18.[151,250,1878,1902]" captionTargetBox="[151,1436,151,1860]" captionTargetId="figure-34@18.[151,1436,151,1860]" captionTargetPageId="18" captionText="FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [ae] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="19" pageNumber="198">Fig. 9ae</figureCitation>
). Thus, the present material largely eliminates the colour differences between the two species listed by
<bibRefCitation id="D45272A5FFA3FFBCFD4EAD9E2D5AFC48" author="Anker, A." box="[735,890,979,1005]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012)</bibRefCitation>
. Nonetheless, while contrasting published colour photographs of
<taxonomicName id="77C374D7FFA3FFBCFEDEADBA2FD3FBB4" authorityName="Banner &amp; Banner" authorityYear="1981" box="[335,499,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFEDEADBA2FD3FBB4" box="[335,499,1015,1041]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFA3FFBCFDA0ADBA2CBFFBB5" authorityName="Anker" authorityYear="2011" box="[561,671,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFDA0ADBA2CBFFBB5" box="[561,671,1015,1041]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
(Anker 2011, 2012) with those available for the present material (
<figureCitation id="28F813D1FFA3FFBCFF0EAA562ED0FB90" box="[159,240,1051,1077]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="19" pageNumber="198">Figs. 8</figureCitation>
,
<figureCitation id="28F813D1FFA3FFBCFF6EAA562F2FFB90" box="[255,271,1051,1077]" captionStart="FIGURE 9" captionStartId="18.[151,250,1878,1902]" captionTargetBox="[151,1436,151,1860]" captionTargetId="figure-34@18.[151,1436,151,1860]" captionTargetPageId="18" captionText="FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [ae] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="19" pageNumber="198">9</figureCitation>
), the author noticed one colour difference that could reveal as diagnostic. In the shrimps identified as
<taxonomicName id="77C374D7FFA3FFBCFF06AA722F1BFBFC" authorityName="Banner &amp; Banner" authorityYear="1981" box="[151,315,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFF06AA722F1BFBFC" box="[151,315,1087,1113]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
from the Red Sea (Farasan Islands, Thuwal),
<collectingCountry id="C8D44FC4FFA3FFBCFCC8AA722DC0FBFC" box="[857,992,1087,1113]" name="Madagascar" pageId="19" pageNumber="198">Madagascar</collectingCountry>
(Nosy Bé), and
<collectingCountry id="C8D44FC4FFA3FFBCFB37AA722AC6FBFC" box="[1190,1254,1087,1113]" name="Japan" pageId="19" pageNumber="198">Japan</collectingCountry>
(
<collectingRegion id="7207C1B6FFA3FFBCFB68AA722B44FBFC" box="[1273,1380,1087,1113]" country="Japan" name="Okinawa" pageId="19" pageNumber="198">Okinawa</collectingRegion>
), the carapace is typically white or white locally speckled with small red chromatophores, especially on the anterolateral surface (
<figureCitation id="28F813D1FFA3FFBCFF69AACA2F6BFB04" box="[248,331,1159,1185]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="19" pageNumber="198">Fig. 8a</figureCitation>
;
<bibRefCitation id="D45272A5FFA3FFBCFEC7AACA2FC0FB04" author="Anker, A." box="[342,480,1159,1185]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
: fig. 7A, B;
<bibRefCitation id="D45272A5FFA3FFBCFDFCAACA2D65FB04" author="Goto, R. &amp; Kato, M." box="[621,837,1159,1185]" pageId="19" pageNumber="198" pagination="319 - 330" refId="ref22204" refString="Goto, R. &amp; Kato, M. (2012) Geographic mosaic of mutually exclusive dominance of obligate commensals in symbiotic communities associated with a burrowing echiuran worm. Marine Biology, 159, 319 - 330. https: // doi. org / 10.1007 / s 00227 - 011 - 1810 - 8" type="journal article" year="2012">Goto &amp; Kato 2012</bibRefCitation>
: fig. 1h). In the
<typeStatus id="6F78B1F6FFA3FFBCFC6CAACA2A7FFB04" box="[1021,1119,1159,1185]" pageId="19" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFA3FFBCFB17AACA2AD1FB05" authorityName="Anker" authorityYear="2011" box="[1158,1265,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFB17AACA2AD1FB05" box="[1158,1265,1159,1185]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
from
<collectingCountry id="C8D44FC4FFA3FFBCFAA7AACA2BBDFB04" box="[1334,1437,1159,1185]" name="Australia" pageId="19" pageNumber="198">Australia</collectingCountry>
(Lizard Island) and specimens from
<collectingCountry id="C8D44FC4FFA3FFBCFDBFAAE62CFBFB60" box="[558,731,1195,1221]" name="New Caledonia" pageId="19" pageNumber="198">New Caledonia</collectingCountry>
(Koumac, Pouébo) herein identified as
<taxonomicName id="77C374D7FFA3FFBCFB0BAAE62B24FB61" authorityName="Anker" authorityYear="2011" box="[1178,1284,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFB0BAAE62B24FB61" box="[1178,1284,1195,1221]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
, the carapace is white with large patches of red chromatophores on the anterolateral surface (
<figureCitation id="28F813D1FFA3FFBCFBBAAA822ABEFB4C" box="[1067,1182,1231,1257]" captionStart="FIGURE 9" captionStartId="18.[151,250,1878,1902]" captionTargetBox="[151,1436,151,1860]" captionTargetId="figure-34@18.[151,1436,151,1860]" captionTargetPageId="18" captionText="FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [ae] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="19" pageNumber="198">Fig. 9a, d</figureCitation>
; Anker 2011: fig. 8G;
<bibRefCitation id="D45272A5FFA3FFBCFF06AABE2F01FAA8" author="Anker, A." box="[151,289,1267,1293]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
: fig. 7E). It also appears that the red patches on the pleon of
<taxonomicName id="77C374D7FFA3FFBCFC44AABE2A60FAA9" authorityName="Anker" authorityYear="2011" box="[981,1088,1267,1293]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFC44AABE2A60FAA9" box="[981,1088,1267,1293]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
extend to additional pleonites. Importantly,
<taxonomicName id="77C374D7FFA3FFBCFEBBAB5A2FEDFA94" authorityName="Banner &amp; Banner" authorityYear="1981" box="[298,461,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFEBBAB5A2FEDFA94" box="[298,461,1303,1329]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFA3FFBCFD99AB5A2C54FA95" authorityName="Anker" authorityYear="2011" box="[520,628,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFD99AB5A2C54FA95" box="[520,628,1303,1329]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
were recovered as sister species in the molecular phylogeny of
<bibRefCitation id="D45272A5FFA3FFBCFAC8AB5A2F3FFAF0" author="Chow, L. H. &amp; De Grave, S. &amp; Anker, A. &amp; Poon, K. K. Y. &amp; Ma, K. &amp; Chu, K. &amp; Chan, T. Y. &amp; Tsang, L. M." pageId="19" pageNumber="198" pagination="2898 - 2910" refId="ref21336" refString="Chow, L. H., De Grave, S., Anker, A., Poon, K. K. Y., Ma, K. Y, Chu, K. H, Chan, T. Y. &amp; Tsang, L. M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898 - 2910. https: // doi. org / 10.1111 / evo. 14351" type="journal article" year="2021">
Chow
<emphasis id="82B7D346FFA3FFBCFF06AB762EF1FAF0" box="[151,209,1339,1365]" italics="true" pageId="19" pageNumber="198">et al.</emphasis>
(2021
</bibRefCitation>
: fig. 3), where the former was represented by a specimen from Okinawa (listed in
<bibRefCitation id="D45272A5FFA3FFBCFB45AB762B7FFAF0" author="Anker, A." box="[1236,1375,1339,1365]" pageId="19" pageNumber="198" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
), and the latter by a specimen from
<collectingCountry id="C8D44FC4FFA3FFBCFE7BAB122CB8FADC" box="[490,664,1375,1401]" name="New Caledonia" pageId="19" pageNumber="198">New Caledonia</collectingCountry>
(MNHN-IU-2018-978). The genetic distance between them was not significant, but sufficient to be considered as interspecific (3.1% for 16S; genetic distance presently unavailable for COI). The next step would be to compare sequences from the
<typeStatus id="6F78B1F6FFA3FFBCFCC2ABEA2D95FA64" box="[851,949,1447,1473]" pageId="19" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFA3FFBCFC4DABEA2A66FA65" authorityName="Anker" authorityYear="2011" box="[988,1094,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFA3FFBCFC4DABEA2A66FA65" box="[988,1094,1447,1473]" italics="true" pageId="19" pageNumber="198">A. saurus</emphasis>
</taxonomicName>
and the western Indian Ocean (Red Sea,
<collectingCountry id="C8D44FC4FFA3FFBCFE9FAB862FB7FA40" box="[270,407,1483,1509]" name="Madagascar" pageId="19" pageNumber="198">Madagascar</collectingCountry>
) material of
<taxonomicName id="77C374D7FFA3FFBCFDBAAB862CECFA40" authorityName="Banner &amp; Banner" authorityYear="1981" box="[555,716,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFA3FFBCFDBAAB862CECFA40" box="[555,716,1483,1509]" italics="true" pageId="19" pageNumber="198">A. brevirostris</emphasis>
</taxonomicName>
, to draw the final conclusion about the taxonomic status of the former species.
</paragraph>
</subSubSection>
</treatment>
</document>