324 lines
48 KiB
XML
324 lines
48 KiB
XML
<document id="EE79B0E4D90240D2BE26DD86996CE284" ID-CLB-Dataset="53821" ID-DOI="10.11646/zootaxa.5227.2.7" ID-GBIF-Dataset="721fa183-7dcf-4346-a4bb-596d88921f09" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518696" ID-ZooBank="143C4EB1-DE9A-4E70-8EA6-3397DC511E48" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673310345010" checkinUser="plazi" docAuthor="Kataev, Boris M. & Wrase, David W." docDate="2023" docId="03A7F672C677FFAFF2BB93A35D9278D0" docLanguage="en" docName="zootaxa.5227.2.7.pdf" docOrigin="Zootaxa 5227 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleolissus brevis Kataev & Wrase 2023, sp. n." docType="treatment" docVersion="5" lastPageNumber="287" masterDocId="FF9E8E0AC672FFA7F22C92505C157F1A" masterDocTitle="Three new brachypterous species of the genus Coleolissus (Coleoptera: Carabidae) from China and India" masterLastPageNumber="289" masterPageNumber="279" pageNumber="284" updateTime="1699445493584" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="564C35FD6DAF1CCAD547862E318DB7E5">
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<mods:title id="378D0BC97C7A1613C8495625567AF18E">Three new brachypterous species of the genus Coleolissus (Coleoptera: Carabidae) from China and India</mods:title>
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<mods:name id="D31ACF2F13D886EF2FBBCBD35F25E95D" type="personal">
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<mods:roleTerm id="F8089F7A6934D80C1353F1858F69E508">Author</mods:roleTerm>
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<mods:namePart id="B3F4F80B41D5A640A3DCCE3F279037C7">Kataev, Boris M.</mods:namePart>
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<mods:affiliation id="BD77C36734FAA60EE74EDE49DFCCE30E">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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</mods:name>
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<mods:namePart id="7A7898A07675B9971C086228B46027B0">Wrase, David W.</mods:namePart>
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<mods:nameIdentifier id="71DE690DD3C84C8E87401FF811093B32" type="ORCID">0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation id="41982302940A2CA8B0D472C702A0CB2B">Oderstr. 2, D- 15306 Gusow-Platkow, Germany. carterus @ gmx. de; https: // orcid. org / 0000 - 0003 - 1779 - 2684</mods:affiliation>
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<mods:nameIdentifier id="8139C69CCF937BD00E0D75247D375842" type="email">carterus@gmx.de</mods:nameIdentifier>
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<mods:typeOfResource id="CE512A961BCE68A21774619D505437F5">text</mods:typeOfResource>
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<mods:title id="9E9FDD53EF057C8A7F98D2A3D7B570DD">Zootaxa</mods:title>
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<mods:part id="E40A6C05425344E3DB20B5B54502E0B1">
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<mods:date id="93688BBDF4F58A71D8F0601984E41C71">2023</mods:date>
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<mods:number id="06A3987095275FBB08347B3F75B1F5A0">2023-01-05</mods:number>
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<mods:number id="20CC1E930D525BA9A1F8860929FFF670">5227</mods:number>
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<mods:identifier id="01306323AFE2C94C38DC3F100C6EA1B8" type="DOI">10.11646/zootaxa.5227.2.7</mods:identifier>
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<treatment id="03A7F672C677FFAFF2BB93A35D9278D0" ID-DOI="http://doi.org/10.5281/zenodo.7518660" ID-GBIF-Taxon="210363608" ID-Zenodo-Dep="7518660" LSID="urn:lsid:plazi:treatment:03A7F672C677FFAFF2BB93A35D9278D0" httpUri="http://treatment.plazi.org/id/03A7F672C677FFAFF2BB93A35D9278D0" lastPageId="8" lastPageNumber="287" pageId="5" pageNumber="284">
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<subSubSection id="C31414EFC677FFA2F2BB93A35DAC7D14" box="[151,441,499,526]" pageId="5" pageNumber="284" type="nomenclature">
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<paragraph id="8BB14764C677FFA2F2BB93A35DAC7D14" blockId="5.[151,462,499,562]" box="[151,441,499,526]" pageId="5" pageNumber="284">
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<heading id="D0F9F008C677FFA2F2BB93A35DAC7D14" bold="true" box="[151,441,499,526]" fontSize="11" level="1" pageId="5" pageNumber="284" reason="1">
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<emphasis id="B97A9B76C677FFA2F2BB93A35DAC7D14" bold="true" box="[151,441,499,526]" pageId="5" pageNumber="284">
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<taxonomicName id="4C0E3CE7C677FFA2F2BB93A35D7A7D14" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[151,367,499,526]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
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<emphasis id="B97A9B76C677FFA2F2BB93A35D7A7D14" bold="true" box="[151,367,499,526]" italics="true" pageId="5" pageNumber="284">Coleolissus brevis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A249260DC677FFA2F35A93A45DAC7D14" box="[374,441,500,526]" pageId="5" pageNumber="284" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31414EFC677FFA2F2BB90485DDB7D28" box="[151,462,536,562]" pageId="5" pageNumber="284" type="description">
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<paragraph id="8BB14764C677FFA2F2BB90485DDB7D28" blockId="5.[151,462,499,562]" box="[151,462,536,562]" pageId="5" pageNumber="284">
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(
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<figureCitation id="13355BE1C677FFA2F2B390485D667D28" box="[159,371,536,562]" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="5" pageNumber="284">Figs 11, 12, 19–21</figureCitation>
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,
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<figureCitation id="13355BE1C677FFA2F35390485DD37D28" box="[383,454,536,562]" captionStart-0="FIGURES 27, 28" captionStart-1="FIGURE 29" captionStartId-0="8.[151,264,1646,1670]" captionStartId-1="9.[151,250,1059,1083]" captionTargetBox-0="[167,1412,359,1612]" captionTargetBox-1="[151,1435,181,1035]" captionTargetId-0="figure-122@8.[151,1436,345,1622]" captionTargetId-1="figure-19@9.[151,1435,181,1035]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 27, 28. Coleolissus brevis sp. n., habitus, paratypes. 27, male; 28, female, paratype. Scale bars = 5.0 mm." captionText-1="FIGURE 29. Distribution of Coleolissus spp." figureDoi-0="http://doi.org/10.5281/zenodo.7518704" figureDoi-1="http://doi.org/10.5281/zenodo.7518706" httpUri-0="https://zenodo.org/record/7518704/files/figure.png" httpUri-1="https://zenodo.org/record/7518706/files/figure.png" pageId="5" pageNumber="284">27–29</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31414EFC677FFA2F2BB90305F8C7DD8" pageId="5" pageNumber="284" type="materials_examined">
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<paragraph id="8BB14764C677FFA2F2BB90305E297D84" blockId="5.[151,1437,608,2039]" pageId="5" pageNumber="284">
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<emphasis id="B97A9B76C677FFA2F2BB90305D517D60" bold="true" box="[151,324,608,634]" pageId="5" pageNumber="284">Type material.</emphasis>
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<materialsCitation id="3B664D39C677FFA2F36690305E297D84" ID-GBIF-Occurrence="4134612304" collectingDate="1999-10-06" collectorName="E. Kucera" elevation="1800" location="Khecheopalri" municipality="Khecheoparin" pageId="5" pageNumber="284" specimenCount="1" stateProvince="Sikkim" typeStatus="holotype">
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<typeStatus id="54B5F9C6C677FFA2F36690305DA27D60" box="[330,439,608,634]" pageId="5" pageNumber="284" type="holotype">Holotype</typeStatus>
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: male, “north INDIA-west
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<collectingRegion id="49CA8986C677FFA2F0C890305F5D7D60" box="[740,840,608,634]" country="India" name="Sikkim" pageId="5" pageNumber="284">SIKKIM</collectingRegion>
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/
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<collectingMunicipality id="6BD5DD1EC677FFA2F17090305FEC7D60" box="[860,1017,608,634]" pageId="5" pageNumber="284">Khecheoparin</collectingMunicipality>
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[=?
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<location id="8ED111BFC677FFA2F61D903058D67D60" LSID="urn:lsid:plazi:treatment:03A7F672C677FFAFF2BB93A35D9278D0:8ED111BFC677FFA2F61D903058D67D60" box="[1073,1219,608,634]" municipality="Khecheoparin" name="Khecheopalri" pageId="5" pageNumber="284" stateProvince="Sikkim">Khecheopalri</location>
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] lake / (
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<quantity id="4CF6EA81C677FFA2F736903059677D60" box="[1306,1394,608,634]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="284" unit="m" value="1800.0">
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<elevation id="0023A057C677FFA2F736903059677D60" box="[1306,1394,608,634]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="284" unit="m" value="1800.0">1800 m</elevation>
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</quantity>
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.m.) 4–
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<date id="FFB061A4C677FFA2F29F90D45D367D84" box="[179,291,644,670]" pageId="5" pageNumber="284" value="1999-10-06">
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<collectingDate id="EFF4984CC677FFA2F29F90D45D367D84" box="[179,291,644,670]" pageId="5" pageNumber="284" value="1999-10-06">10.6.1999</collectingDate>
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</date>
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/ lgt.
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<collectorName id="26FB22B2C677FFA2F34990D55DC27D84" box="[357,471,644,671]" pageId="5" pageNumber="284">E. Kučera</collectorName>
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” (cWR).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB14764C677FFA2F2EB90F95F8C7DD8" blockId="5.[151,1437,608,2039]" box="[199,921,680,707]" pageId="5" pageNumber="284">
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<materialsCitation id="3B664D39C677FFA2F2EB90F95F807DD8" ID-GBIF-Occurrence="4134612306" box="[199,917,680,707]" collectingDate="1999-10-06" collectionCode="ZIN" collectorName="E. Kucera" elevation="1800" location="Khecheopalri" municipality="Khecheoparin" pageId="5" pageNumber="284" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sikkim" typeStatus="paratype">
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<typeStatus id="54B5F9C6C677FFA2F2EB90F95D2D7DD9" box="[199,312,681,707]" pageId="5" pageNumber="284" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D088CEDC677FFA2F36F90F85D847DD8" box="[323,401,680,706]" pageId="5" pageNumber="284" type="male">1 male</specimenCount>
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,
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<specimenCount id="9D088CEDC677FFA2F3B090F85E157DD8" box="[412,512,680,706]" pageId="5" pageNumber="284" type="female">1 female</specimenCount>
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, same data as holotype (cWR,
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<collectionCode id="ED1FDFA1C677FFA2F17690F85F987DD8" box="[858,909,680,706]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="284" type="Museum">ZIN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31414EFC677FFA0F2EB909C58DE7C47" lastPageId="7" lastPageNumber="286" pageId="5" pageNumber="284" type="description">
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<paragraph id="8BB14764C677FFA2F2EB909C59897DFC" blockId="5.[151,1437,608,2039]" box="[199,1436,716,743]" pageId="5" pageNumber="284">
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<emphasis id="B97A9B76C677FFA2F2EB909C5D4C7DFC" bold="true" box="[199,345,716,742]" pageId="5" pageNumber="284">Description.</emphasis>
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Body medium-sized for the genus, length
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<quantity id="4CF6EA81C677FFA2F10F909C5FBC7DFD" box="[803,937,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="9.7" metricValueMin="9.2" pageId="5" pageNumber="284" unit="mm" value="9.45" valueMax="9.7" valueMin="9.2">9.2–9.7 mm</quantity>
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, width
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<quantity id="4CF6EA81C677FFA2F1DA909C586B7DFD" box="[1014,1150,716,743]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="5" pageNumber="284" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.9–4.2 mm</quantity>
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.Habitus as in
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<figureCitation id="13355BE1C677FFA2F73A909C598D7DFC" box="[1302,1432,716,742]" captionStart="FIGURES 27, 28" captionStartId="8.[151,264,1646,1670]" captionTargetBox="[167,1412,359,1612]" captionTargetId="figure-122@8.[151,1436,345,1622]" captionTargetPageId="8" captionText="FIGURES 27, 28. Coleolissus brevis sp. n., habitus, paratypes. 27, male; 28, female, paratype. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518704" httpUri="https://zenodo.org/record/7518704/files/figure.png" pageId="5" pageNumber="284">Figs. 27, 28</figureCitation>
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.
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</paragraph>
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<paragraph id="8BB14764C677FFA2F2EB90A05EAD7C48" blockId="5.[151,1437,608,2039]" pageId="5" pageNumber="284">Body black, shiny, iridescent, with green luster on dorsum; labrum, mandibles, lateral beads of pronotum and lateral furrows of elytra dark reddish brown; elytral epipleurae blackish brown; palps, antennae and legs yellowish brown; femora and coxae more or less infuscate.</paragraph>
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<paragraph id="8BB14764C677FFA2F2EB910C584D7AAC" blockId="5.[151,1437,608,2039]" pageId="5" pageNumber="284">Head medium-sized(HWmax/PWmax 0.56–0.57, HWmin/PWmax 0.45–0.48), impunctate.Eyes medium-sized, moderately convex (HWmax/HWmin 1.17–1.24), separated ventrally from buccal fissure at distance about width of antennomere 2 apically. Genae glabrous. Tempora moderately long, slightly convex, sloping to neck constriction. Labrum flat, markedly emarginate along anterior margin. Clypeus very slightly convex, almost straight or very shallowly emarginate along anterior margin, with one setigerous pore at each outer angle. Fronto-clypeal suture fine, almost straight. Fronto-ocular furrows distinct, reaching supraorbital furrows, markedly deepened at clypeus. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated at level of posterior margin of eyes and removed from supraorbital furrows by distance of about width of antennomere 1 basally. Mentum separated from submentum by complete transverse suture, with a prominent median tooth; epilobes moderately widened apically; submentum with one pair of long lateral setae. Ligular sclerite moderately widened distally, with rounded outer angles and one pair of ventroapical setae. Paraglossae glabrous, narrow, markedly projecting beyond ligular sclerite and separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere slightly longer than apical one. Mandibles elongate, somewhat narrow, evenly curved apically, without wrinkles dorsally; left mandible acute at apex. Dorsal microsculpture distinct throughout, consisting of isodiametric meshes. Antennae slender, surpassing pronotal basal edge by approximately two apical antennomeres (in female slightly shorter than in male), pubescent from the middle of antennomere 3, with antennomeres 4–8 about 2.5–3.0 times as long as wide and basal antennomere about as long as antennomere 3.</paragraph>
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<paragraph id="8BB14764C677FFA2F2EB97915E2678B4" blockId="5.[151,1437,608,2039]" pageId="5" pageNumber="284">Pronotum comparatively large and wide (PWmax/PL 1.51–1.53), widest just behind the middle, much more strongly narrowed apically than basally (PWmax/PWmin-ap 1.62–1.69, PWmax/PWmin-bas 1.15–1.22), with one lateral seta inserted in the end of apical third. Sides evenly rounded along entire length; lateral bead complete, very narrow throughout. Apical margin moderately emarginate, narrowly bordered along entire length. Apical angles slightly protruding anteriorly, narrowly rounded at apex. Basal margin slightly rectilinearly concave in middle portion, slightly rounded laterally, bordered almost along entire length (basal bead obliterated medially and occasionally at basal angles), much longer than apical margin (PWmin-bas/PWmin-ap 1.34–1.48) and as long as base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions vaguely outlined, beginning from apical angles or lateral setae, very shallow, wide, markedly widened posteriorly and fused with wide and shallow basal foveae forming oblique laterobasal depressions isolated from each other by a convex area. Median line fine, superficial, not reaching apical and basal margins. Anterior transverse depression very shallow and short. Disc finely and irregularly punctate in laterobasal depressions, with very fine scattered punctures at basal margin medially. Microsculpture visible throughout, consisting of fine transverse meshes.</paragraph>
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<paragraph id="8BB14764C677FFA1F2EB95E85F347ECB" blockId="5.[151,1437,608,2039]" lastBlockId="6.[151,1437,151,573]" lastPageId="6" lastPageNumber="285" pageId="5" pageNumber="284">
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Elytra convex, steeply declined to apex, wide oval (EL/EW 1.43–1.48, EL/PL 2.45–2.53, EW/PWmax 1.12– 1.13), widest at the middle, comparatively strongly narrowed to apex; subapical sinuation very shallow. Humerus angulate, with small acute denticle at apex. Sutural angle blunted or narrowly rounded at tip, not extended posteriorly. Basal edge slightly arched, forming an almost right angle with lateral margin. Striae very fine, superficial, slightly impressed just at apex. Intervals wide on disc, evenly narrowed apically, mainly flat, slightly convex before apex, impunctate. Parascutellar (abbreviate) striole moderately long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 5–7 small discal setigerous pores mostly adjoining stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series with more or less wide gap at middle, consisting of 8–9 setigerous pores in anterior row and of 11–12 pores in posterior row. Lateral groove very narrow throughout, slightly narrower than in
|
||
<taxonomicName id="4C0E3CE7C674FFA1F0B093C35F107EB6" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[668,773,403,429]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="curtus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C674FFA1F0B093C35F107EB6" box="[668,773,403,429]" italics="true" pageId="6" pageNumber="285">C. curtus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C674FFA1F12293C35F5A7EB7" bold="true" box="[782,847,403,429]" pageId="6" pageNumber="285">
|
||
<taxonomicNameLabel id="A249260DC674FFA1F12293C35F5A7EB7" box="[782,847,403,429]" pageId="6" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0E3CE7C674FFA1F1A593C358057EB6" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[905,1040,403,429]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="cognatus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C674FFA1F1A593C358057EB6" box="[905,1040,403,429]" italics="true" pageId="6" pageNumber="285">C. cognatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C674FFA1F63593C3584E7EB7" bold="true" box="[1049,1115,403,429]" pageId="6" pageNumber="285">
|
||
<taxonomicNameLabel id="A249260DC674FFA1F63593C3584E7EB7" box="[1049,1115,403,429]" pageId="6" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Microsculpture on intervals highly obliterated, at most with indistinct transverse lines.
|
||
</paragraph>
|
||
<paragraph id="8BB14764C674FFA1F2EB938B5E0D7EEF" blockId="6.[151,1437,151,573]" box="[199,536,475,501]" pageId="6" pageNumber="285">Wings reduced to a tiny scale.</paragraph>
|
||
<paragraph id="8BB14764C674FFA1F2EB93AF5D037D27" blockId="6.[151,1437,151,573]" pageId="6" pageNumber="285">Pro- and metasternum with very fine and short setae. Metepisternum short, wider than long, weakly narrowed posteriorly.</paragraph>
|
||
<caption id="DF7117ECC674FFA1F2BB956E5D1978FC" ID-DOI="http://doi.org/10.5281/zenodo.7518700" ID-Zenodo-Dep="7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="6" pageNumber="285" startId="6.[151,264,1854,1878]" targetBox="[161,1425,609,1821]" targetPageId="6">
|
||
<paragraph id="8BB14764C674FFA1F2BB956E5D1978FC" blockId="6.[151,1437,1854,2022]" pageId="6" pageNumber="285">
|
||
<emphasis id="B97A9B76C674FFA1F2BB956E5D52784C" bold="true" box="[151,327,1854,1878]" pageId="6" pageNumber="285">FIGURES 5–21.</emphasis>
|
||
<taxonomicName id="4C0E3CE7C674FFA1F361956E5DE1784C" box="[333,500,1854,1878]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97A9B76C674FFA1F361956E5DD7784C" box="[333,450,1854,1878]" italics="true" pageId="6" pageNumber="285">Coleolissus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, male genitalia. 5–8, 13–15,
|
||
<taxonomicName id="4C0E3CE7C674FFA1F134956E5F61784C" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[792,884,1854,1878]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="curtus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C674FFA1F134956E5F61784C" box="[792,884,1854,1878]" italics="true" pageId="6" pageNumber="285">C. curtus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C674FFA1F157956E5FA1784C" bold="true" box="[891,948,1854,1878]" pageId="6" pageNumber="285">
|
||
<taxonomicNameLabel id="A249260DC674FFA1F157956E5FA1784C" box="[891,948,1854,1878]" pageId="6" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18,
|
||
<taxonomicName id="4C0E3CE7C674FFA1F39895325E397860" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[436,556,1890,1914]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="cognatus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C674FFA1F39895325E397860" box="[436,556,1890,1914]" italics="true" pageId="6" pageNumber="285">C. cognatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C674FFA1F01E95325E7E7860" bold="true" box="[562,619,1890,1914]" pageId="6" pageNumber="285">
|
||
<taxonomicNameLabel id="A249260DC674FFA1F01E95325E7E7860" box="[562,619,1890,1914]" pageId="6" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype); 11, 12, 19–21,
|
||
<taxonomicName id="4C0E3CE7C674FFA1F15195325FC27860" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[893,983,1890,1914]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="285" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="B97A9B76C674FFA1F15195325FC27860" box="[893,983,1890,1914]" italics="true" pageId="6" pageNumber="285">C. brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C674FFA1F1F1953258037860" bold="true" box="[989,1046,1890,1914]" pageId="6" pageNumber="285">
|
||
<taxonomicNameLabel id="A249260DC674FFA1F1F1953258037860" box="[989,1046,1890,1914]" pageId="6" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB14764C675FFA0F2EB92C75D607ECB" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="286">Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with shallow longitudinal excavation on inner side. Metafemur with two long setae at posterior margin ventrally and with two short setae on anterior margin. Protibia without longitudinal sulcus on dorsal side, with one stouter and one thinner spine in male and with three comparatively stout, similar sized spines in female. Tarsi glabrous dorsally. In male, protarsomeres 1–4 and mesotarsomeres 2–4 moderately widened, with biseriate adhesive scales ventrally; mesotarsomere 1 weakly widened, with scales only apically, shorter than (in female about as long as) mesotarsomeres 2 and 3 combined. Metatarsus slender, markedly longer than HWmax, with metatarsomeres 2–4 slightly widened distally; metatarsomere 1 elongate, about as long as metatarsomeres 2–3 combined. Tarsomere 5 with two pairs of ventro-lateral setae.</paragraph>
|
||
<paragraph id="8BB14764C675FFA0F2EB938B59057D03" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="286">Abdominal sternites glabrous, only with obligatory setae; apex of last visible sternite (VII) in male subtruncate, in female rounded, in both sexes with two pairs of marginal setae; these setae not distant from margin.</paragraph>
|
||
<paragraph id="8BB14764C675FFA0F2EB90735F777DD7" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="286">
|
||
Aedeagus (
|
||
<figureCitation id="13355BE1C675FFA0F36A90735E317D27" box="[326,548,547,573]" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="7" pageNumber="286">Figs. 11, 12, 19–21</figureCitation>
|
||
) moderately robust, with basal bulb and parameres comparatively large (
|
||
<figureCitation id="13355BE1C675FFA0F74890735CF77D7B" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="7" pageNumber="286">Figs. 20, 21</figureCitation>
|
||
). Median lobe of aedeagus in lateral view (
|
||
<figureCitation id="13355BE1C675FFA0F0F990175F3C7D7B" box="[725,809,583,609]" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="7" pageNumber="286">Fig. 12</figureCitation>
|
||
) arcuate, with ventral margin slightly convex in apical quarter and with apex slightly swollen (
|
||
<figureCitation id="13355BE1C675FFA0F073903B5EA77D9F" box="[607,690,619,645]" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="7" pageNumber="286">Fig. 11</figureCitation>
|
||
); in dorsal view (
|
||
<figureCitation id="13355BE1C675FFA0F152903B5FC77D9F" box="[894,978,619,645]" captionStart="FIGURES 5–21" captionStartId="6.[151,264,1854,1878]" captionTargetBox="[161,1425,609,1821]" captionTargetId="figure-201@6.[151,1436,597,1830]" captionTargetPageId="6" captionText="FIGURES 5–21. Coleolissus spp., male genitalia. 5–8, 13–15, C. curtus sp. n. (5, 6, holotype; 7, 8, 13–15, paratypes from “31 km E Luxi.”); 9, 10, 16–18, C. cognatus sp. n. (holotype); 11, 12, 19–21, C. brevis sp. n. (holotype). 5, 7, 9, 11, apex of median lobe of aedeagus; 6, 8, 10, 12, 13, 16, 19, median lobe; 14, 17, 20, right paramere; 15, 18, 21, left paramere. Left lateral (5–12, 14, 17, 20), dorsal (13, 16, 19), and right lateral (15, 18, 21) views. Scale bars: A = 0.5 mm (5, 7, 9, 11), B = 1.0 mm (6, 8, 10, 12, 13–21)." figureDoi="http://doi.org/10.5281/zenodo.7518700" httpUri="https://zenodo.org/record/7518700/files/figure.png" pageId="7" pageNumber="286">Fig. 19</figureCitation>
|
||
) slightly curved; terminal lamella wide, slightly wider than long, rounded apically. Apical orifice in dorsal position, wide apically. Internal sac with large longitudinal spiny patches at left side medially (in dorsal view).
|
||
</paragraph>
|
||
<paragraph id="8BB14764C675FFA0F2EB908758DE7C47" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="286">
|
||
Female genitalia (
|
||
<figureCitation id="13355BE1C675FFA0F3BC90875E0C7DEB" box="[400,537,726,753]" captionStart="FIGURES 22–26" captionStartId="7.[151,264,1698,1722]" captionTargetBox="[162,1427,957,1675]" captionTargetId="figure-346@7.[151,1436,957,1675]" captionTargetPageId="7" captionText="FIGURES 22–26. Coleolissus spp., female genitalia, paratypes. 22–24, C. curtus sp. n. (22, Tengchong; 23, 24, “31 km E Luxi.”); 25, 26, C. brevis sp. n. 22, 24, 26, laterotergite, gonosubcoxite, and gonocoxite (ventral view); 23, 25, gonocoxite (lateral view). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7518702" httpUri="https://zenodo.org/record/7518702/files/figure.png" pageId="7" pageNumber="286">Figs. 25, 26</figureCitation>
|
||
). Laterotergite asymmetrical, longer than wide, with membranous apical portion and with one preapical seta. Gonosubcoxite about as long as laterotergite, markedly widened posteriorly, without spines. Gonocoxite narrow, slightly curved, narrowed apically, about 0.8 times as long as gonosubcoxite, with very narrow base and with several fine short setae (spines) on outer side; foramen very close to apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31414EFC675FFAFF2EB91375F877898" lastPageId="8" lastPageNumber="287" pageId="7" pageNumber="286" type="etymology">
|
||
<paragraph id="8BB14764C675FFA0F2EB91375D5A7CBF" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="286">
|
||
<emphasis id="B97A9B76C675FFA0F2EB91375D5B7C9B" bold="true" box="[199,334,871,897]" pageId="7" pageNumber="286">Etymology.</emphasis>
|
||
The specific name is a Latin adjective meaning “wide” and referring to the wide and short body of the new species.
|
||
</paragraph>
|
||
<caption id="DF7117ECC675FFA0F2BB94F25DEC7818" ID-DOI="http://doi.org/10.5281/zenodo.7518702" ID-Zenodo-Dep="7518702" httpUri="https://zenodo.org/record/7518702/files/figure.png" pageId="7" pageNumber="286" startId="7.[151,264,1698,1722]" targetBox="[162,1427,957,1675]" targetPageId="7">
|
||
<paragraph id="8BB14764C675FFA0F2BB94F25DEC7818" blockId="7.[151,1437,1698,1794]" pageId="7" pageNumber="286">
|
||
<emphasis id="B97A9B76C675FFA0F2BB94F25D4279A0" bold="true" box="[151,343,1698,1722]" pageId="7" pageNumber="286">FIGURES 22–26.</emphasis>
|
||
<taxonomicName id="4C0E3CE7C675FFA0F37394F25E1C79A0" box="[351,521,1698,1722]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97A9B76C675FFA0F37394F25DC179A0" box="[351,468,1698,1722]" italics="true" pageId="7" pageNumber="286">Coleolissus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, female genitalia, paratypes. 22–24,
|
||
<taxonomicName id="4C0E3CE7C675FFA0F1A494F35FF279A0" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[904,999,1699,1722]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="curtus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C675FFA0F1A494F35FF279A0" box="[904,999,1699,1722]" italics="true" pageId="7" pageNumber="286">C. curtus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C675FFA0F1DC94F3583979A1" bold="true" box="[1008,1068,1699,1723]" pageId="7" pageNumber="286">
|
||
<taxonomicNameLabel id="A249260DC675FFA0F1DC94F3583979A1" box="[1008,1068,1699,1723]" pageId="7" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(22, Tengchong; 23, 24, “31 km E Luxi.”); 25, 26,
|
||
<taxonomicName id="4C0E3CE7C675FFA0F36E94975D8A79C4" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[322,415,1734,1758]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="B97A9B76C675FFA0F36E94975D8A79C4" box="[322,415,1734,1758]" italics="true" pageId="7" pageNumber="286">C. brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C675FFA0F38494975DF179C5" bold="true" box="[424,484,1735,1759]" pageId="7" pageNumber="286">
|
||
<taxonomicNameLabel id="A249260DC675FFA0F38494975DF179C5" box="[424,484,1735,1759]" pageId="7" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
22, 24, 26, laterotergite, gonosubcoxite, and gonocoxite (ventral view); 23, 25, gonocoxite (lateral view). Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB14764C675FFAFF2EB956059217E5B" blockId="7.[151,1437,1840,2011]" lastBlockId="8.[151,1436,151,321]" lastPageId="8" lastPageNumber="287" pageId="7" pageNumber="286">
|
||
<emphasis id="B97A9B76C675FFA0F2EB95605D777850" bold="true" box="[199,354,1840,1866]" pageId="7" pageNumber="286">Comparison.</emphasis>
|
||
In its general habitus and morphology, including the short, transverse metepisternum, this new species is very similar to the two preceding species,
|
||
<taxonomicName id="4C0E3CE7C675FFA0F0F695055F557875" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[730,832,1877,1903]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="curtus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C675FFA0F0F695055F557875" box="[730,832,1877,1903]" italics="true" pageId="7" pageNumber="286">C. curtus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C675FFA0F16A95055F907875" bold="true" box="[838,901,1877,1903]" pageId="7" pageNumber="286">
|
||
<taxonomicNameLabel id="A249260DC675FFA0F16A95055F907875" box="[838,901,1877,1903]" pageId="7" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0E3CE7C675FFA0F1959505582B7875" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[953,1086,1877,1903]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="286" phylum="Arthropoda" rank="species" species="cognatus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C675FFA0F1959505582B7875" box="[953,1086,1877,1903]" italics="true" pageId="7" pageNumber="286">C. cognatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C675FFA0F668950558967875" bold="true" box="[1092,1155,1877,1903]" pageId="7" pageNumber="286">
|
||
<taxonomicNameLabel id="A249260DC675FFA0F668950558967875" box="[1092,1155,1877,1903]" pageId="7" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but can be distinguished from them by having a slightly larger body size, a green luster on dorsum, a more distinct microsculpture on head and pronotum, the humerus with a denticle at apex, the elytral striae finer, superficial, intervals flat, the lateral furrow narrower, and the apex of median lobe of aedeagus only swollen, not bent dorsally. It additionally differs from
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF2F892C75D2F7FAA" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[212,314,151,177]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="curtus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF2F892C75D2F7FAA" box="[212,314,151,177]" italics="true" pageId="8" pageNumber="287">C. curtus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF36C92C75D6A7FAB" bold="true" box="[320,383,151,177]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF36C92C75D6A7FAB" box="[320,383,151,177]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having a less convex body, the pronotum widest behind the middle, and the female genitalia with a wider basal portion of the laterotergite and with a more markedly curved and apically narrowed gonocoxite (a female of
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF30F928F5DBD7FE2" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[291,424,223,249]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="cognatus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF30F928F5DBD7FE2" box="[291,424,223,249]" italics="true" pageId="8" pageNumber="287">C. cognatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF382928F5DF87FE3" bold="true" box="[430,493,223,249]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF382928F5DF87FE3" box="[430,493,223,249]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unknown).
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF0A6928F5F407FE3" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[650,853,223,249]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF0A6928F5F407FE3" box="[650,853,223,249]" italics="true" pageId="8" pageNumber="287">Coleolissus brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF177928F5F8F7FE3" bold="true" box="[859,922,223,249]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF177928F5F8F7FE3" box="[859,922,223,249]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
additionally differs from
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF69B928F59297FE2" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[1207,1340,223,249]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="cognatus" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF69B928F59297FE2" box="[1207,1340,223,249]" italics="true" pageId="8" pageNumber="287">C. cognatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF76E928F59947FE3" bold="true" box="[1346,1409,223,249]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF76E928F59947FE3" box="[1346,1409,223,249]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the pronotum with a shallower emargination of the apical margin, less protruding apical angles and with shallower and more finely punctate lateral depressions; besides, its eyes are less convex and the elytra are narrower.
|
||
</paragraph>
|
||
<caption id="DF7117ECC67AFFAFF2BB943E592C799F" ID-DOI="http://doi.org/10.5281/zenodo.7518704" ID-Zenodo-Dep="7518704" box="[151,1337,1646,1670]" httpUri="https://zenodo.org/record/7518704/files/figure.png" pageId="8" pageNumber="287" startId="8.[151,264,1646,1670]" targetBox="[167,1412,359,1612]" targetPageId="8">
|
||
<paragraph id="8BB14764C67AFFAFF2BB943E592C799F" blockId="8.[151,1337,1646,1670]" box="[151,1337,1646,1670]" pageId="8" pageNumber="287">
|
||
<emphasis id="B97A9B76C67AFFAFF2BB943E5D41799C" bold="true" box="[151,340,1646,1670]" pageId="8" pageNumber="287">FIGURES 27, 28.</emphasis>
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF376943E5E07799C" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[346,530,1646,1670]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF376943E5E07799C" box="[346,530,1646,1670]" italics="true" pageId="8" pageNumber="287">Coleolissus brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF035943E5E47799C" bold="true" box="[537,594,1646,1670]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF035943E5E47799C" box="[537,594,1646,1670]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus, paratypes. 27, male; 28, female, paratype. Scale bars = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB14764C67AFFAFF2EB94E45F877898" blockId="8.[151,1437,1716,1994]" pageId="8" pageNumber="287">
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF2EB94E45E3979D4" authority="Ito, 2016" authorityName="Ito" authorityYear="2016" box="[199,556,1716,1743]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="nakajimai">
|
||
<emphasis id="B97A9B76C67AFFAFF2EB94E45DAB79D4" box="[199,446,1716,1742]" italics="true" pageId="8" pageNumber="287">Coleolissus nakajimai</emphasis>
|
||
<bibRefCitation id="EF9F3A95C67AFFAFF3E994E55E3979D4" author="Ito, N." box="[453,556,1716,1743]" pageId="8" pageNumber="287" pagination="359 - 372" refId="ref6686" refString="Ito, N. (2016) Six new species of the Selenophori carabids from India and adjacent regions (Coleoptera, Carabidae, Hatrpalini). Elytra, New Series, 6 (2), 359 - 372." type="journal article" year="2016">Ito, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
, which was described from
|
||
<specimenCount id="9D088CEDC67AFFAFF14C94E55FCE79D4" box="[864,987,1716,1743]" pageId="8" pageNumber="287" type="female">one female</specimenCount>
|
||
from the Indian state of
|
||
<collectingRegion id="49CA8986C67AFFAFF6C694E4597B79D4" box="[1258,1390,1716,1742]" country="India" name="Tamil Nadu" pageId="8" pageNumber="287">Tamil Nadu</collectingRegion>
|
||
and known to us only from the original description (
|
||
<bibRefCitation id="EF9F3A95C67AFFAFF09D94885F0079E8" author="Ito, N." box="[689,789,1752,1778]" pageId="8" pageNumber="287" pagination="359 - 372" refId="ref6686" refString="Ito, N. (2016) Six new species of the Selenophori carabids from India and adjacent regions (Coleoptera, Carabidae, Hatrpalini). Elytra, New Series, 6 (2), 359 - 372." type="journal article" year="2016">Ito 2016</bibRefCitation>
|
||
), also has vestigial wings, a pronotum with rounded sides and widely rounded basal angles, as well as oval elytra, but it is distinguished from
|
||
<taxonomicName id="4C0E3CE7C67AFFAFF60394AD5884780C" authority="Kataev & Wrase, 2023" authorityName="Kataev & Wrase" authorityYear="2023" box="[1071,1169,1788,1815]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="287" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="B97A9B76C67AFFAFF60394AD5884780C" box="[1071,1169,1788,1815]" italics="true" pageId="8" pageNumber="287">C. brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A9B76C67AFFAFF6BB94AC58C3780C" bold="true" box="[1175,1238,1788,1814]" pageId="8" pageNumber="287">
|
||
<taxonomicNameLabel id="A249260DC67AFFAFF6BB94AC58C3780C" box="[1175,1238,1788,1814]" pageId="8" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at least by having a larger and more elongate body (length
|
||
<quantity id="4CF6EA81C67AFFAFF04895705ED87820" box="[612,717,1824,1851]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="287" unit="mm" value="12.5">12.5 mm</quantity>
|
||
), the elytral intervals convex, the laterotergite with two setae at apex, the gonosubcoxite with one long seta, and the gonocoxite clearly curved, without spines or seta on the outer side and with a foramen on the inner side slightly apart from the tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31414EFC67AFFAFF2EB95DC5D9278D0" pageId="8" pageNumber="287" type="distribution">
|
||
<paragraph id="8BB14764C67AFFAFF2EB95DC5D9278D0" blockId="8.[151,1437,1716,1994]" pageId="8" pageNumber="287">
|
||
<emphasis id="B97A9B76C67AFFAFF2EB95DC5D7478BC" bold="true" box="[199,353,1932,1958]" pageId="8" pageNumber="287">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54B5F9C6C67AFFAFF07395DD5E9A78BD" box="[607,655,1933,1959]" pageId="8" pageNumber="287">type</typeStatus>
|
||
locality [“Khecheoparin” (=? Khecheopalri) Lake,
|
||
<quantity id="4CF6EA81C67AFFAFF6E695DC593478BD" box="[1226,1313,1932,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="287" unit="m" value="1800.0">1800 m</quantity>
|
||
] in
|
||
<collectingRegion id="49CA8986C67AFFAFF76795DC5CC778D0" country="India" name="Sikkim" pageId="8" pageNumber="287">Sikkim State</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31907F4C67AFFAFF2F195E05D0278D0" box="[221,279,1968,1994]" name="India" pageId="8" pageNumber="287">India</collectingCountry>
|
||
(
|
||
<figureCitation id="13355BE1C67AFFAFF30A95E05D6F78D0" box="[294,378,1968,1994]" captionStart="FIGURE 29" captionStartId="9.[151,250,1059,1083]" captionTargetBox="[151,1435,181,1035]" captionTargetId="figure-19@9.[151,1435,181,1035]" captionTargetPageId="9" captionText="FIGURE 29. Distribution of Coleolissus spp." figureDoi="http://doi.org/10.5281/zenodo.7518706" httpUri="https://zenodo.org/record/7518706/files/figure.png" pageId="8" pageNumber="287">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |