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<document id="3B1A11670FC1F9DA81C2ED59A2AF74A9" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFE77D74AC0BD78EFB729A62" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus thunghaihin Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="D8C75297-3063-4B15-ADA6-D86AD5796B96" docUuidSource="ZooBank" docVersion="8" lastPageNumber="29" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="25" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9A98382480B7CB2466E9EDF86F2FF13B">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="C1C04DAC562985E64D1424CE4AE471C8">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="16F2591FCA8790FF7DEA5F512C96A0BF">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="1BCD7C3AB8EA1A73E85CA53A2F15D036">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="2B791255E84F5201914EE33CFE51551E">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="A1646258DBAEBC40DB88694530630234">Panha, Somsak</mods:namePart>
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<mods:affiliation id="43CEA63B65DFE90758299208C2A2717D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<heading id="D0B55C7AFFE77D70AC0BD78EFC1B9BDD" box="[617,970,991,1018]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="4C429095FFE77D70AC0BD78EFCB69BDE" ID-CoL="7CZP9" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[617,871,991,1018]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFE77D70AC0BD78EFCB69BDE" bold="true" box="[617,871,991,1018]" italics="true" pageId="24" pageNumber="25">Tylopus thunghaihin</emphasis>
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<paragraph id="8BFDEB16FFE77D70ACA0D077FCA09C64" blockId="24.[363,1225,991,1088]" box="[706,881,1062,1088]" pageId="24" pageNumber="25">
<figureCitation id="1379F793FFE77D70ACA0D077FC999C64" box="[706,840,1062,1088]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="25.[189,232,1894,1920]" captionStartId-1="26.[189,232,1928,1954]" captionStartId-2="27.[187,230,1930,1956]" captionTargetBox-0="[189,1398,586,1835]" captionTargetBox-1="[335,1275,1069,1864]" captionTargetBox-2="[189,1398,707,1886]" captionTargetId-0="figure@25.[189,1398,586,1835]" captionTargetId-1="figure@26.[333,1278,1058,1870]" captionTargetId-2="figure@27.[189,1398,707,1886]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." captionText-1="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." captionText-2="Fig. 20. Tylopus thunghaihin sp. nov., holotype, Ƌ, right gonopod. AB. Mesal and lateral views, respectively. CF. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837911" figureDoi-1="http://doi.org/10.5281/zenodo.3837913" figureDoi-2="http://doi.org/10.5281/zenodo.3837915" httpUri-0="https://zenodo.org/record/3837911/files/figure.png" httpUri-1="https://zenodo.org/record/3837913/files/figure.png" httpUri-2="https://zenodo.org/record/3837915/files/figure.png" pageId="24" pageNumber="25">Figs 1820</figureCitation>
,
<figureCitation id="1379F793FFE77D70AD36D077FCA09C64" box="[852,881,1062,1088]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="24" pageNumber="25">26</figureCitation>
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<emphasis id="B9363704FFE77D70AEDFD036FEE79CA5" bold="true" box="[189,310,1127,1153]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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The new species seems to be especially similar to
<taxonomicName id="4C429095FFE77D70AD71D0C6FB8F9C95" authority="(Attems, 1953)" baseAuthorityName="Attems" baseAuthorityYear="1953" box="[787,1118,1174,1201]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nodulipes">
<emphasis id="B9363704FFE77D70AD71D0C6FC729C94" box="[787,931,1174,1200]" italics="true" pageId="24" pageNumber="25">T. nodulipes</emphasis>
(
<bibRefCitation id="EFD396E7FFE77D70ADD7D0C7FB879C95" author="Attems C." box="[949,1110,1174,1201]" pageId="24" pageNumber="25" pagination="133 - 230" refId="ref19454" refString="Attems C. 1953. Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'Histoire naturelle, N. S. 5 A: 133 - 230." type="book chapter" year="1953">Attems, 1953</bibRefCitation>
)
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, but differs in the body tegument being dull, coupled with gonopodal process
<emphasis id="B9363704FFE77D70AD3BD0E8FCA09CF7" bold="true" box="[857,881,1209,1235]" pageId="24" pageNumber="25">m</emphasis>
smaller, process
<emphasis id="B9363704FFE77D70AA2BD0E8FB879CF7" bold="true" box="[1097,1110,1209,1235]" pageId="24" pageNumber="25">z</emphasis>
high and rounded, and process
<emphasis id="B9363704FFE77D70AF7ED08CFEFA9CD3" bold="true" box="[284,299,1245,1271]" pageId="24" pageNumber="25">x</emphasis>
pointed.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFE77D70AEDFD14FFE219DAE" pageId="24" pageNumber="25" type="etymology">
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<heading id="D0B55C7AFFE77D70AEDFD14FFE959D1C" bold="true" box="[189,324,1310,1336]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B9363704FFE77D70AEDFD14FFE959D1C" bold="true" box="[189,324,1310,1336]" pageId="24" pageNumber="25">Etymology</emphasis>
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</paragraph>
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To emphasize the
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locality, in Lao “Thung Hai Hin”, commonly translated as the famous Plain of Jars, a noun in apposition.
</paragraph>
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<heading id="D0B55C7AFFE77D70AEDFD1E0FE7B9DEF" bold="true" box="[189,426,1457,1483]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B9363704FFE77D70AEDFD1E0FE7B9DEF" bold="true" box="[189,426,1457,1483]" pageId="24" pageNumber="25">Material examined</emphasis>
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<collectingCountry id="F355AB86FFE77D70AEDFD252FEDF9E39" box="[189,270,1539,1565]" name="Laos" pageId="24" pageNumber="25">LAOS</collectingCountry>
: Ƌ,
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,
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,
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19°25'50&quot;
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,
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103°09'15&quot;
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, ca
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a.s.l.
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, leg.
<collectorName id="26B78EC0FFE77D70AE8DD277FE4C9E64" box="[239,413,1574,1601]" pageId="24" pageNumber="25">S.I. Golovatch</collectorName>
, C.
<collectorName id="26B78EC0FFE77D70AFA9D277FD8D9E65" box="[459,604,1574,1601]" pageId="24" pageNumber="25">Sutcharit, N</collectorName>
. Likhitrakarn (
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).
</paragraph>
</materialsCitation>
<paragraph id="8BFDEB16FFE77D70AE88D239FE8D9EA6" blockId="24.[189,759,1640,1701]" box="[234,348,1640,1666]" pageId="24" pageNumber="25">
<typeStatus id="54F955B4FFE77D70AE88D239FE8D9EA6" box="[234,348,1640,1666]" pageId="24" pageNumber="25" type="paratype">
<emphasis id="B9363704FFE77D70AE88D239FE8D9EA6" bold="true" box="[234,348,1640,1666]" pageId="24" pageNumber="25">Paratype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BFDEB16FFE77D70AEDFD2DAFD269E81" blockId="24.[189,759,1640,1701]" box="[189,759,1675,1701]" pageId="24" pageNumber="25">
<collectingCountry id="F355AB86FFE77D70AEDFD2DAFEDF9E81" box="[189,270,1675,1701]" name="Laos" pageId="24" pageNumber="25">LAOS</collectingCountry>
:
<specimenCount id="9D44209FFFE77D70AF7BD2DAFE949E81" box="[281,325,1675,1701]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
, same data as for
<typeStatus id="54F955B4FFE77D70AC7BD2DAFD519E81" box="[537,640,1675,1701]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED5373D3FFE77D70ACEDD2DAFD3A9E81" box="[655,747,1675,1701]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="24" pageNumber="25">CUMZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFE77D74AEDFD29DFBF79996" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25" type="description">
<paragraph id="8BFDEB16FFE77D70AEDFD29DFE9F9EC2" blockId="24.[189,334,1740,1766]" box="[189,334,1740,1766]" pageId="24" pageNumber="25">
<heading id="D0B55C7AFFE77D70AEDFD29DFE9F9EC2" bold="true" box="[189,334,1740,1766]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B9363704FFE77D70AEDFD29DFE9F9EC2" bold="true" box="[189,334,1740,1766]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFE77D70AEDFD2AAFD8A9F85" blockId="24.[189,1398,1786,1953]" pageId="24" pageNumber="25">
MEASUREMENTS AND COLOUR. Length 26.1 (Ƌ) or
<quantity id="4CBA46F3FFE77D70AC9CD2ABFCB89F31" box="[766,873,1786,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="24" pageNumber="25" unit="mm" value="26.2">26.2 mm</quantity>
(
<specimenCount id="9D44209FFFE77D70AD1FD2AAFC419F31" box="[893,912,1787,1813]" pageId="24" pageNumber="25" type="female"></specimenCount>
), width of midbody pro- and metazonae 2.02 and
<quantity id="4CBA46F3FFE77D70AF48D34FFE429F1C" box="[298,403,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="3.02" pageId="24" pageNumber="25" unit="mm" value="3.02">3.02 mm</quantity>
(Ƌ) or 2.04 and
<quantity id="4CBA46F3FFE77D70AC3BD34FFD139F1C" box="[601,706,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="3.04" pageId="24" pageNumber="25" unit="mm" value="3.04">3.04 mm</quantity>
(
<specimenCount id="9D44209FFFE77D70ACB6D34FFD369F1C" box="[724,743,1822,1848]" pageId="24" pageNumber="25" type="female"></specimenCount>
), respectively. Live coloration brown black (
<figureCitation id="1379F793FFE77D70AA99D34FFAB89F1C" box="[1275,1385,1822,1848]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="24" pageNumber="25">Fig. 18A</figureCitation>
); antennae, paraterga and legs dark brown. Coloration in alcohol after one year of preservation blackish, paraterga dark brown; venter and a few basal podomeres light brown to yellow-brown, legs increasingly darker brown distally (
<figureCitation id="1379F793FFE77D70AFAAD3D6FDCD9F85" box="[456,540,1927,1953]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="24" pageNumber="25">Fig. 18</figureCitation>
BJ).
</paragraph>
<paragraph id="8BFDEB16FFE67D71AED9D55BFCC799F7" blockId="25.[187,1399,265,468]" pageId="25" pageNumber="26">
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short (
<figureCitation id="1379F793FFE67D71AEA6D57DFEE39962" box="[196,306,300,326]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="25" pageNumber="26">Fig. 18A</figureCitation>
), extending behind body segment 3 (Ƌ) or 2 (
<specimenCount id="9D44209FFFE67D71AD3DD57DFCA39962" box="[863,882,300,326]" pageId="25" pageNumber="26" type="female"></specimenCount>
) when stretched dorsally. In width, head &lt;segment 3 &lt;4 &lt;5 &lt;collum &lt;segment 2 &lt;616 (Ƌ,
<specimenCount id="9D44209FFFE67D71AD49D501FC91994E" box="[811,832,336,362]" pageId="25" pageNumber="26" type="female"></specimenCount>
); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 1+1 intermediate and 3+3 posterior; with a small lateral incision near midway (
<figureCitation id="1379F793FFE67D71AC10D5C7FCD89994" box="[626,777,406,432]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="25" pageNumber="26">Fig. 18B, D</figureCitation>
); caudal corner of paraterga very broadly rounded, declined ventrad, not surpassing rear tergal margin.
</paragraph>
<caption id="DF3DBB9EFFE67D71AEDFD337FC6E9FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837911" ID-Zenodo-Dep="3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1894,1920]" targetBox="[189,1398,586,1835]" targetPageId="25">
<paragraph id="8BFDEB16FFE67D71AEDFD337FC6E9FCE" blockId="25.[189,1399,1894,2026]" pageId="25" pageNumber="26">
<emphasis id="B9363704FFE67D71AEDFD337FECF9FA4" bold="true" box="[189,286,1894,1920]" pageId="25" pageNumber="26">Fig. 18.</emphasis>
<taxonomicName id="4C429095FFE67D71AF48D337FDCF9FA4" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[298,542,1894,1920]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFE67D71AF48D337FDCF9FA4" box="[298,542,1894,1920]" italics="true" pageId="25" pageNumber="26">Tylopus thunghaihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE67D71AC48D336FD5A9FA5" box="[554,651,1895,1921]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE67D71AD5DD337FC859FA4" bold="true" box="[831,852,1894,1920]" pageId="25" pageNumber="26">A</emphasis>
. Habitus, live coloration
<emphasis id="B9363704FFE67D71AAF3D337FB199FA4" bold="true" box="[1169,1224,1894,1920]" pageId="25" pageNumber="26">BC</emphasis>
. Anterior part of body, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE67D71AC9ED3DBFCE29F80" bold="true" box="[764,819,1930,1956]" pageId="25" pageNumber="26">DE</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE67D71AF38D3FCFE409FE3" bold="true" box="[346,401,1965,1991]" pageId="25" pageNumber="26">FH</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively.
<emphasis id="B9363704FFE67D71AA8BD3FCFAC39FE3" bold="true" box="[1257,1298,1965,1991]" pageId="25" pageNumber="26">IJ</emphasis>
. Sternal cones between coxae 4, caudal and sublateral views, respectively.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE57D73AEDFD55DFB5F996D" blockId="26.[189,1399,267,998]" lastBlockId="27.[189,1398,267,329]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
BODY. Tegument dull, shining, prozonae finely shagreened; metaterga often roughly microgranulate and rugulose, leathery; surface below paraterga more delicately, but still sufficiently clearly microgranulate and rugulose (
<figureCitation id="1379F793FFE57D72AF0AD503FE6F9948" box="[360,446,338,364]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18</figureCitation>
AF, H). Metaterga with two transverse rows of small setigerous cones: 2+
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anterior (pre-sulcus) row and 3+3 setae usually borne on low, oblong, rounded tubercles in posterior (post-sulcus) row, except for metaterga 1719, these latter bearing 3(2)+3(2) setae in anterior row and 4+
<quantity id="4CBA46F3FFE57D72AEBED5EAFED699F1" box="[220,263,443,469]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="26" pageNumber="27" unit="in" value="4.0">4 in</quantity>
posterior row. Tergal setae simple, slender, often abraded, about as long as 1/3 of metazonite.Axial line visible both on pro- and metazonae. Paraterga strongly developed (
<figureCitation id="1379F793FFE57D72AA62D58FFB7D99DD" box="[1024,1196,478,505]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18B, D, F</figureCitation>
), especially so in Ƌ, set high (at upper ¼ of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, extending increasingly behind rear tergal margin, posterior edge slightly oblique. Calluses on paraterga delimited by a sulcus only dorsally. Paratega 2 broad, anterior edge angular, lateral edge with three evident incisions. Lateral edge of following paraterga with a clear incision in anterior 1/3 (
<figureCitation id="1379F793FFE57D72AFBCD6DFFD469A8D" box="[478,663,654,681]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18B, D, F</figureCitation>
). Paraterga 1619 with caudal corner extending increasingly caudally, tip clearly curved mesad (
<figureCitation id="1379F793FFE57D72AC09D6E3FD079AE8" box="[619,726,690,716]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18F</figureCitation>
). Ozopores evident, lateral, lying in an ovoid groove at about 1/
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front of posterior edge of metaterga. Transverse sulcus distinct (
<figureCitation id="1379F793FFE57D72AA30D684FAD79ACB" box="[1106,1286,725,751]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18B, D, F</figureCitation>
), slightly incomplete on segments 4 and 18, complete on segments 517, narrow, wave-shaped, rather deep, not reaching bases of paraterga, ribbed at bottom. Stricture between pro- and metazonae evident, wide, deep, ribbed at bottom down to base of paraterga (
<figureCitation id="1379F793FFE57D72AD75D76FFCBD9B7D" box="[791,876,830,857]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18</figureCitation>
BG). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 24 (Ƌ,
<specimenCount id="9D44209FFFE57D72AC91D733FCD69B58" box="[755,775,866,892]" pageId="26" pageNumber="27" type="female"></specimenCount>
), increasingly reduced to a front bulge and a caudal tooth until segment 7, retained only as a sharp caudal tooth until segment 16, thereafter missing (Ƌ), or increasingly reduced and remaining only a sharp caudal tooth until segment 15, thereafter missing (
<specimenCount id="9D44209FFFE57D72AB3AD7F9FABA9BE6" box="[1368,1387,936,962]" pageId="26" pageNumber="27" type="female"></specimenCount>
) (
<figureCitation id="1379F793FFE57D72AEA4D79AFEAD9BC2" box="[198,380,971,998]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18C, E, H</figureCitation>
). Epiproct (
<figureCitation id="1379F793FFE57D72AC68D79AFD8F9BC1" box="[522,606,971,997]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="26" pageNumber="27">Fig. 18</figureCitation>
FH) conical, flattened dorsoventrally, with two evident, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae large, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (
<figureCitation id="1379F793FFE47D73AA76D57FFB50996D" box="[1044,1153,302,329]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="27" pageNumber="28">Fig. 18G</figureCitation>
).
</paragraph>
<caption id="DF3DBB9EFFE57D72AEDFD3D9FEEB9FCD" ID-DOI="http://doi.org/10.5281/zenodo.3837913" ID-Zenodo-Dep="3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1928,1954]" targetBox="[335,1275,1069,1864]" targetPageId="26">
<paragraph id="8BFDEB16FFE57D72AEDFD3D9FEEB9FCD" blockId="26.[189,1399,1928,2025]" pageId="26" pageNumber="27">
<emphasis id="B9363704FFE57D72AEDFD3D9FECF9F86" bold="true" box="[189,286,1928,1954]" pageId="26" pageNumber="27">Fig. 19.</emphasis>
<taxonomicName id="4C429095FFE57D72AF4BD3D9FDCF9F86" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[297,542,1928,1954]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFE57D72AF4BD3D9FDCF9F86" box="[297,542,1928,1954]" italics="true" pageId="26" pageNumber="27">Tylopus thunghaihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE57D72AC4BD3D8FD5A9F87" box="[553,651,1929,1955]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE57D72AD5CD3D9FCA49F86" bold="true" box="[830,885,1928,1954]" pageId="26" pageNumber="27">AB</emphasis>
. Right gonopod, lateral and mesal views, respectively.
<emphasis id="B9363704FFE57D72AF3AD3FAFEBC9FE1" bold="true" box="[344,365,1963,1989]" pageId="26" pageNumber="27">C</emphasis>
. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE47D73AEDFD521FE6A9A79" blockId="27.[189,1399,368,606]" pageId="27" pageNumber="28">
STERNA. Sparsely setose, with a small, but evident cone caudally near each coxa, rear cones being a little larger than fore ones (
<figureCitation id="1379F793FFE47D73AFA1D5C2FDE19989" box="[451,560,403,429]" captionStart="Fig" captionStartId="26.[189,232,1928,1954]" captionTargetBox="[335,1275,1069,1864]" captionTargetId="figure@26.[333,1278,1058,1870]" captionTargetPageId="26" captionText="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="27" pageNumber="28">Fig. 19C</figureCitation>
); cross-impressions shallow; a large, central, slightly bifid, setose lobe between Ƌ coxae 4 (
<figureCitation id="1379F793FFE47D73AFCCD5E7FDF899F5" box="[430,553,438,465]" captionStart="Fig" captionStartId="25.[189,232,1894,1920]" captionTargetBox="[189,1398,586,1835]" captionTargetId="figure@25.[189,1398,586,1835]" captionTargetPageId="25" captionText="Fig. 18. Tylopus thunghaihin sp. nov., holotype, Ƌ. A. Habitus, live coloration BC. Anterior part of body, dorsal and lateral views, respectively. DE. Segments 10 and 11, dorsal and lateral views, respectively. FH. Posterior part of body, dorsal, ventral and lateral views, respectively. IJ. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837911" httpUri="https://zenodo.org/record/3837911/files/figure.png" pageId="27" pageNumber="28">
Fig. 18I
<emphasis id="B9363704FFE47D73AC6ED5E6FDCA99F5" bold="true" box="[524,539,439,465]" pageId="27" pageNumber="28"></emphasis>
J
</figureCitation>
). Legs long and slender, midbody ones ca 1.21.4 (Ƌ) or 1.11.2 (
<specimenCount id="9D44209FFFE47D73AB5AD5E7FA9A99F4" box="[1336,1355,438,464]" pageId="27" pageNumber="28" type="female"></specimenCount>
) as long as body height; legs of Ƌ segments 517 with femur showing a field of ventral microgranulations, while postfemur, tibia and tarsus each with 13 minute adenostyles (tubercles), prefemora not swollen (
<figureCitation id="1379F793FFE47D73AEA4D671FEE39A1E" box="[198,306,544,570]" captionStart="Fig" captionStartId="26.[189,232,1928,1954]" captionTargetBox="[335,1275,1069,1864]" captionTargetId="figure@26.[333,1278,1058,1870]" captionTargetPageId="26" captionText="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="27" pageNumber="28">Fig. 19G</figureCitation>
), Ƌ telopodites with particularly dense short setae ventrally (
<figureCitation id="1379F793FFE47D73AD98D671FBB79A1E" box="[1018,1126,544,570]" captionStart="Fig" captionStartId="26.[189,232,1928,1954]" captionTargetBox="[335,1275,1069,1864]" captionTargetId="figure@26.[333,1278,1058,1870]" captionTargetPageId="26" captionText="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="27" pageNumber="28">Fig. 19G</figureCitation>
), tarsal brushes present until body segment 8.
</paragraph>
<caption id="DF3DBB9EFFE47D73AED9D3DBFEBC9FCF" ID-DOI="http://doi.org/10.5281/zenodo.3837915" ID-Zenodo-Dep="3837915" httpUri="https://zenodo.org/record/3837915/files/figure.png" pageId="27" pageNumber="28" startId="27.[187,230,1930,1956]" targetBox="[189,1398,707,1886]" targetPageId="27">
<paragraph id="8BFDEB16FFE47D73AED9D3DBFEBC9FCF" blockId="27.[187,1399,1930,2027]" pageId="27" pageNumber="28">
<emphasis id="B9363704FFE47D73AED9D3DBFECD9F80" bold="true" box="[187,284,1930,1956]" pageId="27" pageNumber="28">Fig. 20.</emphasis>
<taxonomicName id="4C429095FFE47D73AF4BD3DBFDCF9F80" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[297,542,1930,1956]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="thunghaihin" status="sp. nov.">
<emphasis id="B9363704FFE47D73AF4BD3DBFDCF9F80" box="[297,542,1930,1956]" italics="true" pageId="27" pageNumber="28">Tylopus thunghaihin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE47D73AC48D3DAFD5D9F81" box="[554,652,1931,1957]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, right gonopod.
<emphasis id="B9363704FFE47D73AD9DD3DBFBE79F80" bold="true" box="[1023,1078,1930,1956]" pageId="27" pageNumber="28">AB</emphasis>
. Mesal and lateral views, respectively.
<emphasis id="B9363704FFE47D73AF3AD3FCFE5F9FE3" bold="true" box="[344,398,1965,1991]" pageId="27" pageNumber="28">CF</emphasis>
. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE37D74AEDFD55AFBF79996" blockId="28.[189,1399,267,435]" pageId="28" pageNumber="29">
GONOPOD. Rather complex (
<figureCitation id="1379F793FFE37D74AF9ED55AFD5C9902" box="[508,653,267,294]" captionStart="Fig" captionStartId="26.[189,232,1928,1954]" captionTargetBox="[335,1275,1069,1864]" captionTargetId="figure@26.[333,1278,1058,1870]" captionTargetPageId="26" captionText="Fig. 19. Tylopus thunghaihin sp. nov., holotype, Ƌ. AB. Right gonopod, lateral and mesal views, respectively. C. Leg of segment 9. Scale bars: AB = 0.2 mm; C = 1 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837913" httpUri="https://zenodo.org/record/3837913/files/figure.png" pageId="28" pageNumber="29">
Figs 19A
<emphasis id="B9363704FFE37D74AC0AD55DFDA69902" bold="true" box="[616,631,268,294]" pageId="28" pageNumber="29"></emphasis>
B
</figureCitation>
,
<figureCitation id="1379F793FFE37D74ACF5D55AFD699901" box="[663,696,267,293]" captionStart="Fig" captionStartId="27.[187,230,1930,1956]" captionTargetBox="[189,1398,707,1886]" captionTargetId="figure@27.[189,1398,707,1886]" captionTargetPageId="27" captionText="Fig. 20. Tylopus thunghaihin sp. nov., holotype, Ƌ, right gonopod. AB. Mesal and lateral views, respectively. CF. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3837915" httpUri="https://zenodo.org/record/3837915/files/figure.png" pageId="28" pageNumber="29">20</figureCitation>
); coxa slightly curved caudad, sparsely setose distoventrally. Femorite slightly curved and stout, with an evident mesal groove and a clear distolateral sulcus demarcating a postfemoral part; process
<emphasis id="B9363704FFE37D74ACC3D503FD609948" bold="true" box="[673,689,338,364]" pageId="28" pageNumber="29">h</emphasis>
high, strongly twisted, tip evidently bifid; process
<emphasis id="B9363704FFE37D74AB70D503FAFB9948" bold="true" box="[1298,1322,338,364]" pageId="28" pageNumber="29">m</emphasis>
short; process
<emphasis id="B9363704FFE37D74AF40D524FEFE99AB" bold="true" box="[290,303,373,399]" pageId="28" pageNumber="29">z</emphasis>
rather long, sausage-shaped; solenophore (
<emphasis id="B9363704FFE37D74AD27D524FCA099AB" bold="true" box="[837,881,373,399]" pageId="28" pageNumber="29">sph</emphasis>
) large, strongly coiled, expanded distally, flattened and clearly bifid; process
<emphasis id="B9363704FFE37D74AC35D5C9FDB79996" bold="true" box="[599,614,408,434]" pageId="28" pageNumber="29">x</emphasis>
large, dentiform and pointed dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFE37D74AEDFD58BFB729A62" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BFDEB16FFE37D74AEDFD58BFEFE99D0" blockId="28.[189,303,474,500]" box="[189,303,474,500]" pageId="28" pageNumber="29">
<heading id="D0B55C7AFFE37D74AEDFD58BFEFE99D0" bold="true" box="[189,303,474,500]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B9363704FFE37D74AEDFD58BFEFE99D0" bold="true" box="[189,303,474,500]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFE37D74AEDFD659FB729A62" blockId="28.[189,1398,520,582]" pageId="28" pageNumber="29">
This species has been found under stones, each specimen located in its own burrow of ca
<quantity id="4CBA46F3FFE37D74AA8DD659FAEB9A07" box="[1263,1338,520,547]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="cm" value="30.0">30 cm</quantity>
long under its own stone and forming a kind of small cavern, all in the Plain of Jars,
<collectingCountry id="F355AB86FFE37D74AA00D67DFB4E9A62" box="[1122,1183,556,582]" name="Laos" pageId="28" pageNumber="29">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>